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48 M. J. WIGGINTON Leptocolea mascarena Steph. ex S.W.Arnell 1965, nom. inval. = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). Leptocolea montagnei (Gottsche) Steph. 1916 = Cheilolejeunea montagnei (Gottsche) Schiffn. 1898 (1). Leptocolea nigerica E.W.Jones 1953 = Cololejeunea nigerica (E.W.Jones) R.M.Schust. 1963 (912). Leptocolea nymanii Steph. 1916 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (135). Leptocolea perforata Schiffn. ex S.W.Arnell 1965, nom. inval. = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). Leptocolea perforata sensu S.W.Arnell 1965 [non Schiffn. ex S.W.Arnell 1965, nom. inval.] = Drepanolejeunea madagascariensis (Steph.) Grolle 1974 (924). Leptocolea punctata E.W.Jones 1957 = Cololejeunea vidaliana Tixier 1973 (847). Leptocolea quintasii var mascarenica S.W.Arnell 1965, nom. inval. = Leptolejeunea epiphylla (Mitt.) Steph. 1913 (136). Leptocolea scabrifolia var. africana Herzog 1947, nom. inval. = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (1). Leptocolea subciliata (Mitt.) E.W.Jones 1953 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 (152). Leptocolea tonkinensis (Steph.) Steph. 1916 = Cololejeunea tonkinensis Steph. 1895 (135). Leptolejeunea adhaesiva (Mitt.) Steph. 1895 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963 (912). Leptolejeunea astroidea (Mitt.) Steph. BUR(2), CMN(2), CON(2), GHA(2), GNB(2), GUI(2), IVO(2), NGA(2), RIM(10), SIE(2), UGA(42,50,115), ZAI(2), ZIM(2). Leptolejeunea convexistipa Bischl. 1969 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). Leptolejeunea degelii S.W.Arnell 1962 = Diplasiolejeunea symoensii Vanden Berghen 1960 (495). Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. COM(1). Leptolejeunea epiphylla (Mitt.) Steph. CMN(2), COM(1), CON(2), GHA(2), GUI(2), IVO(2), KEN(2), MAU(1,4), MDG(720), NGA(2), RIM(10), SAO(2), SEY(4,108), SIE(2), TAN(2,736), UGA(42), ZAI(2). Leptolejeunea maculata (Mitt.) Schiffn. CMN(2), COM(1,4), GAB(2), GHA(2), GUI(2), IVO(2), KEN(2,731), MAU(1), NGA(2), RIM(10), ROD(723), SAO(2), SEY(1,4,108), SIE(2), TAN(2), TOG(2), UGA(42,50,96), ZAI(2). Leptolejeunea madagascariensis Steph. 1913 = Drepanolejeunea madagascariensis (Steph.) Grolle 1974 (924). Leptolejeunea mascarena Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). Leptolejeunea papuliflora Steph. 1913 = Leptolejeunea quintasii Steph. 1891 (99) = Leptolejeunea epiphylla (Mitt.) Steph. 1913 (130). Leptolejeunea perforata (Schiffn.) Schiffn. 1897 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). Leptolejeunea quintasii (Steph.) Steph. 1913 = Leptolejeunea epiphylla (Mitt.) Steph. 1913 (130). Vanden Berghen (733) considered that L. epiphylla is perhaps synonymous with L. quintasii, and Tixier (130) treated them as synonymous without further comment. Leptolejeunea quintasii var. mascarenica S.W.Arnell 1965, nom. inval. = Leptolejeunea epiphylla (Mitt.) Steph. 1913 (136). Leptolejeunea repanda (Schiffn.) Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). Leptolejeunea symoensii Vanden Berghen 1961 = Drepanolejeunea symoensii (Vanden Berghen) Grolle 1976 (924). Leptolejeunea thomeensis (Steph.) Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). Leptolejeunea trematodes (Nees) Steph. 1913 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). Leptolejeunea triocellata Steph. 1913 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). Leptolejeunea truncatiflora Steph. 1911 = Leptolejeunea astroidea (Mitt.) Steph. 1913 (99). Leptolejeunea truncatiloba (Steph.) Steph. 1896 = Leptolejeunea thomeensis (Steph.) Steph. (161) = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). Leptolejeunea vitrea (Nees) Schiffn. MAS, with doubt(1). Leptoscyphus bewsii Sim 1926 = Lophocolea bewsii (Sim) Grolle 1959 (471). Leptoscyphus diversifolius (Gottsche) Grolle SAF(2). Leptoscyphus expansus (Lehm.) Grolle KEN(731), LES(75), MDG(1), MLW(29), RWA(2), SAF(2), ZAI(2), ZIM(2). Leptoscyphus gottscheanus (Lindenb. ex Lehm.) Gottsche 1858 = L. expansus (Lehm.) Grolle 1963 (812, 471). Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust. KEN(2,731), RWA(2), TAN(2), UGA(2,96), ZAI(2). Leptoscyphus henkelii Sim 1926 = Heteroscyphus grandistipus subsp. henkelii (Sim) Grolle 1963 (471). Leptoscyphus infuscatus (Mitt.) E.W.Jones BIO(2,33,36), CMN(2), ETH(2), MDG(720), MLW(2,29), REU(1,4), RWA(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2).
CHECKLIST OF THE LIVERWORTS AND HORNWORTS OF SUB-SAHARAN AFRICA 49 Leptoscyphus iversenii (Pearson) Sim 1926 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). Leptoscyphus leightonii Sim 1926 = Lophocolea newtonii Steph. 1907 (812) = Lophocolea congoana Steph. in Renauld 1902/3 (313) = Lophocolea martiana Nees 1845 (974). Leptoscyphus stephensii Sim 1926 = Lophocolea cuspidata (Nees) Limpr. 1876 (812) = Lophocolea bidentata (L.) Dumort. 1835 (975). Lethocolea congesta (Lehm.) S.W.Arnell COM(1,4), KEN(2), LES(75), REU(1,4), RWA(2), SAF(2), TAN(2), UGA(2), ZAI(2). Leucolejeunea capensis S.W.Arnell 1953 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 (806, 867). Leucolejeunea knysnana S.W.Arnell 1953 = Leucolejeunea unciloba (Lindenb.) A.Evans 1907 (867). Leucolejeunea rotalis (Hook.f. et Taylor) S.W.Arnell 1961 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997 (55). Leucolejeunea rotundistipula (Lindenb. ex Lehm.) Steph. LES(75), MAU(1), MLW(563), REU(1), SAF(2,75), ZIM(2). Leucolejeunea unciloba (Lindenb.) A.Evans KEN(731), SAF(2), TAN(2), UGA(776). Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans BIO(2,33), COM(4), ETH(2), KEN(2,731), MDG(4), MLW(563), MOZ(2), REU(1,4), RWA(2), SAF(2), STH(2), TAN(2), UGA(2,42), ZAI(2), ZIM(2). Lindigina renifolia Mitt. 1877 = Gongylanthus renifolius (Mitt.) Steph. 1906 (170). Lindigina scariosa (Lehm.) Mitt. 1877 = Gongylanthus scariosus (Lehm.) Steph. 1906 (170). Lophocolea amanica Steph. 1922 = Lophocolea congoana Steph. in Renauld 1902/3 (313) = Lophocolea martiana Nees 1845 (974). Lophocolea angustistipula Steph. STH(887). Overlooked in (2) and in edition 1 of this checklist. Lophocolea armatistipula Steph. 1911 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (165). Lophocolea ascensionis Steph. ASC(2). Lophocolea bewsii (Sim) Grolle SAF(2). Lophocolea bidentata (L.) Dumort. BIO(2,33,36), BUR(2), CMN(2), CVI(2,37), ETH(2), KEN(2,21,731), MAU(1), MDG(1,720), MLW(2,29), REU(1), RWA(2), SAF(2), STH(49), TAN(2), UGA(2,42,96), ZAI(2). Lophocolea bidentata sensu Sim 1926 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 1845 (313). Lophocolea bidentata var. δ capensis Gottsche et al. 1845 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 1845 (313). Lophocolea borbonica Steph. 1890 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 1845 (313). Lophocolea cagnii Gola 1907 = Lophocolea difformis Nees 1845 (814). Lophocolea cambouena Steph. 1907 = Lophocolea subrotunda Mitt. 1879 (165) = Lophocolea concreta Mont. 1845 (313). Lophocolea concreta Mont. ANN(2,33), BUR(2), GHA(2), KEN(2), LES(75), MAU(1), MDG(1,720), MLW(2,29), NGA(2), REU(1), ROD(1,723), RWA(2), SAF(2,75), SIE(2), SWZ(2), TAN(2), TOG(2), UGA(42,50), ZAI(2), ZAM(2), ZIM(2). Lophocolea congoana Steph. 1905 = Lophocolea martiana Nees 1845 (974). Lophocolea connata (Sw.) Mont. et Nees 1839 MAU(1). [Excludenda(1)]. Lophocolea cuspidata (Nees) Limpr. 1876 = Lophocolea bidentata (L.) Dumort. 1835 (975). Lophocolea devexa Mitt. 1863 = Conoscyphus trapezioides (Sande Lac.) Schiffn. 1898 (916). Lophocolea difformis Nees BIO(2,33,36), CAF(2), CMN(2), ETH(2), GHA(2), IVO(51), KEN(2,731), LES(75), MAU(4), MDG(720), MLW(2,29), NGA(2), REU(1), RWA(2), SAF(2,75), SAO(2), SIE(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), ZIM(2). Lophocolea diversifolia Gottsche 1845 = Leptoscyphus diversifolius (Gottsche) Grolle 1963 (471). Lophocolea diversifolia sensu Sim 1926 [non Gottsche 1845] = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 1845 (313). Lophocolea dusenii Steph. 1907 = Lophocolea congoana Steph. 1905 [= Lophocolea martiana Nees 1845] or Lophocolea opposita Mitt. 1879 [= Lophocolea lucida (Spreng. ex Lehm.) Mont. 1839] (165). Lophocolea esterhuysenii S.W.Arnell ex E.W.Jones 1953 = Lophocolea bewsii (Sim) Grolle 1959 (313). Lophocolea expansa (Lehm.) Drège 1843 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). Lophocolea fasciculata (Nees) S.W.Arnell et Grolle 1959 = Clasmatocolea fasciculata (Nees) Grolle 1960 (314, 97). Lophocolea fragrans (Moris et De Not.) Gottsche et al. COM(1), MLW(29), REU(1), SAF(2), TAN(2), ZAI(2). Lophocolea frappieri Steph. 1907 = Lophocolea fragrans (Moris et De Not.) Gottsche et al. 1845 (313).
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