BRYOZOA: CHEILOSTOMATA Interim classification for Treatise ...

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Incertae sedis 1 Pseudostega Superfamily Conescharellinoidea Levinsen, 1909, p. 308 [nom. transl. d’Hondt 1985, pp. 9, 11] YPR-Rec. 27 Family Batoporidae Neviani, 1900, p. 220 (Batoporideae, nom. transl.) YPR-Rec. 2 Batopora, Lacrimula, Family Conescharellinidae Levinsen, 1909, p. 308 YPR-Rec. 7 Bipora, Conescharellina, Crucescharellina, Flabellopora, Ptoboroa, Trochosodon, Zeuglopora Family Lekythoporidae Levinsen, 1909, p. 313 CHT-Rec. 9 Aulopocellidae Vigneaux, 1949, p. 20 Aulopocella, Catadysis, Harpagozoon, Jugescharellina, Lekythopora, Orthoporidra, Poecilopora, Terataulopocella, Turritigera Family Orbituliporidae Canu & Bassler, 1923, p. 186 YPR-Rec. 8 Atactoporidra, Batoporella, Bicupularia, Conescharellinopsis, Orbitulipora, Sphaerulobryozoon, Stichoporina, Xaveropora Incertae sedis 1 Family Cuvillieridae Annoscia, 1965, p. 6 LUT 1 Cuvilliera SUMMARY OF GENERIC/SUBGENERIC DIVERSITY ORDER CHEILOSTOMATA 1064 Suborder Malacostegina 32 Suborder Inovicellina 2 Suborder Scrupariina 5 Suborder Neocheilostomina 1025 Infraorder Flustrina 372 Infraorder Ascophorina 653 “Grade” Acanthostega 179 “Grade” Hippothoomorpha” 32 "Grade” Umbonulomorpha" 138 "Grade” Lepraliomorpha" 304 RATIO OF “ANASCA” (Malacostegina, Inovicellina, Scrupariina, Flustrina) : ASCOPHORINA = 411 : 653 1. The above list is only an ongoing attempt to sort order out of chaos. Consequently you will probably see immediate areas for improvement. Good. Please send me corrections and improvements. Please understand that the classification is evolving and a rigorous scheme is dependent on phylogenetic analysis. 2. One of the benefits of this list is to give an indication of relative diversity of various groupings, e.g., anascans : ascophorans, or proportions of frontal-shield morphologies among the Ascophorina. 3. At first glance it appears there are a disproportionate number of small, even monogeneric, families (especially among the lepralioid ascophorines). Is this a consequence of oversplitting and too much weight accorded to single characters? On the other hand, some families are rather large, with a range of disparate morphologies, and appear to need splitting, e.g., Calloporidae. The Cribrilinidae needs splitting too, there being several major lineages, and 14
some of Lang's family names will be retainable. For the moment it is worth noting that the oldest-available valid family-group name for any of the Cribrilinoidea is the Steginoporidae d'Orbigny, 1853, which Lang for some curious reason chose to ignore while accepting the genus Steginopora (see also Bassler 1953). 4. I plan on expanding the list of generic synonyms to include those already in Bassler. I haven't done this yet. 5. The suggested new families and superfamilies are just that — suggestions. If others agree that these are appropriate, I don't care who formally establishes any of them, the main job is make progress. 6. The suborder Protocheilostomat[in]a Silén, 1942 has been deleted from this compilation. Its sole genus Labiostomella is uncalcified (Silén 1944, pp. 7, 8), has a ctenostomatous mode of reproduction and closing apparatus, and nests with the Benedeniporidae in cladograms. All available evidence therefore indicates that Labiostomella is a ctenostome and not a cheilostome. It needs to be ascertained if Benedeniporidae Delage & Hérouard, 1897 and Labiostomellidae Silén, 1942 are confamilial. Two superfamilies were proposed ⎯⎯ Labiostomellacea (Phaeophanova 1960, p. 155) and Benedeniporoidea (Jebram 1973, p. 34). 7. Priority of taxon names. Article 23.1. [Statement of the Principle of Priority] in the fourth edition of the ICZN (1999) reads: “The valid name of a taxon is the oldest available name applied to it, unless that name has been invalidated or another name is given precedence by any provision of the Code or by any ruling of the Commission.” Note also Article 23.9. [Reversal of precedence] which reverses the Principle of Priority if the senior synonym does not now satisfy two conditions: 1) it has not been used as a valid name after 1899, and 2) the junior synonym has been used in at least 25 works published by at least 10 authors in the immediately preceding 50 years and encompassing a span of not less than 10 years. 8. Note Article 35.5. Precedence for names in use at higher rank: “If after 1999 a name in use for a family-group taxon (e.g. for a subfamily [or superfamily]) is found to be older than a name in prevailing usage for a taxon at higher rank in the same family-group taxon … the older name is not to displace the younger name.” This helpful ruling affects some names of family-group taxa in recent versions of the cheilostome classification scheme that have not yet been formally established through publication. For example, when I proposed the superfamily Calloporoidea (Gordon 1989), I chose not to adopt the Antroporacea of Vigneaux (1949) because the family Antroporidae Vigneaux, 1949 was not in current use, Antropora being traditionally classified with calloporids. In his discussion of the familial relationships of Antropora, Tilbrook (1998) remarked on the heterogeneous composition of the Calloporidae (see also note 3 above) and suggested that adoption of the Antroporidae, inter alia, would ease the problem. He did not comment on the use of Antroporacea (Antroporoidea). The classification given herein uses the Antroporidae following Tilbrook’s suggestion but, following Article 35.5, it is not necessary to overturn Calloporoidea in favour of Antroporoidea. Taxa added 15 September 2011: Arbocuspis, Arbopercula, Osburnea Taxa added 25 August 2011: Favosthimosia, Mourellina, Mourellinidae, Rectimicropora, Schizocoryne Scriblitopora; Crepis transferred to Calloporidae Taxa added 15 December 2010: Corbuliporina, Eutaleola (replaces Euteleia), Uharella Taxon reinstated 29 June 2010: Scorpiodinipora Taxon reinstated 30 April 2010: Rhammatopora 15
Page 1 and 2: BRYOZOA: CHEILOSTOMATA Interim clas
Page 3 and 4: Acanthoporella, Acanthoporidra, Ade
Page 5 and 6: Monoporella Family Macroporidae Utt
Page 7 and 8: Rhabdopora, Rhacheopora, Rhiniopora
Page 9 and 10: Family Inversiulidae Vigneaux, 1949
Page 11 and 12: Family Margarettidae Harmer, 1957,
Page 13: Incertae sedis 1 Bimicroporella Sup

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