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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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278<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

19. RHYNCHOSIA<br />

Herbaceous or woody vines, twining. Leaves alternate, trifoliolate, <strong>the</strong> lower surface with numerous<br />

yellow resinous dots; stipules deciduous; stipels minute. Inflorescences <strong>of</strong> axillary racemes, with <strong>the</strong><br />

flowers sparse or clustered; bracts minute, persistent or deciduous. Calyx campanulate, with 4-5 elongate<br />

lobes; corolla yellow, <strong>the</strong> st<strong>and</strong>ard obovate or rounded, slightly retuse at <strong>the</strong> apex, unguiculate <strong>and</strong><br />

auriculate at <strong>the</strong> base, <strong>the</strong> wings unguiculate, with a curved appendage at <strong>the</strong> base <strong>of</strong> <strong>the</strong> inner margin,<br />

<strong>the</strong> keel scarcely longer than <strong>the</strong> wings; stamens 10, monadelphous or diadelphous; ovary superior,<br />

short-stipitate, with few or numerous ovules, <strong>the</strong> style glabrous, curved, <strong>the</strong> stigma capitate. Fruit an<br />

oblong legume, flattened, apiculate at <strong>the</strong> apex, usually dehiscent by valves that twist on opening;<br />

seeds few or numerous, flattened, rounded or elliptical. A genus <strong>of</strong> about 200 species, <strong>of</strong> pantropical<br />

distribution.<br />

Key to <strong>the</strong> species <strong>of</strong> Rhynchosia<br />

1a. Robust lianas that attain 8-10 m in length; mature stems flattened, forming a ribbon up to 2.5 cm<br />

wide; seeds red <strong>and</strong> black ….………………………………….... 2. R. phaseoloides<br />

1b. Slightly woody vines, usually less than 5 m in length; mature stems cylindrical, less than 5 mm in<br />

diameter; seeds brown ……………………………………………………………. 2<br />

2a. Lower surface <strong>of</strong> <strong>the</strong> leaflets with <strong>the</strong> tertiary venation not prominent; calyx 2.5-3 mm long, <strong>the</strong><br />

sepals short-subulate, as long as or twice as long as <strong>the</strong> tubular portion <strong>of</strong> <strong>the</strong> calyx<br />

..............…………………..…..……………………………...……………………1. R. minima<br />

2b. Lower surface <strong>of</strong> <strong>the</strong> leaflets with <strong>the</strong> reticulate tertiary venation prominent; calyx 6-10 mm long,<br />

<strong>the</strong> sepals linear-lanceolate, three times as long as <strong>the</strong> tubular portion <strong>of</strong> <strong>the</strong> calyx<br />

...................……………..…….…………………………………….…………….3. R. reticulata<br />

1. Rhynchosia minima (L.) DC., Prodr. 2: 385.<br />

1825.<br />

Fig. 111. E-H<br />

BASIONYM: Dolichos minimus L.<br />

SYNONYMS: Rhynchosia minima var. lutea Eggers<br />

Rhynchosia minima var. pauciflora Kuntze<br />

Dolicholus minimus (L.) Medic.<br />

Frijolillo, Habilla<br />

Non-woody vine, twining, attainig 1-4 m in<br />

length. Stems slender, densely pubescent, striate,<br />

cylindrical, with numerous resinous dots in <strong>the</strong><br />

furrows. Leaves alternate, trifoliolate,<br />

chartaceous; margins entire, slightly revolute;<br />

upper surface dull, glabrous or white-pubescent,<br />

especially on <strong>the</strong> veins, with numerous orange<br />

resinous dots; lower surface pale green, glabrous,<br />

puberulent or pubescent, with minute orange<br />

resinous dots <strong>and</strong> <strong>the</strong> primary <strong>and</strong> secondary<br />

venation prominent; terminal leaflet rhombic, 0.9-<br />

2.8(7.5) × 0.6-2.8(6.3) cm, <strong>the</strong> apex acute or<br />

obtuse, <strong>the</strong> base cuneate; lateral leaflets<br />

asymmetrical, elliptical-ovate, slightly smaller<br />

than <strong>the</strong> terminal leaflet, <strong>the</strong> base obtuse;<br />

petiolules densely pubescent, 1-1.5 mm long;<br />

petiole striate, densely pubescent, 1.5-2 cm long;<br />

stipules subulate, 2-2.5 mm long, densely<br />

pubescent; stipels subulate, ca. 3 mm long.<br />

Inflorescence a raceme, as long as or longer than<br />

<strong>the</strong> leaves. Calyx green, <strong>the</strong> lobes lanceolate, 2-3<br />

mm long; corolla yellow with purple or brown<br />

venation, <strong>the</strong> st<strong>and</strong>ard 5 mm long, orbicularovate;<br />

wings oblong, unguiculate, 4-5.5 mm long,<br />

with resinous dots. Legume falcate to oblongovate,<br />

flattened, minutely villous, 9-20 × 3-5 mm,<br />

with a beak at <strong>the</strong> apex. Seeds 2-3, ovate-reniform,<br />

dark brown to almost black, 3-4 mm long.<br />

Phenology: Flowering <strong>and</strong> fruiting throughout<br />

<strong>the</strong> year.<br />

Status: Probably exotic, naturalized, very<br />

common.<br />

Distribution: Abundant on fences <strong>and</strong> in<br />

weedy places at middle <strong>and</strong> lower elevations. Also<br />

on Mona, Vieques, St. Croix, St. John, St. Thomas,<br />

Tortola, <strong>and</strong> <strong>Virgin</strong> Gorda; generalized in <strong>the</strong><br />

tropics <strong>and</strong> subtropics <strong>of</strong> <strong>the</strong> world. Although this

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