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Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

Early Cretaceous Freshwater Fish Fauna in Kyushu, Japan

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120 Yoshitaka Yabumoto<br />

Systematic Paleontology<br />

Class Osteichthyes<br />

Subclass Act<strong>in</strong>opterygii<br />

Infraclass Neopterygii<br />

Division Halecostomi<br />

Order Semionotiformes<br />

Family Semionotidae<br />

Genus Lepidotes Agassiz, 1832<br />

Diagnosis. It differs from the other genus, Semionotus, <strong>in</strong> hav<strong>in</strong>g the comb<strong>in</strong>ation of<br />

follow<strong>in</strong>g characters. A median vomer and tritoral dentition are present. Two or<br />

more anamestic suborbitals are present (Schaeffer and Dunkle, 1950; McCune,<br />

1986; Olsen and McCune, 1991). The fossils are found from the Upper Triassic<br />

<strong>in</strong>to the <strong>Cretaceous</strong> <strong>in</strong> Europe, North America, South America, Asia, Africa and<br />

Madagascar.<br />

Type species. Lepidotes gigas Agassiz, 1832.<br />

Lepidotes macropterussp. nov.<br />

(PI. 36)<br />

Diagnosis. It differs from other species of the genus Lepidotes <strong>in</strong> hav<strong>in</strong>g the<br />

comb<strong>in</strong>ation of follow<strong>in</strong>g characters. The dorsal f<strong>in</strong> is situated at about the middle<br />

of the body. The dorsal f<strong>in</strong> base is long and its length is conta<strong>in</strong>ed 1.4 times <strong>in</strong> the<br />

body depth. The pelvic f<strong>in</strong> is below the dorsal f<strong>in</strong>. The number of dorsal f<strong>in</strong> rays is<br />

13 with 4 f<strong>in</strong> fulcra. The number of scales from the dorsal marg<strong>in</strong> to the ventral<br />

marg<strong>in</strong> at the caudal immediately beh<strong>in</strong>d the dorsal f<strong>in</strong> is 11 to 13.<br />

Holotype. KMNH VP 100,146, the middle part of the body. The anterior part<br />

and the posterior part of the body are miss<strong>in</strong>g. The length of the preserved portion<br />

is 78.0 mm along the body axis.<br />

Locality. Tokuriki (TO-1), Kokura-m<strong>in</strong>ami-ku (Kokura Southern Ward), Kitakyu<br />

shu City, Fukuoka Prefecture, <strong>Japan</strong>.<br />

Horizon. The First Formation (the lower formation, W|, correlated to the Sengoku<br />

Formation) of the Wak<strong>in</strong>o Subgroup <strong>in</strong> the Kwanmon Group, the Lower <strong>Cretaceous</strong>.<br />

Etymology. The speciese name, macropterus, means a long or large f<strong>in</strong>, which refers<br />

to the fact that this species has the long dorsal f<strong>in</strong> base.

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