scale insects and mealy bugs - Zoological Survey of India

SCALE INSECTS AND MEALY BUGS 

(INSECTA: HOMOPTERA: COCCOIDEA) 

R.K.VARSHNEY*, M.J. JADHAV AND R.M.SHARMA 

Zoological Survey of India, Western Regional Centre, Pune-411044 

*Raj Bhawan, Manik Chowk, Aligarh-202001 

The Scale insects and mealy bugs fall under the Superfamily Coccoidea, Order 

Homoptera. The name refers to their secretion of waxy scales or mealy that serves as a protective 

covering. They range in size from 1.5 mm to 25 mm in length. They are economically important 

insects, since many species attack different parts of plant and feed by sucking the plant sap. On 

the other hand a few species of scale insects are beneficial to mankind, e.g. the cochineal insect 

(Dactylopius indicus) yields a carmine dye and the lac insect (Kerria lacca) is the source of 

shellac. They are found on nearly all parts of the host plant including the leaves, branches, 

trunks, fruits, and roots. They sometimes occur under bark and may cause various kinds of plant 

deformities including chlorotic spots, pits, and galls. They cause severe damage to economically 

important plants like sugarcane, tea, coffee, cotton, sorghum, vegetables, greenhouse plants and 

almost all kinds of fruit trees. 

About 7,500 species of Scale insects and Mealy bugs are known at present from the 

world which inhabits almost all parts of the globe, in all floral habitats, from the tundra to the 

tropics. The Indian diversity accounts to 409 species under 166 genera spread over 14 families 

and 14 subfamilies (Varshney; 1992, 2002) and other scattered literature. 

1. Genus: Kuwania Cockerell in Fernald, 1903 

SYSTEMATIC ACCOUNT 

Superfamily: Coccoidea 

Family: Margarodidae 

Subfamily: Margarodinae 

Tribe: Kuwanini 

1. Kuwania zeylanica (Green) Fernald, 1903 

Distribution: Kerala (Calicut). 

Tribe: Margarodini 

1

2. Genus: Margarodes Guilding, 1829 

2. Margarodes formicarum Guilding, 1829 

Distribution: South India. 

3. Margarodes indicus Green, 1912 

Distribution: Mumbai and Bengaluru. 

4. Margarodes papillosus Green, 1912 

Distribution: Karnataka (Bengaluru, Bellary). 

3. Genus: Neomargarodes Green, 1914 

5. Neomargarodes niger Morrison, 1928 

Subfamily: Neomargarodinae 

Distribution: Karnataka (Mysore, Bellary), Tamil Nadu (Coimbatore). 

Family: Monophlebidae 

Subfamily: Monophlebinae 

Tribe: Iceryini 

4. Genus: Crypticerya Cockerell, 1895 

Crypticerya sp. 

Distribution: Odisha (Bhubaneshwar, Taptapani, Talcher and Angul). 

6. Crypticerya jacobsoni Morrison, 1928 

Distribution: Assam (Dibrugarh, Jokai Forest), Karnataka (Bengaluru), Kerala (Cochin). 

7. Crypticerya jaihind Rao, 1951 

Distribution: Karnataka (Kanara Dist., Mangalore), Kerala (Cochin, Nelliampathy hills, 

Travancore), Tamil Nadu (Nilambur). 

8. Crypticerya kumari Rao, 1951 

Distribution: Karnataka (Gopalswami hills). 

5. Genus: Icerya Signoret, 1875 

Icerya sp. 

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Distribution: Odisha (Bhubaneshwar, Taptapani, Dhauli, Konark, Rambha, Gopalpur-on-sea, 

Kuliana and Baripada). 

9. Icerya aegyptiaca (Douglas, 1890) 

Distribution: Bihar, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, Tripura, West Bengal. 

10. Icerya formicarum Newstead, 1897 

Distribution: Andhra Pradesh, Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, 

Tamil Nadu, Uttar Pradesh, West Bengal and North-East India. 

11. Icerya menoni Rao, 1951 

Distribution: Kerala (Maradu). 

12. Icerya minor Green, 1908 

Distribution: Assam (Haflong), Bihar (Pusa), Uttar Pradesh (Varanasi). 

13. Icerya pilosa Green, 1896 

Distribution: Andhra Pradesh (Sarola), Bihar (Shahabad, Juneshwargarh farm), Delhi, Haryana 

(Karnal), Madhya Pradesh (Gwalior), Kerala (Perungaloor, Travancore), Tamil Nadu (Chennai, 

Hasanoor, Coimbatore, Kanyakumari, Satyamangalam, Coromandel coast), Uttar Pradesh 

(Aligarh, Meerut, Shahjahanpur) and West Bengal. 

14. Icerya pulcher Morrison, 1928 

Distribution: Bihar (Pusa). 

15. Icerya purchase Maskell, 19879 

Distribution: Karnataka, Kerala, Maharashtra, Odisha, and Tamil Nadu. 

16. Icerya seychellarum (Westwood, 1855) 

Distribution: Andaman Is., Karnataka (Bellary, Kottur), Kerala, Madhya Pradesh, Maharashtra, 

Odisha, and Tamil Nadu. 

17. Icerya travancorensis Rao, 1951 

Distribution: Kerala, Tamil Nadu. 

18. Icerya zimmermani Green, 1932 

Distribution: Bihar (Bhagalpur). 

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6. Genus: Steatococcus Ferris, 1921 

19. Steatococcus assamensis Rao, 1951 

Distribution: Meghalaya (Dawki). 

7. Genus: Drosicha Walker, 1858 

Drosicha sp. 

Tribe: Drosichini 

Distribution: Tamil Nadu (Chennai), Bihar (Pusa). 

20. Drosicha burmeisteri Cockerell, 1902 

Distribution: North India. 

21. Drosicha contrahens Walker, 1858 

Distribution: North-East India. 

22. Drosicha dalbergiae (Green, 1902) 

Distribution: Bihar, Punjab (Sutlej Valley) and Uttarakhand (Dehra Dun). 

23. Drosicha leachii (Westwood, 1845) 

Distribution: Kerala, Uttarakhand (Dehra Dun). 

24. Drosicha mangiferae (Green, 1903) 

Distribution: Bihar (Bhagalpur, Pusa, Sitamarhi, Dalsingsarai, Santhal Paraganas, Darbhanga, 

Muzaffarpur), Delhi, Uttar Pradesh (Fatehpur), Uttarakhand (Dehra Dun). 

25. Drosicha quadricaudata (Green, 1922) 

Distribution: Uttar Pradesh (Varanasi). 

26. Drosicha saundersii (Westwood) Foldi, 2001 


27. Drosicha stebbingii (Green); Morrison, 1928 

Distribution: Bihar (Darbhanga, Saran, Muzaffarpur), Haryana (Karnal), Himachal Pradesh 

(Shimla), Madhya Pradesh, Rajasthan (Jodhpur), Uttarakhand (Dehra Dun), Uttar Pradesh 

(Saharanpur, Kanpur, Varanasi), West Bengal (Howrah & Darjeeling Dist.) 

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28. Drosicha tectonae (Green); Morrison, 1928 

Distribution: Madhya Pradesh (Damoh), Punjab, Uttar Pradesh. 

8. Genus: Aspidoproctus Newstead, 1901 

Tribe: Monophlebini 

29. Aspidoproctus pertinax Newstead, 1901 

Distribution: Karnataka (Bengaluru). 

9. Genus: Hemaspidoproctus Morrison, 1927 

30. Hemaspidoproctus cinereus Morrison, 1927 

Distribution: Bihar (Pusa), Karnataka, Kerala, Gujarat, Rajasthan and Tamil Nadu. 

10. Genus: Labioproctus Green, 1922 

31. Labioproctus poleii Green, 1896 

Distribution: Karnataka (Mysore, Coorg), West Bengal (Darjeeling Dist.). 

11. Genus: Misracoccus Rao, 1950 

32. Misracoccus assamensis Rao, 1950 

Distribution: Meghalaya (Shilong). 

33. Misracoccus xyliae Rao, 1950 

Distribution: Karnataka (Molakalmurn), Kerala (Malabar) and Tamil Nadu (Coimbatore). 

12. Genus: Monophlebus Guerin-Meneville, 1827 

Monophlebus sp. 

Distribution: Tamil Nadu. 

34. Monophlebus atripennis Burmeister, 1835 


13. Genus: Monophlebidus Morrison, 1927 

35. Monophlebidus indicus Morrison, 1927 

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14. Genus: Perissopneumon Newstead, 1900 

36. Perissopneumon ferox Newstead, 1900 

Distribution: Kerala, Maharashtra, Tamil Nadu, Uttar Pradesh. 

37. Perissopneumon phyllanthi (Green, 1923) 

Distribution: Andhra Pradesh (Godavari Dist.), Karnataka (Bengaluru), Kerala (Malabar, 

Walayar Forest) and West Bengal (Kolkata). 

38. Perissopneumon tamarinda Rao, 1950 

Distribution: Andhra Pradesh (Godavari Dist.) and West Bengal (Kolkata). 

15. Genus: Walkeriana Signoret, 1876 

39. Walkeriana senex Green, 1896 

Distribution: Kerala (Wynaad) and Tamil Nadu. 

16. Genus: Orthezia Bosc d’ Antic, 1784 

40. Orthezia insignis Browne, 1887 

Family: Ortheziidae 

Distribution: Karnataka (Bengaluru), Kerala (Wynad), Tamil Nadu (Salem Dist.), Sikkim 

(Rangpo) and West Bengal. 

17. Genus: Kerria Targioni-Tozzetti, 1884 

Kerria sp. Varshney, 2000 

Family: Tachardiidae 

Subfamily: Tachardiinae 

Tribe: Tachardini 

Subgenus: (i) Kerria 

Distribution: Odisha (Bhubaneshwar) and Tamil Nadu (Jawalgiri). 

41. Kerria (K.) albizziae (Green, 1911) 

Distribution: Bihar, Uttarakhand (Dehra Dun), West Bengal. 

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42. Kerria (K.) brancheata Varshney, 1966 

Distribution: Chhatisgarh, Jharkhand. 

43. Kerria (K.) chamberlini Varshney, 1966 

Distribution: Rajasthan (Jodhpur). 

44. Kerria (K.) chinensis chinensis (Mahdihassan, 1923) 

Distribution: North- East India. 

45. Kerria (K.) chinensis kydia (Misra, 1930) 

Distribution: Assam (Cachar Dist.). 

46. Kerria (K.) communis (Mahdihassan, 1923) 

Distribution: Andhra Pradesh, Goa, Karnataka, Kerala, Maharashtra, Tamil Nadu. 

47. Kerria (K.) ebrachiata (Chamberlin, 1923) 

Distribution: Chhatisgarh, Jharkhand, Karnataka (Bengaluru). 

48. Kerria (K.) fici fici (Green, 1903) 

Distribution: Bihar (Patna), Delhi, Madhya Pradesh, Uttar Pradesh, Jammu & Kashmir, 

Rajasthan, Tamil Nadu (Coimbatore), West Bengal. 

49. Kerria (K.) fici jhasiensis (Misra, 1930) 

Distribution: Uttar Pradesh (Jhansi). 

50. Kerria (K.) indicola (Kapur, 1958) 

Distribution: Bihar. 

51. Kerria (K.) lacca lacca (Kerr, 1782) 

Distribution: Andhra Pradesh, Bihar, Karnataka, Odisha, Tamil Nadu, Uttar Pradesh and West 

Bengal. 

52. Kerria (K.) lacca ambigua (Misra, 1930) 

Distribution: Uttar Pradesh (Guna, Jhansi). 

53. Kerria (K.) lacca mysorensis (Mahadihassan, 1923) 

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Distribution: Karnataka (Bengaluru, Mysore). 

54. Kerria (K.) nagoliensis (Mahadihassan, 1923) 

Distribution: Madhya Pradesh (Raipur, Sohagpur). 

55. Kerria (K.) nepalensis Varshney, 1977 


56. Kerria (K.) pusana (Misra, 1930) 


57. Kerria (K.) laca mysorensis (Mahadihassan, 1923) 

Distribution: Karnataka. 

58. Kerria (K.) sharda Misra & Sushil, 2000 rangoonensis (Chamberlin, 1925) 

Distribution: Odisha. 

Subgenus: (ii) Chamberliniella Varshney, 1987 

59. Kerria (C.) rangoonensis (Chamberlin, 1925) 

Distribution: Assam. 

18. Genus: Paratachardina Balachowsky, 1950 

Subfamily: Tachardininae 

60. Paratachardina mahdihassani (Kondo & Gullan, 2007) 


61. Paratachardina mithila Varshney, 1968 

Distribution: Meghalaya (Shillong). 

62. Paratachardina silvestri (Mahdihassan, 1923) 

Distribution: Andhra Pradesh, Karnataka (Bengaluru) and Tamil Nadu. 

63. Paratachardina ternate (Chamberlin, 1923) 

Distribution: Kerala (Travancore). 

64. Paratachardina theae (Green) in Green & Mann, 1907 

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Distribution: Assam, Sikkim and West Bengal. 

19. Genus: Eriococcus Targioni-Tozzetti, 1869 

65. Eriococcus araucariae Maskell, 1879 

Family: Eriococcidae 

Distribution: Karnataka (Bengaluru) and Tamil Nadu. 

66. Eriococcus lagerstroemiae Kuwana, 1907 


67. Eriococcus phyllanthi Ferris, 1957 


68. Eriococcus rhodomyrti Green, 1922 

Distribution: Maharashtra. 

69. Eriococcus transwersus Green, 1922 

Distribution: India 

70. Eriococcus beesoni Green, 1926 

Distribution: Odisha (Ganjam Dist.) and Tamil Nadu 

20. Genus: Kermes Boitard, 1828 

71. Kermes himalayensis Greee, 1909 

Distribution: North-West India. 

21. Genus: Pseudopulvinaria Atkinson, 1889 

Family: Kermesidae 

72. Pseudopulvinaria sikkimensis Atkinson, 1889 

Distribution: Assam, Meghalaya, Sikkim, Shillong. 

22. Genus: Dactylopius Costa, 1835 

Family: Dactylopiidae 

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Dactylopius sp. 

Distribution: India: Odisha (Bhubaneswar). 

73. Dactylopius coccus Costa, 1835 

Distribution: India. 

74. Dactylopius indicus Green, 1922 

Distribution: Andhra Pradesh (Godavari Dist.), Bihar (Patna, Gaya), Himachal Pradesh, Odisha, 

Tamil Nadu and West Bengal. 

75. Dactylopius tomentosus (Lamarck, 1801) 

Distribution: Odisha, Karnataka, Maharashtra, Tamil Nadu and West Bengal. 

23. Genus: Birendracoccus Ali, 1975 

Family: Pseudococidae 

Sub family: Pseudococcinae 

Tribe: Phenacoccinini 

76. Birendracoccus saccharifolii (Green, 1908) 

Distribution: Andhra Pradesh, Bihar, Delhi, Uttar Pradesh (Aligarh, Meerut) and West Bengal. 

24. Genus Brevennia Goux, 1940 

77. Brevennia rehi (Lindinger, 1943) 

Distribution: Andhra Pradesh, Bihar, Delhi, Karnataka, Kerala, Madhya Pradesh, Maharashtra, 

Odisha, Tamil Nadu and West Bengal. 

25. Genus: Coccidohystrix Lindinger, 1943 

Coccidohystrix sp. 

Distribution: Punjab (Hoshiarpur). 

78. Coccidohystrix insolita (Green, 1908) 

Distribution: Andhra Pradesh, Bihar, Gujarat, Goa, Kerala, Haryana, Tamil Nadu and West 

Bengal. 

26. Genus: Helicoccus Sulc, 1912 

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79. Helicoccus singularis Avasthi & Shafee, 1982 

Distribution: Andhra Pradesh. 

27. Genus: Naiacoccus Green, 1919 

80. Naiacoccus serpentines Green, 1919 

Distribution: Punjab and Uttar Pradesh. 

28. Genus: Nipacoccus Sulc, 1945 

Nipacoccus sp. 

Distribution: Punjab, Karnataka and Odisha. 

81. Nipacoccus viridis (Newstead, 1894) 

Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Himachal Pradesh, Maharashtra, Kerala, 

Karnataka, Uttar Pradesh and West Bengal. 

29. Genus: Novonilacoccus Ghosh & Ghose, 1987 

82. Novonilacoccus oryzae Ghosh & Ghose, 1987 

Distribution: West Bengal (Kalyani). 

30. Genus: Paraputo Laing, 1929 

83. Paraputo taraktogeni Rao, 1950 

Distribution: Assam (Dibrugarh). 

31. Genus: Phenacoccus Cockerell, 1893 

Phenacoccus sp. 

Distribution: Kashmir and Uttarkhand (Dehra Dun). 

84. Phenacoccus indicus (Avashti & Shafee, 1980) 

Distribution: Karnataka (Mysore, Bengaluru). 

85. Phenacoccus quaternus (Green, 1912) 

Distribution: Tamil Nadu (Coromandel Coast). 

32. Genus: Rastrococcus Ferris, 1954 

11

Rastrococcus sp. 

Distribution: Bihar (Patna). 

86. Rastrococcus iceryoides (Green, 1908) 

Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Himachal Pradesh, Maharashtra, Kerala, 

Karnataka, Uttar Pradesh and West Bengal. 

87. Rastrococcus invadens Williams, 1986 


88. Rastrococcus mangiferae Ferris, 1954 

Distribution: Andhra Pradesh, Karnataka, Odisha and Tamil Nadu. 

89. Rastrococcus ornatus Ferris, 1954 


90. Rastrococcus spinosus (Robinson, 1918) 

Distribution: Odisha (Bhubaneswar). 

91. Rastrococcus resinophila Green, 1916 

Distribution: Himachal Pradesh, Jammu & Kashmir and Uttar Pradesh. 

33. Genus: Adolesoma Borchsenius, 1948 

Tribe: Pseudococcinini 

92. Adolesoma phragmitidis Borchsenius, 1948 

Distribution: West Bengal. 

34. Genus: Cataenococcus Ferris, 1955 

Cataenococcus sp. 

Distribution: Andaman Is. 

93. Cataenococcus theaecola Das & Ganguli, 1962 

Distribution: Assam and West Bengal. 

35. Genus: Chorizococcus McKenzie, 1960 

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94. Chorizococcus sorghi Williams, 1970 

Distribution: Madhya Pradesh. 

36. Genus: Crisicococcus Ferris, 1950 

95. Crisicococcus dischidae Ezzat & McConnell, 1959 

Distribution: Tripura. 

37. Genus: Dysmicoccus Ferris, 1950 

96. Dysmicoccus breviceps (Cockerell, 1893) 

Distribution: Andhra Pradesh, Karnataka, Kerala, Odisha, Tripura and West Bengal. 

97. Dysmicoccus carens Williams, 1970 


98. Dysmicoccus cocotis (Maskell, 1890) 

Distribution: Kerala. 

99. Dysmicoccus cucurbitae Avashti & Shafee, 1986 

Distribution: Uttar Pradesh. 

100. Dysmicoccus triadus Williams, 1987 


101. Dysmicoccus williamsi Avashti & Shafee, 1981 


38. Genus: Ferrisia Fullaway, 1923 

Ferrisia sp. 

Distribution: Odisha (Bhubaneswar, Puri). 

102. Ferrisia virgata (Cockerell, 1893) 

Distribution: Andhra Pradesh, Bihar, Goa, Maharashtra, Karnataka, Kerala, Madhya Pradesh, 

Tamil Nadu, West Bengal and Uttar Pradesh. 

39. Genus: Indococcus Ali, 1968 

13

Indococcus sp. 


103. Indococcus pipalae Ali, 1968 

Distribution: Bihar and Odisha. 

40. Genus: Kermicus Newstead, 1897 

104. Kermicus wroughtoni Newstead, 1897 


41. Genus: Kiritshenkella Borchsenius, 1948 

105. Kiritshenkella sacchari (Green, 1900) 

Distribution: Andhra Pradesh, Bihar, Assam, Maharashtra, Karnataka, Kerala, Madhya Pradesh, 


42. Genus: Maconellicoccus Ezzat, 1958 

106. Maconellicoccus hirsutus Green , 1908 

Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Maharashtra, Karnataka, Kerala, Madhya 

Pradesh, Tamil Nadu, Uttar Pradesh. 

107. Maconellicoccus perforates (De Lotto 1954) 


43. Genus: Paracoccus Ezzat & McConnell, 1956 

108. Paracoccus burnerae Ezzat & McConnell, 1956 


109. Paracoccus nellorensis Avashti & Shafee, 1984 


44. Genus: Planococcoides Ezzat & McConnell, 1956 

110. Planococcoides bengalensis Ghosh & Ghose, 1988 


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111. Planococcoides robustus Ezzat & McConnell, 1956 

Distribution: Andhra Pradesh, Bihar, Gujarat, Karnataka, Kerala, Maharashtra and Uttar Pradesh. 

45. Genus: Planococcus Ferris, 1950 

Planococcus sp. 


112. Planococcus citri Ferris, 1950 

Distribution: Karnataka, Kerala and Tamil Nadu. 

113. Planococcus ficus Fernald, 1903 


114. Planococcus indicus Avashti & Shafee, 1987 

Distribution: Kerala (Ernaculum) and Tamil Nadu (Coimbatore: Botanical Garden) 

115. Planococcus lilacinus (Cockerell, 1905) 

Distribution: Andhra Pradesh, Karnataka, Kerala, Odisha and Tamil Nadu. 

46. Genus: Pseudococcus Westwood, 1840 

Pseudococcus sp. 

Distribution: Andhra Pradesh, Maharashtra and Odisha. 

116. Pseudococcus formiceticola (Newstead, 1900) 

Distribution: Maharashtra (Konkan). 

117. Pseudococcus longispinus (Targioni-Tozzetti, 1867) 

Distribution: Andhra Pradesh, Karnataka, Kerala, Tamil Nadu and West Bengal. 

118. Pseudococcus saccharicola Takahashi, 1928 

Distribution: Tamil Nadu (Coimbatore). 

47. Genus: Saccharococcus Ferris, 1950 

119. Saccharicoccus sacchari (Cockerell, 1895) 

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Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Haryana, Maharashtra, Karnataka, Kerala, 

Madhya Pradesh, Odisha, Tamil Nadu and Uttar Pradesh. 

48. Genus: Trionymus Berg, 1899 

120. Trionymus ceres Williams, 1970 

Distribution: Madhya Pradesh. 

49. Genus: Geococcus Green, 1902 

121. Geococcus citrinus Kuwana, 1923 

Distribution: Tamil Nadu (North Arcot Dist.). 

122. Geococcus coffeae Green, 1933 


Tribe: Rhizoecini 

50. Genus: Rhizoecus Kunckel d’Herculais, 1878 

123. Rhizoecus amorphophalli Betram, 1940 


124. Rhizoecus cocois Williams, 1985 

Distribution: Kerala (Thiruvananthapuram Dist.). 

125. Rhizoecus cynodontis Green, 1931 

Distribution: Andhra Pradesh (Godavari Dist.) and Tamil Nadu (Coimbatore). 

51. Genus: Xenococcus Silvestri, 1924 

126. Xenococcus annandalei Silvestri, 1924 

Distribution: Andhra Pradesh, Karnataka, Odisha (Berkuda Is. In Chilka lake) and Tamil Nadu. 

127. Xenococcus scorpioides De Lotto, 1977 


52. Genus: Antonina Signoret, 1875 

Tribe: Sphaerococcini 

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128. Antonina anceps Green, 1914 

Distribution: Maharashtra (Probably Pune). 

129. Antonina graminis (Maskell, 1897) 

Distribution: Bihar, Delhi, Karnataka, Odisha, Uttar Pradesh and Tamil Nadu. 

130. Antonina maritime Green, 1922 

Distribution: Tamil Nadu (Coimbatore) and West Bengal. 

131. Antonina zonata Green, 1919 

Distribution: Bihar and Tamil Nadu (Coimbatore). 

53. Genus: Chaetococcus Maskell, 1898 

132. Chaetococcus bambusae (Maskell, 1893) 


54. Genus: Aclerda Signoret, 1874 

Aclerda sp. 

Distribution: Odisha (Bhubaneswar, Taptapani). 

133. Aclerda distorta Green, 1909 

Family: Aclerdidae 


134. Aclerda inermis Ayyar, 1936 


135. Aclerda ischaemum Ayyar, 1930 

Distribution: Tamil Nadu (Coimbatore) and Uttar Pradesh. 

136. Aclerda tokionis (Cockerell, 1896) 

Distribution: Bihar, Odisha and Tamil Nadu (Salem). 

Family: Asterolecaniidae 

Subfamily: Asterolecaniinae 

17

55. Genus: Asterolecanium Targioni-Tozzetti, 1868 

Asterolecanium sp. 

Distribution: Tripura (Agartala) and West Bengal. 

137. Asterolecanium longum Green, 1909 


138. Asterolecanium minutum Takahashi, 1930 


139. Asterolecanium pseudomilliaris Green, 1922 

Distribution: Assam and West Bengal. 

140. Asterolecanium tenuissimum Green, 1909 

Distribution: Kerala (Malabar, Dhoney valley). 

56. Genus: Bambusaspis Cockerell, 1902 

141. Bambusaspis bambusae (Boisduval, 1869) 


57. Genus: Asterococcus Borchsenius, 1960 

Family: Cerococcidae 

142. Asterococcus ramakrishnai Lambdin, 1983 

Distribution: Tamil Nadu (Coimbatore), Maharashtra (Mumbai). 

58. Genus: Cerococcus Comstock, 1882 

Cerococcus sp. 

Distribution: Uttar Pradesh (Aligarh). 

143. Cerococcus bryoides (Maskell, 1894) 

Distribution: Tamil Nadu (Chennai). 

144. Cerococcus indicus (Maskell, 1897) 

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Distribution: Andhra Pradesh, Bihar, Karnataka, Maharashtra (Mumbai), Odisha and West 

Bengal. 

145. Cerococcus ornatus Green, 1909 


59. Genus: Anomalococcus Green, 1902 

146. Anomalococcus indicus Ayyar, 1930 

Family: Lecanodiaspididae 

Distribution: Karnataka, Odisha and Tamil Nadu. 

60. Genus: Cosmococcus Borchsenius, 1959 

147. Cosmococcus erythrinae Borchsenius, 1959 


61. Genus: Stictacanthus Lambdin & Kosztarab, 1973 

148. Stictacanthus azadirachtae Green, 1909 

Distribution: South India and Uttar Pradesh. 

62. Genus: Ceroplastes Gray, 1928 

Ceroplastes sp. 

Distribution: Delhi, Odisha and Tripura. 

149. Ceroplastes actiniformes Green, 1896 

Family: Coccidae 

Subfamily: Ceroplastinae 

Distribution: Andhra Pradesh, Bihar, Maharashtra, Odisha, Uttar Pradesh, Tamil Nadu and West 

Bengal 

150. Ceroplastes ajmerensis (Avasthi & Shafee, 1979) 

Distribution: Rajasthan (Ajmer). 

151. Ceroplastes alami Avasthi & Shafee, 1986 

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Distribution: Uttar Pradesh (Aligarh) and Tamil Nadu (Coimbatore). 

152. Ceroplastes ceriferus (Fabricius, 1798) 

Distribution: Bihar, Uttar Pradesh, Himachal Pradesh, Maharashtra (Mumbai), Madhya Pradesh, 

Tamil Nadu (Coimbatore, Tanjore) and West Bengal. 

153. Ceroplastes floridensis Comstock, 1881 

Distribution: Assam, Bihar, Uttar Pradesh, Himachal Pradesh, Karnataka, Maharashtra 

(Mumbai), Madhya Pradesh, Odisha, Tamil Nadu (Coimbatore, Tanjore) and West Bengal. 

154. Ceroplastes pseudoceriferus (Fabricius, 1798) 

Distribution: Bihar, Delhi, Odisha and Uttar Pradesh. 

155. Ceroplastes rubens Maskell, 1893 

Distribution: Andhra Pradesh (Chicacole), Assam, Kerala, Tamil Nadu (Coimbatore) and West 

Bengal. 

156. Ceroplastes rusci Signoret, 1972 

Distribution: Uttarakhand (Dehra Dun). 

63. Genus: Vinsonia Signoret, 1972 

157. Vinsonia stellifera Westwood, 1871 

Distribution: Andhra Pradesh, Karnataka, Kerala, Uttar Pradesh, Maharashtra (Mumbai), Odisha, 


64. Genus: Akermes Cockerell, 1902 

158. Akermes montanus (Green, 1908) 

Distribution: Himachal Pradesh. 

65. Genus: Ceronema Maskell, 1895 

159. Ceronema koebeli Green, 1909 


Subfamily: Coccinae 

Tribe: Coccini 

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66. Genus: Coccus Linnaeus, 1758 

Coccus spp. 

Distribution: Andhra Pradesh, Karnataka, Tripura, Uttar Pradesh and Odisha. 

160. Coccus acutissimus Green, 1896 

Distribution: Kerala (Malabar) and Tamil Nadu (Coimbatore). 

161. Coccus adersi (Newstead, 1917), Ali, 1971 


162. Coccus almoraensis Avasthi & Shafee, 1984 

Distribution: Bihar and Uttar Pradesh (Almora). 

163. Coccus discrepans Green, 1904 

Distribution: Andhra Pradesh, Assam, Bihar and Kerala. 

164. Coccus gymnospori Sanders, 1909 

Distribution: Andhra Pradesh and Maharashtra (Pune). 

165. Coccus hesperidium Linnaeus, 1758 

Distribution: Andhra Pradesh, Assam, Bihar, Goa, Gujarat, Karnataka, Kerala, Odisha, Jammu & 

Kashmir, Tripura and West Bengal. 

166. Coccus kosztarabi Avasthi & Shafee, 1984 


167. Coccus latioperculatum Green, 1922 


168. Coccus longulus (Douglas, 1887) 

Distribution: Andhra Pradesh, Assam, Karnataka and Odisha. 

169. Coccus ophiorrhizae Green, 1896 

Distribution: Andhra Pradesh (Kurnool). 

170. Coccus ramakrishnae (Ayyar, 1919) 

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Distribution: Andhra Pradesh and Himachal Pradesh. 

171. Coccus viridis (Green, 1889) 

Distribution: Assam, Karnataka, Kerala, Odisha, Tamil Nadu and Tripura. 

172. Coccus watti (Green, 1900) 

Distribution: Assam, Meghalaya and West Bengal. 

173. Coccus radicicola Green, 1921 


174. Coccus olivaceum Green, 1922 


67. Genus: Eriochiton Maskell, 1887 

175. Eriochiton tessellates Signoret, 1873 


68. Genus: Eulecanium Cockerell, 1893 

Eulecanium sp. 


176. Eulecanium capreae (Linnaeus, 1767) 

Distribution: Jammu & Kashmir. 

177. Eulecanium coryli (Linnaeus, 1758) 

Distribution: Jammu & Kashmir and Uttar Pradesh. 

69. Genus: Hemilecanium Newstead, 1906 

178. Hemilecanium imbricans (Green, 1903) 

Distribution: Tamil Nadu (Coimbatore, Nilgiris) and Karnataka (Mysore, Belur). 

70. Genus: Maacoccus Tao et al., 1983 

179. Maacoccus bicruciatus Tao et al., 1983 

22


71. Genus: Marsipococcus Cockerell & Bueker, 1930 

180. Marsipococcus marsupiale (Green, 1904) 

Distribution: Kerala and Tamil Nadu. 

72. Genus: Paralecanium Cockerell & Parrot, 1899 

181. Paralecanium expansum expansum (Green, 1896) 

Distribution: Karnataka and Tamil Nadu (Chennai). 

182. Paralecanium maritimum Fernald, 1903 


73. Genus: Parthenolecanium Sulc, 1908 

183. Parthenolecanium persicae Fabricius, 1776 


74. Genus: Saissetia Deplanche, 1859 

Saissetia sp. 

Distribution: Odisha (Bhubaneswar, Baripada, Puri ). 

184. Saissetia coffeae (Walker, 1852) 

Distribution: Andhra Pradesh, Assam, Bihar, Goa, Karnataka, Maharashtra, Manipur, Odisha, 


185. Saissetia crassum Ali, 1971 


186. Saissetia depressa King, 1902 


187. Saissetia filicum Fernald, 1903 

Distribution: Karnataka, Odisha and Tamil Nadu. 

188. Saissetia miranda De Lotto, 1970 

23

Distribution: Tamil Nadu (Palani hills). 

189. Saissetia nigra King, 1902 

Distribution: Assam, Bihar, Jammu & Kashmir, Tamil Nadu, Uttar Pradesh and West Bengal. 

190. Saissetia lieae Cockerell, 1901 

Distribution: Assam (Tocklai). 

191. Saissetia privigna De Lotto, 1965 

Distribution: Andhra Pradesh (Guntur). 

75. Genus: Taiwansaissetia Tao et al., 1983 

192. Taiwansaissetia formicarii Tao et al., 1983 

Distribution: Assam and Karnataka. 

76. Genus: Chloropulvinaria Borchsenius, 1952 

Tribe: Pulvinariini 

193. Chloropulvinaria cellulose Varshney, 1985 

Distribution: Bihar, Uttar Pradesh and West Bengal. 

194. Chloropulvinaria floccifera (Westwood, 1870) 


195. Chloropulvinaria psidii Borchsenius, 1957 

Distribution: Andhra Pradesh, Bihar, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, 

Tripura and West Bengal. 

196. Chloropulvinaria venkatramani Varshney, 1992 [nomen novum] 


77. Genus: Micropulvinaria Hodgson, 1968 

197. Micropulvinaria maxima thespesiae (Green); Varshney, 1992 [n. comb.] 

Distribution: Andhra Pradesh and Tamil Nadu. 

78. Genus: Protopulvinaria Cockerell, 1894 

24

198. Protopulvinaria mangiferae Steinweden, 1929 

Distribution: Bihar, Tamil Nadu and West Bengal. 

79. Genus: Pulvinaria Targioni Tozzetti, 1866 

Pulvinaria spp. 

Distribution: Bihar and Odisha 

199. Pulvinaria aligarhensis Avasthi & Shafee, 1980 

Distribution: Uttar Pradesh (Aligarh). 

200. Pulvinaria burkilli Green, 1908 


201. Pulvinaria indica Avasthi & Shafee, 1980 


202. Pulvinaria obscura Newstead, 1894 


80. Genus: Takahashia Cockerell, 1986 

203. Takahashia japonica (Cockerell, 1896) 


81. Genus: Cardiococcus Cockerell, 1903 

Subfamily: Filipiinae 

204. Cardiococcus bivalvata (Green, 1903) 

Distribution: Andhra Pradesh and Tamil Nadu. 

82. Genus: Ceroplastodes Cockerell, 1893 

Ceroplastodes sp. 

Distribution: Bihar, Jammu & Kashmir, Odisha and Uttar Pradesh. 

205. Ceroplastodes cajani Cockerell, 1900 

25

Distribution: Andhra Pradesh, Bihar, Himachal Pradesh, Kerala, Punjab, Tamil Nadu and West 

Bengal. 

206. Ceroplastodes chiton Green, 1909 

Distribution: Himachal Pradesh, Uttar Pradesh and West Bengal. 

83. Genus: Dicyphoccus Borchsenius, 1959 

207. Dicyphococcus castilloae Borchsenius, 1959 


84. Genus: Inglisia Maskell, 1879 

208. Inglisia chelonioides Green, 1903 

Distribution: Kerala, Maharashtra (Shiroda) and Tamil Nadu (Coimbatore). 

85. Genus: Lecanopsis Targioni Tozzetti, 1868 

209. Lecanopsis ceylonica Green, 1922 


86. Genus: Metaceronema Takahashi, 1955 

210. Metaceronema japonica (Maskell, 1897) 

Distribution: West Bengal (Darjeeling). 

87. Genus: Leucaspis Signoret, 1869 

Superfamily: Diaspidoidea 

Family: Diaspididae 

Subfamily: Parlatoriinae 

Tribe: Leucaspidini 

211. Leucaspis manii (Borchsenius, 1964) 

Distribution: Meghalaya. 

212. Leucaspis riccae Targioni Tozzetti, 1881 

Distribution: Andhra Pradesh (Hyderabad). 

88. Genus: Lopholeucaspis Balachowsky, 1953 

26

Lopholeucaspis sp. 

Distribution: Karnataka (Anekal). 

213. Lopholeucaspis cockerelli (Grandpre & De Charmoy, 1899) 


214. Lopholeucaspis excoecariae Borchsenius, 1964 


215. Lopholeucaspis japonica (Cockerell, 1897) 

Distribution: Karnataka (Vasanthapura). 

216. Lopholeucaspis menoni Borchsenius, 1964 

Distribution: Rajasthan, Uttar Pradesh and West Bengal. 

89. Genus: Radionaspis Ferris, 1942 

217. Radionaspis indica (Marlatt, 1908) 

Distribution: Maharashtra (Pune). 

90. Genus: Salicicola Lindinger, 1905 

218. Salicicola indiaeorientalis (Lindinger, 1911) 

Distribution: Himachal Pradesh (Shimla). 

91. Genus: Apteronidia Berlese, 1895 

Tribe: Parlatoriini 

219. Apteronidia blanchardi (Targioni Tozzetti, 1892) 

Distribution: Andhra Pradesh (Hyderabad). 

92. Genus: Cryptoparlatoria Lindinger, 1905 

220. Cryptoparlatoria pini Takahashi, 1938 

Distribution: Himachal Pradesh (Manali). 

93. Genus: Genaparlatoria MacGillivray, 1921 

27

221. Genaparlatoria pseudaspidiotus (Lindinger, 1905) 

Distribution: Andhra Pradesh (Godavari Dist.). 

94. Genus: Greeniella Cockerell, 1897 

222. Greeniella cornigera (Green, 1896) 


223. Greeniella dentata (Lindinger, 1911) 


224. Greeniella ferreae (Rutherford, 1914) 


225. Greeniella viridis (Lindinger, 1911) 


95. Genus: Gymnaspis Newstead, 1898 

Gymnaspis sp. 

Distribution: Kerala (Kattuthurai). 

226. Gymnaspis affinis Ayyar, 1930 

Distribution: Tamil Nadu (Tirunelveli). 

227. Gymnaspis cassida Hall & Williams, 1962 


228. Gymnaspis ficus Green, 1919 


229. Gymnaspis mangifera Ayyar, 1924 

Distribution: Tamil Nadu (Tirunelveli) 

230. Gymnaspis ramakrishnai Green, 1919 


28

96. Genus: Neoleucaspis Green, 1926 

231. Neoleucaspis parallela Green, 1926 


97. Genus: Parlaspis McKenzie, 1945 

Parlaspis spp. 

Distribution: Karnataka and Kerala. 

232. Parlaspis papillosa (Green, 1919) 

Distribution: Kerala (Palghat). 

98. Genus: Parlatorearum Lindinger, 1911 

Parlatorearum sp. 

Distribution: Andhra Pradesh (Cuddapah). 

99. Genus: Parlatoreopsis Lindinger, 1911 

233. Parlatoreopsis chinensis (Marlatt, 1908) 

Distribution: Tamil Nadu (Coonoor, Coimbatore). 

100. Genus: Parlatoria Targioni Tozzetti, 1868 

Parlatoria sp. 

Distribution: Karnataka and Tamil Nadu. 

234. Parlatoria artocarpi Green, 1919 

Distribution: Kerala (Peria Ghat). 

235. Parlatoria atalantiae Green, 1905 


236. Parlatoria boycei MacKenzie, 1952 


237. Parlatoria camelliae Comstock, 1883 

29


238. Parlatoria cinerea Hadden in Doane & Hadden, 1909 

Distribution: West Bengal and South India. 

239. Parlatoria crotonis Douglas, 1887 

Distribution: Bihar 

240. Parlatoria crypta McKenzie, 1943 

Distribution: Andhra Pradesh, Delhi, Karnataka, Maharashtra and Punjab. 

241. Parlatoria marginalis McKenzie, 1945 

Distribution: Delhi. 

242. Parlatoria morrisoni McKenzie, 1943 


243. Parlatoria mytilaspiformis Green, 1899 


244. Parlatoria oleae (Colvee, 1880) 

Distribution: Bihar, Kashmir, Maharashtra, Rajasthan and West Bengal. 

245. Parlatoria orientalis Rao, 1953 


246. Parlatoria pergandii Comstock, 1881 

Distribution: Bihar, Karnataka, Kerala and Tamil Nadu (Coimbatore, Mundathorai, Tirunelvelly) 

247. Parlatoria proteus (Curtis, 1843) 

Distribution: Karnataka, Tamil Nadu (Coimbatore, Nilgiris), Tripura and West Bengal. 

248. Parlatoria vateriae Green, 1919 

Distribution: Kerala (Travancore, Kollam). 

249. Parlatoria ziziphi (Lucas, 1853) 

30

Distribution: Tamil Nadu (Coimbatore, Ramanathapuram, Tanjore) and West Bengal. 

101. Genus: Tamilparla Takagi, 1987 

250. Tamilparla smilacis Takagi, 1987 

Distribution: Tamil Nadu (Coonoor, Nilgiris). 

102. Genus: Mangaspis Takagi & Kondo, 1997 

Subfamily: Odonaspidinae 

Tribe: Rugaspidiotini 

251. Mangaspis Bengalurunsis Takagi & Kondo, 1997 


103. Genus: Natalaspis MacGillivray, 1921 

252. Natalaspis simplex (Green, 1899); Ben-Dov & Takagi, 1974 


104. Genus: Ramachandraspis Rao, 1953 

253. Ramachandraspis fenestrata Rao, 1953 


Tribe: Odonaspidini 

105. Genus: Berlesaspidiotus MacGillivray, 1921 

254. Berlesaspidiotus crenulatus Ben-Dev, 1988 

Distribution: Tamil Nadu (Chennai, Palani). 

106. Genus: Froggattiella Leonardi, 1900 

255. Froggattiella penicillata (Green, 1905); Rutherford, 1915 


107. Genus: Odonaspis Leonardi, 1897 

Odonaspis spp. 


31

256. Odonaspis paucipora Ben-Dov, 1988 

Distribution: West Bengal (Kolkata). 

257. Odonaspis saccharicaulis (Zehntner, 1897); Fernald, 1903 

Distribution: Karnataka, Odisha and Tamil Nadu (Coimbatore). 

108. Genus: Abgrallaspis Balachowsky, 1948 

Subfamily: Aspidiotinae 

Tribe: Aspidiotini 

258. Abgrallaspis cyanophylli Balachowsky, 1948 

Distribution: Karnataka (Bengaluru Kerala and Tamil Nadu (Coimbatore). 

109. Genus: Aonidia Targioni - Tozzetti, 1868 

259. Aonidia crenulata Green, 1900 


260. Aonidia indica Green, 1919 


261. Aonidia loranthi Green, 1896 


262. Aonidia tentaculata Green, 1919 


110. Genus: Aonidiella Berlese & Leonardi, 1895 

263. Aonidiella andersoni (Laing, 1929); McKenzie, 1938 


264. Aonidiella aurantii (Maskell, 1879) 

Distribution: Andhra Pradesh, Assam, Bihar, Delhi, Kanataka, Kerala, Odisha, Punjab and Tamil 

Nadu. 

265. Aonidiella citrine Nel, 1933 

32


266. Aonidiella comperei McKenzie, 1937 

Distribution: Maharashtra (Mumbai, Colaba). 

267. Aonidiella inortata McKenzie, 1938 


268. Aonidiella misrai Laing, 1929 


269. Aonidiella orientalis McKenzie, 1937 

Distribution: Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Punjab, Tamil 

Nadu, Tripura and West Bengal. 

111. Genus: Aspidiella Leonardi, 1898 

270. Aspidiella hartii (Cockerell, 1895) 

Distribution: Maharashtra (Pune) and Tamil Nadu (Coimbatore). 

271. Aspidiella sacchari MacGillivray, 1921 [comb. new.] 


112. Genus: Aspidiotus Bouche, 1833 

Aspidiotus spp. 

Distribution: Andhhra Pradesh, Odisha and Karnataka. 

272. Aspidiotus destructor Signoret, 1869 

Distribution: Andhra Pradesh, Kerala, Jammu & Kashmir, Maharashtra (Mumbai), Rajasthan, 

Tripura, West Bengal and Lakshadweep Is. 

273. Aspidiotus hederae (Vallot, 1829) 


274. Aspidiotus pandani (Boisduval, 1868) 


33

113. Genus: Borchseniaspis Zahradnik, 1959 

275. Borchseniaspis palmae Zahradnik, 1959 


114. Genus: Brainaspis MacGillivray, 1921 

276. Brainaspis kellyi MacGillivray, 1921 


115. Genus: Clavaspis MacGillivray, 1921 

Clavaspis sp. 


116. Genus: Chrysomphalus Ashmead, 1880 

277. Chrysomphalus aonidum (Linnaeus, 1758) 

Distribution: Andhra Pradesh, Assam, Bihar, Kerala, Jammu & Kashmir, Maharashtra 

(Mumbai), Madhya Pradesh, Gujarat, Tamil Nadu, Tripura and West Bengal. 

278. Chrysomphalus dictyospermi (Morgan, 1889) 

Distribution: Andhra Pradesh, Assam, Meghalaya, Tamil Nadu and Tripura. 

279. Chrysomphalus diversicolor (Green, 1923) 

Distribution: Karnataka and Tamil Nadu. 

117. Genus: Cryptopalratoreopsis Borchsenius, 1947 

280. Cryptopalratoreopsis euphorbiae Hall & Williams, 1962 


281. Cryptopalratoreopsis longispina (Takahashi, 1939) 


282. Cryptopalratoreopsis spinosissima (Lindinger, 1911) 

Distribution: India (Central India). 

283. Cryptopalratoreopsis targionipsis (Lindinger, 1911) 

34


118. Genus: Diaspidiotus Berlese & Leonardi, 1896 

284. Diaspidiotus pseudocamelliae Borchsenius, 1966 


119. Genus: Duplaspidiotus MacGillivray, 1921 

285. Duplaspidiotus tesseratus (Grandpre & de Charmoy, 1899) 

Distribution: Karnataka and Maharashtra (Mumbai, Lonavala). 

120. Genus: Hemiberlesia Cockerell, 1897 

286. Hemiberlesia lataniae (Signoret, 1869) 

Distribution: Assam, Odisha, Karnataka, Maharashtra (Mumbai), Tamil Nadu, Tripura and West 

Bengal. 

287. Hemiberlesia rapax (Comstock, 1881) 

Distribution: Assam, Odisha, Karnataka, Tamil Nadu, Tripura and West Bengal. 

121. Genus: Lindingaspis MacGillivray, 1921 

Lindingaspis sp. 


288. Lindingaspis floriadana Ferris, 1942 


289. Lindingaspis fusca McKenzie, 1943 

Distribution: Maharashtra (Pune) and South India. 

290. Lindingaspis rossi (Maskell, 1891) 

Distribution: Andhra Pradesh, Maharashtra (Pune) and Madhya Pradesh. 

122. Genus: Melanaspis Cockerell, 1897 

Melanaspis sp. 


35

291. Melanaspis glomerata (Green, 1903) 

Distribution: Andhra Pradesh, Bihar, Odisha, Gujarat, Tamil Nadu and Uttar Pradesh. 

292. Melanaspis smilacis (Comstock, 1883) 


123. Genus: Metaspidiotus Takagi, 1957 

293. Metaspidiotus sp. 

Distribution: Karnataka and Goa. 

124. Genus: Morganella Cockerell, 1897 

294. Morganella cueronsis (Cockerell, 1899) 

Distribution: Tamil Nadu and South India. 

295. Morganella longispina (Morgan, 1889) 


125. Genus: Neofurcaspis Green, 1926 

296. Neofurcaspis andamanensis Green, 1926 

Distribution: Andaman Is. 

126. Genus: Octaspidiotus MacGillivray, 1921 

297. Octaspidiotus stauntoniae (Takahashi, 1933) 

Distribution: Karnataka and Goa. 

298. Octaspidiotus australiensis (Kuwana, 1931) 


299. Octaspidiotus calophylli var. symplocos (Green, 1924) 

Distribution: Tamil Nadu (Nilgiris, Coonoor, Kotagiri). 

300. Octaspidiotus tamarindi (Green, 1919) 

Distribution: Karnataka (Nandi) and Tamil Nadu (Coimbatore). 

36

301. Octaspidiotus tripurensis Takagi, 1984 

Distribution: Tripura (Agartala). 

127. Genus: Parrottia MacGillivray, 1921 

Parrottia sp. 

Distribution: Karnataka (Devarayanadurga). 

302. Parrottia moorei (Green, 1896) 


128. Genus: Pseudaonidia Cockerell, 1897 

Pseudaonidia spp. 


303. Pseudaonidia duplex (Cockerell, 1896) 


304. Pseudaonidia trilobitiformis (Green, 1896) 

Distribution: Andhra Pradesh, Bihar, Kerala, Tamil Nadu and West Bengal. 

129. Genus: Quadraspidiotus MacGillivray, 1921 

305. Quadraspidiotus perniciosus (Comstock, 1881) 

Distribution: Andhra Pradesh, Bihar, Himachal Pradesh, Jammu & Kashmir, Karnataka, Kerala, 

Maharashtra (Mahabaleshwar), Meghalaya, Odisha, Punjab, Sikkim, Tamil Nadu, Uttar Pradesh 

and West Bengal. 

130. Genus: Semelaspidus MacGillivray, 1921 

306. Semelaspidus artocarpi (Green, 1896) 

Distribution: Karnataka (Bengaluru, Coorg) and Maharashtra (Mahabaleshwar). 

131. Genus: Temnaspidiotus MacGillivray, 1921 

307. Temnaspidiotus excisus (Green, 1896) 

Distribution: South India and Tripura (Agartala). 

37

132. Genus: Anaimalaia Takagi, 1995 

308. Anaimalaia scabra Takagi, 1995 

Distribution: Tamil Nadu (Anaimalai Hills). 

133. Genus: Kuwanaspis MacGillivray, 1921 

Subfamily: Diaspidinae 

Tribe: Protodiaspidini 

Tribe: Kuwanaspidini 

309. Kuwanaspis annandalei (Green, 1919) 

Distribution: Bihar (Parasnath Hills). 

134. Genus: Xiphuraspis Borchsenius & Williams, 1963 

310. Xiphuraspis spiculata Borchsenius & Williams, 1963 

Distribution: Kerala (N. Malabar). 

135. Genus: Coronaspis MacGillivray, 1921 

311. Coronaspis malabarica Takagi, 1999 

Distribution: Kerala (Periyar Tiger Reserve) and Tamil Nadu (Anaimalai Hills). 

136. Genus: Acanthomytilus Borchsenius, 1947 

Tribe: Lepidosaphidini 

312. Acanthomytilus sacchari (Hall, 1923) 

Distribution: Bihar, Tamil Nadu (Coimbatore), Uttar Pradesh and West Bengal. 

313. Acanthomytilus vermiformis (Takahashi, 1931) 

Distribution: Kerala (Dhony Valley). 

137. Genus: Andaspis MacGillivray, 1921 

Andaspis sp. (nr. leucophloeae Rao) 


314. Andaspis artocarpi Borchsenius, 1967 

38


315. Andaspis betulae (Borchsenius, 1967) 

Distribution: Meghalaya (Shilong). 

316. Andaspis laingi Rao, 1952 

Distribution: Tamil Nadu (Coimbatore, Makkalidurga). 

317. Andaspis leucophleae Rao, 1952 

Distribution: Tamil Nadu (Coimbatore, Chennai) and Madhya Pradesh. 

318. Andaspis meliae (Green, 1919) 

Distribution: Andhra Pradesh, Karnataka (Mandya), Tamil Nadu (Coimbatore) and West Bengal 

(Kolkata). 

319. Andaspis numerate Brimblecombe, 1959 


320. Andaspis retrusa (Green, 1919) 


138. Genus: Aonidomytilus Leonardi, 1903 

321. Aonidomytilus albus (Cockerell, 1893) 

Distribution: Karnataka (Bengaluru), Tamil Nadu (Coimbatore, Salem, Puttukottai) and 

Puducherry. 

139. Genus: Cornuaspis MacGillivray, 1921 

322. Cornuaspis beckii (Newmann, 1869) 

Distribution: Assam, Karnataka, Kerala and West Bengal. 

323. Cornuaspis piperis (Green, 1908) 

Distribution: Western Ghats; Kerala (N. Malabar, Wynaad, Travancore). 

140. Genus: Eucornuaspis Borchsenius, 1963 

324. Eucornuaspis pinnaeformis Borchsenius, 1966 

39


325. Eucornuaspis pseudomachili Borchsenius, 1964 


141. Genus: Howardia Berlese & Leonardi, 1896 

326. Howardia biclavis (Comstock, 1883) 

Distribution: Karnataka, Kerala (Wynaad) and Tamil Nadu (Chennai, Anaimalai Hills). 

142. Genus: Insulaspis Mamet, 1950 

327. Insulaspis citrine Borchsenius, 1964 

Distribution: New Delhi. 

328. Insulaspis gloverii (Packard, 1869) 

Distribution: Bihar, Karnataka and West Bengal. 

329. Insulaspis mcgregori (Banks, 1906) 

Distribution: Kerala (Thiruvananthapuram Dist.). 

330. Insulaspis pallidula Williams, 1969 

Distribution: Andhra Pradesh (Godavari Dist.). 

331. Insulaspis tapleyi (Williams, 1960); Borchsenius, 1963 

Distribution: Karnataka and Uttar Pradesh. 

332. Insulaspis tokionis (Kuwana, 1902); Borchsenius, 1963 


143. Genus: Lepidosaphes Shimer, 1868 

333. Lepidosaphes cornutus Green in Ayyar, 1937 

Distribution: Kerala and Tamil Nadu (North Arcot Dist., South Arcot Dist., Tanjore, Salem). 

334. Lepidosaphes malicola Borchsenius, 1947 

Distribution: Jammu & Kashmir (Anantnag). 

40

335. Lepidosaphes ulmi (Linnaeus, 1758) 


144. Genus: Neopinnaspis McKenzie, 1949 

336. Neopinnaspis travancorensis (Lindinger, 1911) 


145. Genus: Velataspis Ferris, 1937 

337. Velataspis serrulata Ganguli, 1958 


146. Genus: Africaspis MacGillivray, 1921 

Tribe: Diaspidini 

338. Africaspis chionaspiformis (Newstead, 1910) 

Distribution: Tamil Nadu (Coimbatore, Tirunelveli). 

147. Genus: Aulacaspis Cockerell, 1893 

Aulacaspis spp. 

Distribution: Karnataka and Kerala. 

339. Aulacaspis gudalura Ferris, 1956 

Distribution: Tamil Nadu (Nilgiris). 

340. Aulacaspis hedyotidis Takagi, 1985 


341. Aulacaspis herbae Hall, 1946 


342. Aulacaspis litzeae (Green, 1896) 


343. Aulacaspis loranthi Cockerell, 1902 

41

Distribution: Bihar (Parasnath Hill). 

344. Aulacaspis madiunensis (Zehtner, 1898) 

Distribution: Andhra Pradesh and Tamil Nadu (Chengalpattu, North & South Arcot Dist., 

Tanjore, Tiruchirapalli). 

345. Aulacaspis malayala Varshney, 2002 

Distribution: Kerala (Periyar Tiger Reserve). 

346. Aulacaspis mangiferae (Newstead, 1908) 

Distribution: Andhra Pradesh, Karnataka and Tamil Nadu (Tirunelveli). 

347. Aulacaspis orientalis (Green, 1922) 

Distribution: Tamil Nadu (Courtallum, Tirunelveli). 

348. Aulacaspis rosae (Bouche, 1833) 

Distribution: Tamil Nadu (Courtallum, Tirunelveli). 

349. Aulacaspis tubercularis (Newstead, 1908) 

Distribution: Karnataka, Kerala and Uttar Pradesh. 

350. Aulacaspis vitis (Green, 1896) 

Distribution: Assam, Bihar, Himachal Pradesh, Odisha, Karnataka and Tamil Nadu. 

148. Genus: Chionaspis Signoret, 1869 

Chionaspis sp. 


351. Chionaspis engeddensis Bodenheimer, 1924 

Distribution: Punjab. 

352. Chionaspis furfura (Fitch, 1857) 

Distribution: Jammu & Kashmir. 

353. Chionaspis himalaica Takagi, 1985 


42

354. Chionaspis ramakrishnai Rao, 1953 


355. Chionaspis salicis (Linnaeus, 1758) 

Distribution: Himalayas and Jammu & Kashmir. 

149. Genus: Contigaspis MacGillivray, 1921 

356. Contigaspis coimbatorensis Borchsenius & Williams, 1963 

Distribution: Karnataka and Tamil Nadu (Coimbatore). 

357. Contigaspis indigoferae (Hall, 1928) 

Distribution: Tamil Nadu (Hosur). 

150. Genus: Dentachionaspis MacGillivray, 1921 

358. Dentachionaspis centripetalis Rao, 1953 

Distribution: Jammu & Kashmir (Srinagar). 

151. Genus: Diaspis Costa, 1828 

Diaspis sp. 

Distribution: Odisha and Tamil Nadu. 

359. Diaspis boisduvali Signoret, 1869 

Distribution: Tamil Nadu (Nilgiris). 

360. Diaspis echinocacti (Bouche, 1833) 

Distribution: Karnataka, Kerala, Maharashtra (Pune), Odisha, Tamil Nadu and Uttar Pradesh. 

152. Genus: Duplachionaspis MacGillivray, 1921 

361. Duplachionaspis divergens (Green, 1899) 

Distribution: Kerala, Punjab and Tamil Nadu. 

153. Genus: Fiorinia Targioni- Tozzetti, 1868 

Fiorinia spp. 

43

Distribution: Andhra Pradesh and Tripura. 

362. Fiorinia fioriniae (Targioni Tozzetti, 1867) 


363. Fiorinia fletcheri Laing, 1929 


364. Fiorinia frontecontracta Green, 1919 

Distribution: Maharashtra (Mumbai). 

365. Fiorinia himalaica Takagi, 1975 

Distribution: Himachal Pradesh and Uttar Pradesh. 

366. Fiorinia japonica Kuwana, 1902 

Distribution: West Bengal (Kolkata). 

367. Fiorinia multipora Lindinger, 1911 


368. Fiorinia pakistanensis Ahmad, 1970 

Distribution: Kashmir. 

369. Fiorinia plana Green, 1919 

Distribution: Tamil Nadu (Coimbatore, Tirunelveli). 

370. Fiorinia proboscidaria Green, 1920 


371. Fiorinia sapindi Green, 1919 

Distribution: Maharashtra (Pune). 

372. Fiorinia theae Green, 1900 

Distribution: Assam, North India and West Bengal (Darjeeling). 

154. Genus: Greenaspis MacGillivray, 1921 

44

373. Greenaspis decurvata (Green, 1903) 

Distribution: Kerala, Odisha, Tamil Nadu and West Bengal. 

374. Greenaspis elongata (Green, 1896) 


155. Genus: Lineaspis MacGillivray, 1921 

375. Lineaspis caroli (Green, 1919) 


156. Genus: Myrtaspis Takagi, 1999 

376. Myrtaspis syzygii (Takagi, 1985) 


157. Genus: Pinnaspis Cockerell, 1892 

Pinnaspis sp. 

Distribution: Karnataka (Thalghatpura, Pannayanpalaya, Shivganga). 

Pinnaspis sp. [nr. strachani (Cooley, 1899)]. 

Distribution: West Bengal (Barrackpore) and Karnataka (Gokarna). 

377. Pinnaspis aspidistrae (Signoret, 1869) 

Distribution: Andhra Pradesh, Karnataka (Shivganga), Kerala (Cochin, Malabar), Madhya 

Pradesh, Maharashtra (Janjira Estate), Tamil Nadu (Coimbatore, Nilgiris) and Tripura. 

378. Pinnaspis buxi (Bouche, 1851) 


379. Pinnaspis dracaenae (Cooley, 1899) 

Distribution: Kerala (South Malabar, Vadanapalle). 

380. Pinnaspis exercitata (Green, 1896) 

Distribution: Karnataka (Coorg), Odisha, Tamil Nadu (Coimbatore, Chennai) and West Bengal. 

381. Pinnaspis fici (Green, 1908) 

45


382. Pinnaspis minima (Green, 1908) 


383. Pinnaspis strachani (Cooley, 1899) 

Distribution: Andhra Pradesh, Bihar, Karnataka (Bengaluru, Devanahally, Bellari, Vasanthapura, 

Hoskote, Nandi, Koramangala), Kerala (Kondayur), Jammu & Kashmir, Tamil Nadu 

(Coimbatore, Nilgiris, Marudamali) and West Bengal. 

384. Pinnaspis theae (Maskell, 1891) 

Distribution: North India (Assam) and West Bengal (Darjeeling). 

385. Pinnaspis uniloba (Kuwana, 1909) 

Distribution: Assam and Tamil Nadu (Chennai, Pudukottai). 

158. Genus: Poliaspis Maskell, 1880 

386. Poliaspis cycadis Comstock, 1883 

Distribution: Maharashtra (Kolhapur). 

159. Genus: Pseudaulacaspis MacGillivray, 1921 

Pseudaulacaspis sp. 

Distribution: Karnataka (Kodigehalli, Namadachiluve). 

387. Pseudaulacaspis barberi (Green, 1908) 

Distribution: Andhra Pradesh (Visakhapatnam, Alamanda), Karnataka (Bengaluru, Bellari) and 

Tamil Nadu (Tanjore Dist.). 

388. Pseudaulacaspis cockerelli (Cooley, 1897) 

Distribution: Andhra Pradesh (Godavari Dist.), Karnataka (Bengaluru, Lal Bagh, Koramangala), 

Kerala (Malabar, Wynaad, Kondayur), Maharashtra (Pune), Tamil Nadu (Hosur, Nilgiris) and 

West Bengal (Kolkata). 

389. Pseudaulacaspis gynandropsidis (Green, 1922) 

Distribution: Karnataka (Bengaluru, Jalahalli). 

46

390. Pseudaulacaspis manni (Green, 1907) 

Distribution: Assam, Himachal Pradesh (Kangra), Tamil Nadu (Hosur) and West Bengal 

(Kolkata, Darjeeling). 

391. Pseudaulacaspis megaloba (Green, 1899) 


392. Pseudaulacaspis pentagona pentagona (Targioni- Tozzetti, 1886) 

Distribution: Assam (Maungep, Sonapari), Karnataka (Byadarahalli, Vasanthapura), Meghalaya 

(Shilong) and Uttar Pradesh (Chaubattia). 

393. Pseudaulacaspis pentagona theae (Maskell, 1896) 


394. Pseudaulacaspis polygoni (Green, 1899) 

Distribution: Karnataka (Mallandur). 

395. Pseudaulacaspis pusa (Rao, 1953) 


396. Pseudaulacaspis subcorticalis (Green, 1905) 

Distribution: Karnataka (Jayanthigrama, Kuppehalli, Kobbinahalli). 

397. Pseudaulacaspis tenera (Green, 1922) 

Distribution: Karnataka (Karkada). 

398. Pseudaulacaspis varicosa (Green, 1896) 

Distribution: Karnataka (Coorg, Sidapur) and Tamil Nadu (Mundathurai, Nilgiris, Tirunelveli). 

399. Pseudaulacaspis venue (Menon & Khan, 1961) 

Distribution: Himachal Pradesh (Kangra, Palampur). 

160. Genus: Pseudoparlatoria Cockerell, 1892 

400. Pseudoparlatoria parlatorioides (Comstock, 1883) 


47

161. Genus: Rutherfordia MacGillivray, 1921 

401. Rutherfordia major (Cockerell, 1894) 

Distribution: Kerala (Periyar). 

162. Genus: Sinistraspis MacGillivray, 1921 

402. Sinistraspis unilateralis (Newstead, 1913) 

Distribution: Kerala (Thiruvananthapuram). 

163. Genus: Tenuiaspis MacGillivray, 1921 

403. Tenuiaspis minuta (Green, 1896) 


164. Genus: Thysanofiorinia Balachowsky, 1954 

404. Thysanofiorinia leei Williams, 1971 

Distribution: Tripura (Agartala) and Uttarakhand (Dehra Dun). 

405. Thysanofiorinia nephelii (Maskell, 1897) 

Distribution: Karnataka (Coorg, Siddapur) and West Bengal (Krishnanagar). 

165. Genus: Trullifiorinia Leonardi, 1906 

406. Trullifiorinia geloniae (Green, 1900) 

Distribution: Tamil Nadu (Ooty). 

166. Genus: Unaspis MacGillivray, 1921 

Unaspis sp. 

Distribution: Karnataka (Nandi). 

407. Unaspis acuminate (Green, 1896) 

Distribution: Kerala (Chalikudi, Malabar, Peria Ghat) and Tamil Nadu (Madurai). 

408. Unaspis atricolor (Green, 1922) 

Distribution: Andhra Pradesh (Hyderabad) and Tamil Nadu (Coimbatore, Dharmapuri). 

48

409. Unaspis permutans (Green, 1899) 

Distribution: Kerala (Malabar, Peria Pass). 

REFERENCES 

Varshney R. K. 1992. A Check list of Scale insects and mealy bugs of South Asia, (Part-1), 

Rec. zool. Surv. India, Occ. Paper, 139: 152. 

Varshney R. K. 2002. A Check list of Scale insects and mealy bugs of South Asia, (Part-2), 

Rec. zool. Surv. India, Occ. Paper, 191: 147. 

Williams D. J. 1978. The anomalous ant-attended mealy bugs (Homoptera: Pseudococcidae) 

of South-east Asia, Bull. Br. Mus. nat. (Ent.), 37 (1): 1-72. 

Y. Ben-Dev, C.J.Hodgson and D.R.Miller. 1997. Changes and Comments on the Taxonomy and 

Nomenclature of Some Taxa in the Families Coccidae, Eriococcidae and 

Pseudococcidae (Homoptera: Coccoidae). Phytoparasitica, 25(3): 199-206. 

Internet Source : http://www.sel.barc.usda.gov/SCALENET/SCALENET.HTM 

49

scale insects and mealy bugs - Zoological Survey of India

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