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Pleocoma linsleyi Hovore, 1971 (Pleocoma) Pleocoma marquai Hovore, 1972 (Pleocoma) Pleocoma minor Linsley, 1938 (Pleocoma) Pleocoma octopagina Robertson, 1970 (Pleocoma) Pleocoma oregonensis Leach, 1933 (Pleocoma) Pleocoma remota Davis, 1934 Pleocoma puncticollis Rivers, 1889 (Pleocoma) Pleocoma rickseckeri Horn, 1888 (Pleocoma) Pleocoma rubiginosa rubiginosa Hovore, 1972 (Pleocoma) Pleocoma rubiginosa transsierrae Hovore, 1977 (Pleocoma) Pleocoma shastensis Van Dyke, 1933 (Pleocoma) Pleocoma simi Davis, 1934 (Pleocoma) Pleocoma sonomae Linsley, 1935 (Pleocoma) Pleocoma staff Schaufuss, 1870 (Pleocoma) Pleocoma adjuvans Crotch, 1873 Pleocoma edwardsii LeConte, 1874 Pleocoma ulkei Horn, 1888 Pleocoma trifoliata Linsley, 1938 (Pleocoma) Pleocoma tularensis Leach, 1933 (Pleocoma) GEOTRUPIDAE Latreille, 1802 BOLBOCERATINAE Mulsant, 1842 ATHYREINI Howden and Martínez, 1963 Neoathyreus Howden and Martínez, 1963 Neoathyreus fissicornis (Harold, 1880) (Athyreus) Neoathyreus mixtus Howden, 1964 (Neoathyreus) BOLBOCERATINI Mulsant, 1842 Bolbelasmus Boucomont, 1911 Kolbeus Boucomont, 1911 Bolbelasmus arcuatus (Bates, 1887) (Bolboceras) Bolbelasmus bajaensis Howden, 1964 (Bolbelasmus) Bolbelasmus hornii (Rivers, 1886) (Bradycinetus) Bolbelasmus minor (Linell, 1896) (Bradycinetus) Bolbelasmus variabilis Howden, 1964 (Bolbelasmus) Bolbocerastes Cartwright, 1953 Bolbocerastes imperialis imperialis Cartwright, 1953 (Bolbocerastes) Bolbocerastes imperialis kansanus Cartwright, 1953 (Bolbocerastes) Bolbocerastes peninsularis Schaeffer, 1906 (Bradycinetus) Bolbocerastes regalis Cartwright, 1953 (Bolbocerastes) 2
Bolbocerastes serratus (LeConte, 1854) (Athyreus) Bolbocerosoma Schaeffer, 1906 Bolbocerosoma (Bolbocerosoma) Schaeffer, 1906 Bolbocerosoma (Bolbocerosoma) biplagiatum Dawson and McColloch, 1924 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) bruneri Dawson and McColloch, 1924 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) cartwrighti Howden, 1955 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) confusum Brown, 1928 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) elongatum Howden, 1955 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) farctum (Fabricius, 1775) (Scarabaeus) Scarabaeus cephus Fabricius, 1775 Bolbocerosoma (Bolbocerosoma) hamatum Brown, 1929 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) lepidissimum Brown, 1928 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) mexicanum Howden, 2005 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) pusillum pusillum Dawson and McColloch, 1924 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) pusillum townesi Howden, 1955 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) quadricornum Robinson, 1941 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) ritcheri Howden, 1955 (Bolbocerosoma) Bolbocerosoma (Bolbocerosoma) tumefactum (Palisot de Beauvois, 1809) (Scarabaeus) Bolborhombus Cartwright, 1953 Bolborhombus angulus (Robinson, 1947) (Bolboceras) Bolborhombus magnus Howden, 1964 (Bolborhombus) Bolborhombus parvulus Cartwright, 1953 (Bolborhombus) Bolborhombus sallaei (Bates, 1887) (Bolboceras) Bradycinetus carinatus Schaeffer, 1906 Bolboceras schaefferi Boucomont, 1911 Bradycinetulus Cockerell, 1906 Amechanus Horn, 1870 Bradycinetus Horn, 1871 Bradycinetulus ferrugineus (Palisot de Beauvois, 1809) (Scarabaeus) Bolboceras furcicollis Laporte, 1840 Bradycinetulus fossatus (Haldeman, 1853) (Bolboceras) Bradycinetulus rex Cartwright, 1953 (Bradycinetulus) Eucanthus Westwood, 1848 Eucanthus alutaceus Cartwright, 1944 (Eucanthus) Eucanthus greeni Robinson, 1948 (Eucanthus) Eucanthus impressus Howden, 1964 (Eucanthus) Eucanthus lazarus (Fabricius, 1775) (Scarabaeus) 3
Page 1: Checklist and Nomenclatural Authori
Page 5 and 6: Geotrupes (Anoplotrupes) hornii Bla
Page 7 and 8: Passalus punctiger LePeletier and S
Page 9 and 10: Trox triestinae Pittino, 1987 Omorg
Page 11 and 12: Nicagus LeConte, 1861 Nicagus occul
Page 13 and 14: Codocera Eschscholtz, 1821 Stomphax
Page 15 and 16: Lichnanthe Burmeister, 1844 Dasyder
Page 17 and 18: Agrilinellus nuriae Dellacasa, Dell
Page 19 and 20: Aphodius retusus Waltl, 1835 Aphodi
Page 21 and 22: Coelotrachelus kuntzeni (Schmidt, 1
Page 23 and 24: Dialytes striatulus (Say, 1825) (Tr
Page 25 and 26: Lechorodius Gordon and Skelley, 200
Page 27 and 28: Oscarinus silvanicus (Cartwright, 1
Page 29 and 30: Scabrostomus bakeri (Skelley and Go
Page 31 and 32: Trichonotus setiger Mulsant, 1842 A
Page 33 and 34: Ataenius hirsutus Horn, 1871 (Ataen
Page 35 and 36: Sayloria Chalumeau, 1981 Haroldiata
Page 37 and 38: Platytomus caelicollis (Cartwright,
Page 39 and 40: Canthon (Canthon) pilularius (Linna
Page 41 and 42: Dichotomius Hope, 1838 Homocopris B
Page 43 and 44: Copris metallicus Fabricius, 1798 O
Page 45 and 46: Phanaeus (Phanaeus) adonis Harold,
Page 47 and 48: Serica aemula Dawson, 1947 (Serica)
Page 49 and 50: Serica errans Blatchley, 1929 Seric
Page 51 and 52: Hypothyce osburni (Cartwright, 1967
Page 53 and 54:
Lachnosterna generosa Horn, 1887 Ph
Page 55 and 56:
Phyllophaga (Phyllophaga) glabricul
Page 57 and 58:
Phyllophaga (Phytalus) obsoleta (Bl
Page 59 and 60:
Phyllophaga (Phyllophaga) squamipil
Page 61 and 62:
Polyphylla alleni Cazier, 1939 Poly
Page 63 and 64:
Diplotaxis brevicollis LeConte, 185
Page 65 and 66:
Diplotaxis mima Vaurie and Cazier,
Page 67 and 68:
Diplotaxis morula LeConte, 1856 Dip
Page 69 and 70:
Dichelonyx testacea Kirby, 1837 Dic
Page 71 and 72:
Hoplia (Hoplia) dispar LeConte, 188
Page 73 and 74:
Plusiotina subenodis Casey, 1915 Ch
Page 75 and 76:
Pelidnota pallidipes Casey, 1915 Pe
Page 77 and 78:
Anomala exigua (Schwarz, 1878) (Str
Page 79 and 80:
Strigoderma floridana Ohaus, 1905 S
Page 81 and 82:
Spilosota magister Casey, 1915 Spil
Page 83 and 84:
Aphonus brevicruris Cartwright, 194
Page 85 and 86:
Ligyrus farctus Casey, 1915 Ligyrus
Page 87 and 88:
Xyloryctes thestalus borealis Endr
Page 89 and 90:
Apeltastes Howden, 1968 Apeltastes
Page 91 and 92:
Myrmeceicon Mann, 1914 Cremastochei
Page 93 and 94:
Euphorhipis Casey, 1915 Euphoriaspi
Page 95 and 96:
Cetonia fictilis Newman, 1838 Prota
Page 97:
Gymnetis aequalis Casey, 1915 Gymne
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