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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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290<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

drying. Seeds rectangular, ca. 4.5 mm long, black,<br />

shiny.<br />

Phenology: Collected in fruit in March.<br />

Status: Native, very rare.<br />

Distribution: Known from a single collection<br />

from Boca de Cangrejos, apparently along <strong>the</strong><br />

s<strong>and</strong>y coast. Predominant along <strong>the</strong> tropical coasts<br />

<strong>of</strong> <strong>the</strong> Old World. In <strong>the</strong> New World, it is known<br />

from a few collections from Panama <strong>and</strong> Brazil.<br />

Commentary: The application <strong>of</strong> <strong>the</strong> name V.<br />

marina is ra<strong>the</strong>r confused, <strong>and</strong> needs an<br />

explanatory note. This epi<strong>the</strong>t was originally<br />

published by Burman based on <strong>the</strong> work <strong>of</strong><br />

Rumphius (Index Alt. Herb. Amboin. 5: 391, tab.<br />

141 fig. 2, 1750). As a result, <strong>the</strong> name Vigna<br />

marina is typified by <strong>the</strong> illustration presented in<br />

that work. Never<strong>the</strong>less, <strong>the</strong> examination <strong>of</strong> this<br />

illustration reveals that in reality it represents <strong>the</strong><br />

species that today bears <strong>the</strong> name V. luteola. Due<br />

to <strong>the</strong> code <strong>of</strong> nomenclature’s principle <strong>of</strong> priority,<br />

<strong>the</strong> name V. luteola should be replaced with V.<br />

marina. Never<strong>the</strong>less, <strong>the</strong> name Vigna luteola<br />

could be maintained if <strong>the</strong> name V. marina was<br />

conserved with a different type, so as to preserve<br />

<strong>the</strong> traditional use <strong>of</strong> both names. In this treatment<br />

I use <strong>the</strong> traditional nomenclature in anticipation<br />

<strong>of</strong> a proposal to conserve <strong>the</strong> name V. marina with<br />

a different type. As a result, V. retusa becomes a<br />

synonym <strong>of</strong> V. marina.<br />

7. Vigna peduncularis (Kunth) Fawcett &<br />

Rendle, Fl. Jam. 4(2): 68. 1920.<br />

Fig. 116. A-D<br />

BASIONYM: Phaseolus peduncularis Kunth<br />

Herbaceous vine, creeping or climbing<br />

twining, attainig 2 m in length. Stems branched<br />

from <strong>the</strong> base, slender, almost cylindrical,<br />

puberulent, usually producing roots in <strong>the</strong> area <strong>of</strong><br />

<strong>the</strong> nodes. Leaves alternate, trifoliolate; leaflets<br />

chartaceous, ovate-deltate or lanceolate, 3-8 × 1.5-<br />

5 cm, both surfaces glabrous, <strong>the</strong> venation slightly<br />

prominent on <strong>the</strong> lower surface, <strong>the</strong> apex acute,<br />

acuminate, or less frequently obtuse, <strong>the</strong> base<br />

cuneate, <strong>the</strong> margins entire, slightly ciliate;<br />

terminal leaflet larger than <strong>the</strong> lateral ones; lateral<br />

leaflets asymmetrical at <strong>the</strong> base, one side obtuse,<br />

<strong>the</strong> o<strong>the</strong>r cuneate or truncate; petiolules ca. 3 mm<br />

long, puberulent; rachis 5-10 mm long, striate;<br />

petioles 2-7 cm long, striate, slender, glabrous;<br />

stipels obovate, ca. 1 mm long; stipules oblong or<br />

lanceolate, 2-2.5 mm long. Pseudoracemes<br />

axillary, up to 26 cm long, <strong>the</strong> flowers 2 per node,<br />

in <strong>the</strong> distal portion <strong>of</strong> <strong>the</strong> inflorescence;<br />

peduncles thick, glabrous or puberulent; pedicels<br />

ca. 1 mm long, erect, with <strong>the</strong> calyx in an<br />

ascending position. Calyx green, campanulate,<br />

puberulent, 2.2-2.5 mm long, <strong>the</strong> sepals more or<br />

less equal, broadly deltate, except for <strong>the</strong> abaxial<br />

one, which is narrower <strong>and</strong> slightly longer; corolla<br />

pale pink, st<strong>and</strong>ard obovate, unguiculate, ca. 8<br />

mm long, <strong>the</strong> wings <strong>and</strong> <strong>the</strong> keel as long as <strong>the</strong><br />

st<strong>and</strong>ard, <strong>the</strong> wings oblong, sinuate, unguiculate,<br />

<strong>the</strong> keel spirally twisted. Legumes ascendent,<br />

linear, flattened, straight, with <strong>the</strong> margins<br />

slightly undulate, 4-5.5 cm long, ca. 3 mm wide,<br />

puberulent, punctate. Seeds oblong, ca. 3.5 mm<br />

long, light brown with blackish spots, <strong>the</strong> hilum<br />

white, ca. 2 mm long.<br />

Phenology: Collected in flower <strong>and</strong> fruit in<br />

November <strong>and</strong> January.<br />

Status: Native, uncommon.<br />

Distribution: In disturbed places like roadsides<br />

<strong>and</strong> pastures. Also in <strong>the</strong> Antilles, Central<br />

America, <strong>and</strong> South America.<br />

Public Forest: Río Abajo.<br />

8. Vigna unguiculata (L.) Walp., Repert. Bot.<br />

Syst. 1: 779. 1842.<br />

Fig. 116. E-F<br />

BASIONYM: Dolichos unguiculatus L.<br />

Frijol, Cowpea<br />

Herbaceous vine, climbing <strong>and</strong> twining,<br />

attainig 3-5 m in length. Stems up to 5 mm in<br />

diameter, cylindrical, glabrous, wine-colored<br />

when young. Leaves alternate, trifoliolate; leaflets<br />

chartaceous, 5-14.6 × 3-10.5 cm, both surfaces<br />

glabrous, with slightly prominent venation, <strong>the</strong><br />

apex obtuse or acute, mucronate, <strong>the</strong> margins<br />

sinuate to lobate; terminal leaflet rhombic-ovate<br />

or rhombic-lanceolate, <strong>the</strong> base cuneate, with <strong>the</strong><br />

point rounded or sometimes subcordiform; lateral<br />

leaflets markedly asymmetrical, one side oblong,<br />

<strong>the</strong> o<strong>the</strong>r ovate or lanceolate, <strong>the</strong> base cuneate or<br />

truncate; petiolules 4-5 mm long, puberulent,<br />

winged; rachis 0.5-3.5 cm long, canaliculate,<br />

marginate or winged; petioles 1-10 cm long,<br />

canaliculate, marginate or winged, glabrous,<br />

thick, broadened at <strong>the</strong> base; stipels oblong or<br />

obovate, ca. 2 mm long; stipules peltate,

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