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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 277<br />

purple, <strong>the</strong> st<strong>and</strong>ard semicircular, ca. 10 mm long,<br />

glabrous, <strong>the</strong> wings obovate, unguiculate, longer<br />

than <strong>the</strong> st<strong>and</strong>ard, <strong>the</strong> keel spirally twisted, ca. 1<br />

cm long; stamens 10, diadelphous, <strong>the</strong> vexillar<br />

stamen broadened at <strong>the</strong> base; ovary strigulose.<br />

Fruit an oblong legume, slightly flattened, straight<br />

or slightly curved, 9-12 × 1-1.2 cm, glabrous,<br />

long-apiculate at <strong>the</strong> apex, dehiscent by valves<br />

that separate longitudinally. Seeds oblongreniform,<br />

1-1.2 mm long, <strong>of</strong> various colors,<br />

reddish brown, grayish with dark spots, pink.<br />

Phenology: Collected in flower <strong>and</strong> fruit from<br />

May to July <strong>and</strong> in November.<br />

18. PUERARIA<br />

Status: Exotic, cultivated <strong>and</strong> naturalized,<br />

uncommon.<br />

Distribution: In areas <strong>of</strong> degraded vegetation,<br />

along trails, on fences, <strong>and</strong> in thickets. Also<br />

throughout <strong>the</strong> Antilles. Native to Mesoamerica,<br />

with numerous forms cultivated for millennia in<br />

tropical continental America. Widely cultivated<br />

throughout <strong>the</strong> world.<br />

Excluded Species:<br />

Phaseolus polystachyos (L.) Britton was<br />

reported for <strong>Puerto</strong> <strong>Rico</strong> by Liogier (1982, as P.<br />

polystachios), based on an erroneous<br />

identification <strong>of</strong> Phaseolus lunatus (Liogier, et al.<br />

31642).<br />

Herbaceous or woody vines, twining. Leaves alternate, trifoliolate; stipules ovate to linear, persistent;<br />

stipels minute. Inflorescences <strong>of</strong> axillary or terminal pseudoracemes; bracts minute. Calyx campanulate,<br />

with 5 unequal lobes; corolla blue or violet, <strong>the</strong> st<strong>and</strong>ard obovate, retuse at <strong>the</strong> apex, unguiculate <strong>and</strong><br />

auriculate at <strong>the</strong> base, <strong>the</strong> wings unguiculate, with a curved appendage at <strong>the</strong> base <strong>of</strong> <strong>the</strong> inner margin,<br />

<strong>the</strong> keel slightly longer than <strong>the</strong> wings; stamens 10, monadelphous or diadelphous; ovary superior,<br />

sessile, with several ovules, <strong>the</strong> style glabrous, curved, <strong>the</strong> stigma capitate. Fruit a linear legume,<br />

flattened, dehiscent by valves that twist on opening; seeds oblong, numerous. A genus <strong>of</strong> 6 species<br />

native to sou<strong>the</strong>rn Asia, with one species introduced in <strong>the</strong> New World as a forage plant.<br />

1. Pueraria phaseoloides (Roxb.) Benth., J.<br />

Linn. Soc., Bot. 9: 125. 1865.<br />

Fig. 111. A-D<br />

BASIONYM: Dolichos phaseoloides Roxb.<br />

Kudzu tropical, Corsú<br />

Herbaceous vine, twining, much branched,<br />

attainig 15 m in length. Stems cylindrical, pilose.<br />

Leaves alternate, trifoliolate; leaflets 3-12(14) ×<br />

2.9-8.7(13) cm, chartaceous, ovate or rhombic,<br />

<strong>the</strong> lateral ones asymmetrical, <strong>the</strong> apex acute, <strong>the</strong><br />

base cuneate on <strong>the</strong> central leaflet, rounded-obtuse<br />

on <strong>the</strong> lateral ones, <strong>the</strong> margins entire; upper<br />

surface dark green, dull, pubescent, especially on<br />

<strong>the</strong> veins; lower surface pale green, strigose, with<br />

prominent venation; petiolules swollen, 4-5 mm<br />

long, pubescent; petioles sulcate, pubescent, up<br />

to 12 cm long, with <strong>the</strong> base swollen; stipules<br />

narrowly lanceolate, 3-5 mm long; stipels<br />

subulate, minute, persistent. Pseudoracemes<br />

axillary, up to 25 cm long, with 2-3 flowers per<br />

node; bracts minute, persistent; peduncles<br />

pubescent. Calyx campanulate, ca. 5 mm long.<br />

Legume linear, flattened, 6-9 cm long, slightly<br />

curved, dehiscent by valves that open in a spiral,<br />

<strong>the</strong> valves septate between <strong>the</strong> seeds. Seeds<br />

numerous, ca. 3 mm long, oblong, dark brown to<br />

almost black.<br />

Phenology: Flowering <strong>and</strong> fruiting from<br />

November to March.<br />

Status: Exotic, naturalized, very aggressive<br />

<strong>and</strong> very common.<br />

Distribution: In moist disturbed areas, at<br />

middle <strong>and</strong> lower elevations. Also on St. John.<br />

Native to tropical Asia, introduced in Africa <strong>and</strong><br />

<strong>the</strong> Americas as a forage plant.<br />

Public Forests: Carite, El Yunque, Maricao,<br />

Río Abajo, <strong>and</strong> Tortuguero.<br />

Commentary: Apparently originally<br />

introduced on l<strong>and</strong>s <strong>of</strong> <strong>the</strong> Agricultural<br />

Experiment Station in Mayagüez in <strong>Puerto</strong> <strong>Rico</strong><br />

in 1940 from material from Malaya; today this<br />

species has a wide distribution throughout <strong>Puerto</strong><br />

<strong>Rico</strong>.

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