Stratigraphic data of the Middle â Late Permian on ... - Sylvie Crasquin

Geobios 36 (2003) 533–558 

www.elsevier.com/locate/geobio 

<strong>Stratigraphic</strong> <strong>data</strong> <strong>of</strong> <strong>the</strong> <strong>Middle</strong> – <strong>Late</strong> <strong>Permian</strong> on Russian Platform 

Données stratigraphiques sur le Permien moyen et supérieur de la 

Plate-forme russe 

Vladimir P. Gorsky a , Ekaterina A. Gusseva a,† , Sylvie Crasquin-Soleau b,* , Jean Broutin c 

a All-Russian Geological Research Institute (VSEGEI), Sredny pr. 74, St. Petersburg, 199106, Russia 

b CNRS, FRE2400, université Pierre-et-Marie-Curie, département de géologie sédimentaire, T.15–25, E.4, case 104, 75252 Paris cedex 05, France 

c Université Pierre-et-Marie-Curie, laboratoire de paléobotanique et paléoécologie, IFR101–CNRS, 12, rue Cuvier, 75005 Paris, France 

Received 12 November 2001; accepted 2 December 2002 

Abstract 

This paper presents <strong>the</strong> litho– and biostratigraphic <strong>data</strong> and correlations <strong>of</strong> <strong>the</strong> <strong>Middle</strong> and <strong>Late</strong> <strong>Permian</strong> (Ufimian, Kazanian and Tatarian) 

on <strong>the</strong> Russian Platform. The lithological descriptions and <strong>the</strong> paleontological content (foraminifera, bivalves, ostracods, brachiopods, 

vertebrates, plants and acritarchs) <strong>of</strong> <strong>the</strong> different units are exposed from <strong>the</strong> Barents Sea up to <strong>the</strong> Caspian Sea. 

© 2003 Éditions scientifiques et médicales Elsevier SAS. All rights reserved. 

Résumé 

Cet article présente les descriptions et les corrélations litho– et biostratigraphiques du Permien moyen et supérieur (Ufimien, Kazanien, 

Tatarien) de la Plate-forme russe depuis la mer de Barents jusqu’à la mer Caspienne. Les descriptions lithologiques et le contenu 

paléontologique (foraminifères, bivalves, ostracodes, brachiopodes, vertébrés, plantes et acritarches) des différentes unités sont exposés. 

© 2003 Éditions scientifiques et médicales Elsevier SAS. All rights reserved. 

Keywords: <strong>Stratigraphic</strong> <strong>data</strong>; Correlations; <strong>Middle</strong> and <strong>Late</strong> <strong>Permian</strong>; Russian Platform; Ostracods; Plants 

Mots clés : Données stratigraphiques ; Corrélations ; Permien moyen et supérieur ; Plate-forme russe ; Ostracodes ; Plantes 

1. Introduction 

* Corresponding author. 

E-mail address: crasquin@ccr.jussieu.fr (S. Crasquin-Soleau). 

† Dr. Ekaterina A. Gusseva passed away at <strong>the</strong> end <strong>of</strong> 2000. 

During <strong>the</strong> International Peri-Tethys Programme (1995– 

2000) and for <strong>the</strong> realisation <strong>of</strong> <strong>the</strong> paleogeographic Atlas 

(Dercourt et al., 2000), very efficient collaborations were 

undertaken between <strong>the</strong> French and Russian scientists, and 

particularly between <strong>the</strong> VSEGEI <strong>of</strong> St. Petersburg and <strong>the</strong> 

Pierre et Marie Curie University in Paris. Several proposals 

<strong>of</strong> <strong>the</strong> Peri-Tethys Programme were focussed on <strong>the</strong> <strong>Permian</strong> 

<strong>of</strong> <strong>the</strong> Russian Platform. During <strong>the</strong> last years, both teams 

worked toge<strong>the</strong>r on <strong>the</strong> project <strong>of</strong> publishing in English <strong>the</strong> 

VSEGEI internal report <strong>of</strong> <strong>the</strong> “Decisions <strong>of</strong> <strong>the</strong> interministerial 

meeting on <strong>the</strong> regional stratigraphy concerning <strong>the</strong> 

<strong>Middle</strong> and <strong>Late</strong> Paleozoic <strong>of</strong> <strong>the</strong> Russian Platform: <strong>the</strong> 

<strong>Permian</strong>” (Gorsky and Gusseva, 1990). This report is here 

completed by <strong>the</strong> studies realized during <strong>the</strong> Programme. 

This paper presents <strong>the</strong> stratigraphy <strong>of</strong> <strong>the</strong> <strong>Middle</strong> and <strong>Late</strong> 

<strong>Permian</strong>. The Early <strong>Permian</strong> part, included in <strong>the</strong> Gorsky and 

Gusseva report (1990), will be published later. 

The ostracod fauna is updated by <strong>the</strong> local works in Sou<strong>the</strong>rn 

Cis-Urals and Precaspian Depression (E. Gusseva † and S. 

Crasquin-Soleau in this paper; Molostovskaya, 1997; Molostovsky 

et al., 1997; Kukhtinov and Crasquin-Soleau, 1999; 

Crasquin-Soleau, 2003). The paleobotanical <strong>data</strong> are thoroughly 

vetted by without systematic revision (i.e., synonymy 

is not keeping up to date).All <strong>the</strong> o<strong>the</strong>r fossil <strong>data</strong> are listed as 

<strong>the</strong>y were quoted in <strong>the</strong> original descriptions <strong>of</strong> sections. 

These checklists have historical value. 

©2003Éditions scientifiques et médicales Elsevier SAS. All rights reserved. 

doi:10.1016/S0016-6995(03)00057-3

534 V.P. Gorsky et al. / Geobios 36 (2003) 533–558 

It is not possible to cite here all <strong>the</strong> bibliographic references 

(all in Russian) used to draw up <strong>the</strong> VSEGEI report. We 

refer to this report (Gorsky and Gusseva, 1990). 

During <strong>the</strong> <strong>Middle</strong> – <strong>Late</strong> <strong>Permian</strong> time, almost all <strong>the</strong> 

Nor<strong>the</strong>rn Peri-Tethyan platform is under continental or marginal 

conditions (Gaetani, 2000). The final stages <strong>of</strong> <strong>the</strong> convergence 

between Laurussia, Kazakhstania and Angara result 

in <strong>the</strong> Ural Orogenic belt. Between 20°E (Baltic Sea) and 

60°E (<strong>the</strong> Urals), <strong>the</strong> Russian Platform lies from <strong>the</strong> Barents 

Sea to <strong>the</strong> North up to <strong>the</strong> Caspian Sea to <strong>the</strong> South (Fig. 1). 

Fig. 1. Location <strong>of</strong> <strong>the</strong> geographical reference points 

Fig. 1. Localisation des repères géographiques

V.P. Gorsky et al. / Geobios 36 (2003) 533–558 

535 

2. <strong>Middle</strong> – <strong>Late</strong> <strong>Permian</strong> stratigraphical correlations 

The time slice considered here covered <strong>the</strong> Ufimian, Kazanian 

and Tatarian. The precise correlations with <strong>the</strong> international 

scale (IUGS 2000) are not easy. Indeed, quite all <strong>the</strong> 

Russian series are under continental environments and North 

American series (for <strong>the</strong> <strong>Middle</strong> <strong>Permian</strong>: Guadalupian) and 

<strong>the</strong> Chinese series (for <strong>the</strong> <strong>Late</strong> <strong>Permian</strong>: Lopingian) are 

under marine environments. The main useful tool for <strong>the</strong> 

correlations between <strong>the</strong> different scales could be <strong>the</strong> paleomagnetism 

with <strong>the</strong> reversal between <strong>the</strong> Kiaman Permo- 

Triassic superchron and <strong>the</strong> Illawara mixed polarity superchron. 

The base <strong>of</strong> <strong>the</strong> Illawara mixed zone is recognized in 

<strong>the</strong> Tethyan domain (Haag and Heller, 1991; Baud et al., 

1996). The K/I boundary is also recognized on <strong>the</strong> Russian 

Platform in <strong>the</strong> nor<strong>the</strong>rnmost part <strong>of</strong> western Urals, <strong>the</strong> western 

and central part <strong>of</strong> <strong>the</strong> Syktyokar area, in Perm – Viatka 

area and in Samara area (Gorsky and Gusseva, 1990). Gialanella 

et al. (1997) published developed results obtained in 

Tataria (Kazan area) and presented <strong>the</strong> correlations with <strong>the</strong> 

Tethyan domain. The limit is located at <strong>the</strong> boundary between 

<strong>the</strong> Urzhumian and <strong>the</strong> Severodvinian series i.e. at <strong>the</strong> 

boundary between early and late Tatarian (Table 2)asitwas 

presented by Lozovsky and Yaroshenko (1994) for <strong>the</strong> Moscow 

syncline. The correlations between <strong>the</strong> international 

scale (IUGS 2000) and <strong>the</strong> Russian one are summarized on 

Table 1. 

Two main domains <strong>of</strong> sedimentation are distinguished 

during <strong>the</strong> <strong>Middle</strong> and <strong>Late</strong> <strong>Permian</strong>: <strong>the</strong> Eastern European 

Basin (noted A on Fig. 2) and <strong>the</strong> Western European one 

(noted B on Fig. 2). They are subdivided in different facies 

belts presented on Figure 2. These two domains are heritage 

<strong>of</strong> <strong>the</strong> anterior paleogeographic configuration. The Baltic 

Shield extended eastwards with two major positive structures, 

namely <strong>the</strong> Voronezh High and <strong>the</strong> Ukrainian High. 

These highs separated <strong>the</strong> Eastern European Basin (Western 

Urals, Precaspian and Dniepr-Donetz basins connected toge<strong>the</strong>r) 

from <strong>the</strong> Western one which is related to <strong>the</strong> Polish 

Basin (Gaetani et al., 2000). 

The correlations are presented by a series <strong>of</strong> Tables 

(Tables 2 to 11) from North to South, belt by belt, for <strong>the</strong> 

Eastern European basin (Tables 2 to 10 p.p.) and after for <strong>the</strong> 

Western European Basin (Tables 10 p.p. and 11). The lithological 

descriptions (when <strong>the</strong> room was not sufficient in <strong>the</strong> 

Table an asterisk refers to <strong>the</strong> Table explanation) and <strong>the</strong> 

paleontological content (<strong>the</strong> list is always in <strong>the</strong> Table explanation 

and is refers by a number in <strong>the</strong> Table) are given for all 

<strong>the</strong> intervals. The member “aleurolite” is <strong>of</strong>ten used in <strong>the</strong> 

facies description. “Aleurolite” is equivalent to siltstone in 

Russian literature. 

Table 1 

Correlations <strong>of</strong> International, Tethyan and Russian <strong>Middle</strong> and <strong>Late</strong> <strong>Permian</strong> scales. No vertical scale. 1. After <strong>the</strong> International stratigraphic Chart (IUGS 2000). 

2. After Baud et al. (1993) and Davidov (1996) 

Corrélations des échelles internationales, téthysienne et russe pour le Permien moyen et supérieur. Pas d’échelle verticale. 1.D’après la Charte stratigraphique 

internationale (IUGS 2000). 2.D’après Baud et al. (1993) et Davidov (1996)


Fig. 2. Regional distribution <strong>of</strong> <strong>Middle</strong> and <strong>Late</strong> <strong>Permian</strong> deposits on <strong>the</strong> Russian Platform (on a present day geographical map, identical to Fig. 1. For clarity 

<strong>of</strong> <strong>the</strong> map, a lot <strong>of</strong> geographic names, present on Figure 1 are not reported here). 

Fig. 2. Répartition régionale des dépôts du Permien moyen et supérieur de la Plate-forme russe (sur un fond géographique actuel, identique à celui de la Fig. 1). 

Par souci de clarté, bon nombre de noms géographiques présents sur la Figure 1 ne sont pas reportés ici.


537 

• A. Eastern European sedimentation basin. 

I. Arkhangelsk Belt (Arkhangelsk – Narian-Mar – Adz’Va – Pechora – Ukhta) (Tables 3 and 4); I.1. North <strong>of</strong> Arkhangelsk (lower courses <strong>of</strong> Mezen and 

Pinega rivers, Kouloi river); Kanin Peninsula; I.2. North East <strong>of</strong> Mezen area (Pesa river area; Beluse – Safonovo area); I.3. West <strong>of</strong> Pechora depression 

(Indiga – Izma river – Ukhta area); I.4. Upper course <strong>of</strong> Pechora river; south-eastern part <strong>of</strong> Pechora depression; I.5. North East <strong>of</strong> Pechora depression 

(Narian Mar – Adz’Va – Usinsk area); NE <strong>of</strong> Pechora syneclise; I.6. Tchernishova chain (Adak – Usinsk – Pechora area; Adz’va river). II. Syktyokar Belt 

(South <strong>of</strong> Pechora and Mezen syneclises and Nor<strong>the</strong>rn part <strong>of</strong> Volga and Urals anteclises) (Table 5). II.1.a. South <strong>of</strong> Arkhangelsk area and West <strong>of</strong> Vologda 

area, North East <strong>of</strong> Moscow syneclise, lower course <strong>of</strong> <strong>the</strong> Dvina river; II.1.b. South East <strong>of</strong> Arkhangelsk area and East <strong>of</strong> Vologda area (Sukona river, 

upper courses <strong>of</strong> Pinega and Dvina rivers); South <strong>of</strong> axial part <strong>of</strong> Mezen syneclise and NE <strong>of</strong> Moscow syneclise; II.2. South <strong>of</strong> Komi Republic, North <strong>of</strong> 

Viatka (ex Kirov) area, upper course <strong>of</strong> Vycegda river; II.3. South East <strong>of</strong> Komi Republic, North <strong>of</strong> Perm area, West <strong>of</strong> Ural chain, South <strong>of</strong> Pechora 

syneclise. III. Perm belt (South East <strong>of</strong> Moscow syneclise and North <strong>of</strong> Volga and Ural syneclises) (Table 6); III.1. Volga area between Nijni Novgorod (ex 

Gorky) and Iaroslav (between Volga and Oka rivers, lower course <strong>of</strong> Kostroma river); South West and central Moscow syneclise; III.2. East <strong>of</strong> Unja river, 

North <strong>of</strong> Nijni Novgorod – Tcheboksary area (lower and middle course <strong>of</strong> Unja river and Vetlonga river area), East <strong>of</strong> Moscow syneclise; III.3. Area <strong>of</strong> 

Viatka (upper and middle course <strong>of</strong> Viatka river and Kobra river), structural shoulder <strong>of</strong> Viatka; III.4. Oudmourtia, Perm area; Ufa – Solimansk – Glasov 

area; Kama river, North West <strong>of</strong> Tataria shoulder, shoulders <strong>of</strong> Perm and Bashkiria (with structural shoulder <strong>of</strong> Ufa), Birsk – Kungur depressions. IV. Kazan 

belt (middle part <strong>of</strong> Volga and Ural syneclises) (Table 7); IV.1. South <strong>of</strong> Nijni-Novgorod – Tcheboksary area, Chuvachia (area between Oka and Sviaga 

rivers); IV.2. South East <strong>of</strong> Mari Republic and East <strong>of</strong> Chuvachia, West <strong>of</strong> Tataria, Sviaga river area, West <strong>of</strong> Kazan; IV.3. Tataria (lower course <strong>of</strong> <strong>the</strong> Kama 

and Viatka rivers), shoulder <strong>of</strong> Tataria, North <strong>of</strong> Kuybychev Lake; IV.4. Bashkiria, Sterlitamak – Ufa – Birsk area, lower course <strong>of</strong> Belaya river. V. Samara 

(ex Kuybychev) Belt (South <strong>of</strong> Volga and Ural anticlise) (Table 8). Sou<strong>the</strong>rn part <strong>of</strong> Kuybychev Lake, Samara (ex Kuybychev) – Saratov – Orenburg area, 

Samara river area. VI. Volgograd belt (Precaspian syneclise) (Tables 8 and 9); VI.1. Volgograd – Ural’sk – Orenburg Arc, Obchyi syrt; external part <strong>of</strong> 

western and nor<strong>the</strong>rn Precaspian depression; VI.2. Volgograd – Ural’sk – Orenburg Arc, Obchyi syrt; internal part <strong>of</strong> western and nor<strong>the</strong>rn Precaspian 

depression; VI.3. Volga area near Astrakhan, inner zone <strong>of</strong> sou<strong>the</strong>rn part <strong>of</strong> Precaspian depression; VI.4. West <strong>of</strong> Volga in Astrakhan area, external zone <strong>of</strong> 

sou<strong>the</strong>rn part <strong>of</strong> Precaspian depression; VI.5. Western Kazakhstan area (lower course <strong>of</strong> Ural river), central part <strong>of</strong> Precaspian depression; VI.6. West <strong>of</strong> 

Mugodzar Mounts, eastern part <strong>of</strong> Precaspian depression; VI.7. West <strong>of</strong> Ural chain, near Aktubynsk, NE part <strong>of</strong> eastern part <strong>of</strong> Precaspian depression. VII. 

Mazyr area (lower course <strong>of</strong> Pripyat river), Prypiat depression (Table 10). 

• B. Western European sedimentation basin. 

VIII. Brest area, Brest depression (eastern part) (Table 11); IX. Kaliningrad area (Polish – Lithuanian depression, sou<strong>the</strong>rn part) (Table 11). 

• A. Bassin de sédimentation Est Européen.I. Ceinture d’Arkhangelsk (Arkhangelsk – Narian-Mar – Adz’Va – Pechora – Ukhta) (Tableaux 3 et 4) ; I.1. 

Nord d’Arkhangelsk (cours inférieur des rivières Mezen et Pinega, rivière Kouloi), presqu’île de Kanin ; I.2. Nord Est de Mezen (valléedelaPesa;région 

de Beluse – Safonovo) ; I.3. Ouest de la dépression de Pechora (région de Indiga – rivière Izma – Ukhta) ; I.4. Cours supérieur de la rivière Pechora ; partie 

SudEstdeladépression de Pechora ; I.5. NordEstdeladépression de Pechora (région de Narian Mar – Adz’Va – Usinsk) ; I.6. Chaîne Tchernishova 

(région d’Adak – Usinsk – Pechora; rivière Adz’Va). II. Ceinture de Syktyokar (Sud des dépressions de Pechora et de Mezen et partie nord des antiformes 

de la Volga et de l’Oural) (Tableau 5) ; II.1.a.Régions au Sud d’Arkhangelsk et à l’Ouest de Vologda, Nord Est de la dépression de Moscou, cours inférieur 

de la Dvina ; II.1.b. Régions Sud Est d’Arkhangelsk et Est de Vologda (rivière Sukhona, cours supérieur des rivières Pinega et Dvina) ; Sud de la partie 

axiale de la dépression de Mezen et Nord Est de la dépression de Moscou ; II.2. Sud de la République Komi, Nord de la région de Viatka (ex Kirov), cours 

supérieur de la rivière Vycegda ; II.3. SudEstdelaRépublique Komi, Nord de la région de Perm, Ouest de la chaîne Oural, Sud de la dépression de 

Pechora ; III. Ceinture de Perm (Sud Est de la dépression de Moscou et Nord des dépressions des rivières Volga et Oural) (Tableau 6) ; III.1.Région de la 

Volga entre Nijni Novgorod (ex Gorky) et Iaroslav (entre les rivières Volga et Oka, cours inférieur de la Kostroma) ; Sud Ouest et partie centrale de la 

dépression de Moscou ; III.2. Est de la rivière Unja, Nord de la région de Nijni Novgorod – Tcheboksary (cours inférieur et moyen de la rivière Unja et 

régiondelarivière Vetlonga), Est de la dépression de Moscou ; III.3.Région de Viatka (rivière Koba et cours supérieur et moyen de la Viatka), épaulement 

structural de la Viatka ; III.4. Oudmourtie, région de Perm ; région de Ufa – Solimansk – Glasov ; rivière Kama, Nord Ouet de l’épaulement de Tatarie, 

épaulements de Perm et de la Bashkirie (avec épaulement structural de Ufa), dépressions de Birsk – Kungur. IV. Ceinture de Kazan (cours moyen de la 

Volgaetdépression de la rivière Oural) (Tableau 7) ; IV.1. Sud de la région de Nijni Novgorod – Tcheboksary, Chouvachie (région entre les rivières Oka et 

Sviaga) ; IV.2.SudEstdelaRépublique Mari et Est de la Chouvachie, Ouest de la Tatarie, région de la rivière Sviaga, Ouest de Kazan ; IV.3. Tatarie (cours 

inférieur des rivières Kama et Viatka), épaulement de Tatarie, Nord du lac Kuybychev ; IV.4. Bachkirie, région de Sterlitamak – Ufa – Birsk, cours inférieur 

de la Belaya. V. Ceinture de Samara (ex Kuybychev) (sud de l’antiforme des rivières Volga et Oural) (Tableau 7). Partie Sud du lac de Kuybychev, région 

de Samara (ex Kuybychev) – Saratov – Orenburg, région de la rivière Samara. VI. Ceinture de Volgograd (dépression Pré Caspienne) (Tableaux 8 et 9) ; 

VI.1. Arc de Volgograd – Ural’sk – Orenburg, Obchyi syrt ; partie externe des limites ouest et nord de la dépression Pré Caspienne ; VI.2. Arc de Volgograd 

– Ural’sk – Orenburg, Obchyi syrt, partie interne des limites Ouest et Nord de la dépression Pré Caspienne ; VI.3. Région de la Volga près d’Astrakhan, 

zone interne de la partie Sud de la dépression Pré Caspienne ; VI.4. Ouest de la Volga dans la région d’Astrakhan, partie externe de la région Sud de la 

dépression Pré Caspienne ; VI.5. Ouest du Kazakhstan (cours inférieur de la rivière Oural), partie centrale dépression Pré Caspienne ; VI.6. Ouest des 

Monts Mugodzar, partie Est de la dépression Pré Caspienne ; VI.7. Ouest de la chaîne de l’Oural, près d’Aktubynsk, partie Nord Est de l’Estdela 

dépression Pré Caspienne ; VII.Région de Mazyr (cours inférieur de la rivière Pripyat), dépression de Prypiat (Tableau 10). 

• B. Bassin de sédimentation Ouest Européen.VIII. Région de Brest, dépression de Brest (partie orientale) (Tableau 11) ; IX. Région de Kaliningrad 

(dépression Lithuano-Polonaise, partie sud) (Tableau 11).


Table 2 

Regional stratigraphic subdivisions <strong>of</strong> <strong>the</strong> <strong>Middle</strong> – <strong>Late</strong> <strong>Permian</strong> on <strong>the</strong> Russian Platform 

Subdivisions stratigraphiques régionales du Permien moyen et supérieur de la Plate-forme russe 

• Foraminifera: 1. Digitina rara UCH., Cornuspira megasphaerica GERKE, Nodosaria elabugae TSCHERD., N. suchonensis K.M. MACL, Pseudonodosaria 

lata K.M. MACL., Ps. pygmaeaformis K.M. MACL., Rectoglandulina borealis GERKE. 2. Saccammina rodun<strong>data</strong> K.M. MACL., Nodosaria 

hexagona TSCHERD., N. noinskyi TSCHERD., N. pseudoconcinna K.M. MACL., Lingulonodosaria kamensis K.M. MACL., Pseudonodosaria 

nodosariaeformis K.M. MACL., Ps. lata K.M. MACL., Ps. pygnaeaformis K.M. MACL., Spandelina longissima (K.M. MACL.), S. tzaregradskyi (K.M. 

MACL.), S. fallax (K.M. MACL.) 

• Bivalves: 3. Palaeomutela curiosa Zone. Palaeomutela curiosa AMAL., P. abunda NETSCH., P. parva AMAL., P. oleniana GUS., P. plana AMAL., P. 

keyserlingi AMAL., P. ovalis AMAL., P. inostranzevi AMAL., Oligodontella geinitzi (AMAL.), O. tetraedroides (PLOT.), O.? obliqua (AMAL.), 

Anthraconaia solemyaeformis (NETSCH.), Palaeanodonta parallela AMAL., P. okensis AMAL., P. fischeri (AMAL.), Opokiella tschernyschevi PLOT. 

4. Palaeomutela wohrmani Zone. Palaeomutela wohrmani NETSCH., P. ulemensis GUS., P. numerosa GUS., P. marposadica GUS., P. orthodorsala 

GUS., P. subparallela AMAL., P. verneuili AMAL., P. semilunulata AMAL., P. orthodonta AMAL., P. keyserlingi AMAL., P. ovalis AMAL., 

Oligodontella geinitzi (AMAL.), O. tetraedroides (PLOT.), Anthraconaia verneuili (AMAL.), A. tschuvashica GUS., A. concavocarinata (NETSCH.), 

A. trapezoidalis (AMAL.), A. sambulacovi KUL., Palaeanodonta parallela AMAL., P. okensis AMAL., P. fischeri AMAL., Opokiella tschernyachevi 

PLOT., O. ignatjevi GUS. 5. Palaeomutela vjatkensis Zone. Palaeomutela vjatkensis GUS., P. extensiva GUS., P. visenda GUS., P. corpulenta GUS., P. 

monstruosa GUS., P. celebrata GUS., P. krotowi NETSCH., P. dorati<strong>of</strong>ormis GUS., P. olgae GUS., P. ulemensis GUS., P. verneuili AMAL., P. 

semilunulata AMAL., Prilukiella lata (NETSCH.), P. nitida GUS., P. mirabilis (GUS.), P. pugnatoria GUS., P. rukhini PLOT., P. subovata (JONES), 

Anthraconaia tschuvashica GUS., A. aktubensis GUS., A. castor (EICHW.), A. longissima (NETSCH.), Anthraconauta volgensis GUS., A. uslonensis 

GUS., A. tscherdynzevi GUS., A. iljinskiensis FED., Sinomya gracilenta GUS. 6. Nuculavus trivialis (EICHW.), Schizodus rossicus VERN., 

Siphogramyssia kasanensis (GEIN.), Alula kutorgi (VERN.), Streblochondria sericea (VERN.), Pseudobakewellia ceratophagaeformis NOIN., 

Pseudomonotis permianus MASL., Netschajewia tschernyshewi LICH., Lithophaga consobrina (EICHW.). 7. Polidevcia kasanensis (VERN.), Schizodus 

rossicus VERN., Pseudobakewellia ceratophagaeformis NOIN., Pseudomonotis speluncaria SCHLOTH., P. garfor<strong>the</strong>nsis (KING.), Netachajewia 

globosa (NETSCH.), Wilkingia komiensis (MASL.). 8. Palaeomutela pseudoumbonata Zone. Palaeomutela umbonata FISCH., P. pseudoumbonata 

GUS., P. attenuata GUS., P. olgae GUS., P. stegocephalum NETSCH., P. sintaensis GUS., P. celebrata GUS., P. visenda GUS., P. krotowi NETSCH., P. 

doratiformis GUS., P. quadriangularis (NETSCH.), Concinnella angulata POG., Anthraconaia castor (EICHW.), A. rhomboidea (NETSCH.), A. 

longissima (NETSCH.), A. kamae GUS. 9. Promytilus sicaricus (LUTK. and LOB.), Cyrtorostra laticostata NETSCH., Aviculopecten talis (LUT. and 

LOB.), Astartella raricostata DEMB. 10. Palaeomutela stegocephalum NETSCH., P. attenuata GUS., P. sintanensis GUS., P. ovatiformis GUS., 

Prilukiella alta (POG.), Concinnella concinnaeformis POG., C. angulata POG., Anthraconaia castor (EICHW.), A. rhomboidea (NETSCH.), A. 

longissima (NETSCH.), Anthraconauta subacuta POG., Mrassiella trapezoidalis (KROT.). 

• Ostracods: 11. Suchonellina trapezoidea Zone. Suchonellina trapezoidea (SHARAP.), S. parvaeformis (KASH.), S. fragilis (SHARAP.), S. fragilina 

(BEL.), S. ignatjevi ZEK. and JAN., S. fragiloides ZEK., S. parallela SPIZH., S.? praelongata ZEK., Suchonella cyrta ZEK., S. auriculata (SHARAP.), 

S. typica SPIZH., Darwinuloides svijazicus (SHARAP.), D. tataricus (POSN.). 12. Suchonella futschiki Zone. Suchonellina futschiki (KASH.), S. 

parallela SPIZH., S. cultella MISH., S. daedala MISH., S. digitalis MISH., Prasuchonella stelmachovi (SPISH.), P. ovalis (KOTSCH.), P. sulacensis


539 

KOTSCH., Darwinuloides buguruslanicus KASH., Tscherdynzeviena pulchra (BEL.), T. busulucensis KASH. 13. Paleodarwinula fragiliformis Zone. 

Paleodarwinula fragiliformis (KASH.), P. elongata (LUN.), P. chramovi (GLEB.), P. fragilis var. angusta (KASH.), P. torensis (KOTSCH.), P. kassini 

(BEL.), P. faba (MISH.), P. defluxa (MISH.), P. imposter (MISH.), P. arida (MOLOST.), P.? gariainovi (MOLOST.), P.? tuba (MISH.), P. teodorovichi 

(BEL.), Prasuchonella nasalis (SHARAP.), P. calantarae (BEL.), Darwinuloides tscherdynzevi KASH., <strong>Permian</strong>a obloga (POSN.), Tscherdynzeviana 

aquamosa KOTSCH. 14. Paleodarwinula fainae Zone. Paleodarwinula alexandrinae (BEL.), Paleodarwinula fainae (BEL.), P. irinae (BEL.), P. 

aronovae (BEL.), P. tichonovichi (BEL.), P. inornatinaeformis (BEL.), P. belousovae (KASH.), P. jatskovae (KASH.), P. actubinensis (STAROZH.), P. 

cuneata (KOTSCH.), P. tuimazensis (KOTSCH.), P. pigmaea (STAROZH.), Prasuchonella onega (BEL.), P. vinocurovi BEL., P. tichvinskaja BEL., P. 

belebeica BEL., Darwinuloides sentjakensis (SHARAP.), D. edmistonae BEL., Vymella dobrynini (KASH.), Schneideria kazanica KOTSCH. 14bis. 

Amphissites tscherdynzevi Zone. Pseudoparaparchites furmidabilis SOHN, Amphissites tscherdynzevi POSN., Kirkbya ingeniosa SOHN, Fascianella 

notabilis KOTSCH., F. spinosa KOTSCH., Graphiodactylus uchtensis MART., Cavellina grandis SOHN, C. edmistonae HAR. and L., Healdia oblonga 

KOTSCH., H. postcornuta SOHN, H. pseudosimplex KOTSCH., H. subtriangula KOTSCH., Healdianella vulgaris KOTSCH., Bairdia beedei ULR. and 

B., B. sobolevensis KHIV., Acratia baschkirica KOTSCH., Fabalicypris crespidabilis KOTSCH., Actuaria diffusa KOTSCH., Bairdianella semilunulata 

KOTSCH., Moorea facilis SOHN, Monoceratina fascigiata KOTSCH. 15. Paleodarwinula angusta Zone. Paleodarwinula angusta (MAND.), P. abunda 

(MAND.), P. parphenovae (BEL.), P. ufimica (BEL.), P. inerta (KASH.), P. buraevoensis (PAL.), P. biriensis (PAL.), P. ovataeformis (KASH.), 

Prasuchonella stelmarta KASH., Sinusuella pergrafica MAND., Bairdia kamaensis GUSS., B. dobrjankaensis GUSS., B. plebeia REUSS, B. cultellata 

GUSS., B. cuneiformis GUSS., B. keltmanensis MAND. 

• Bryozoans: 16. Fistulipora dybowskii GOR., Cyclotripa multiformis GOR., Ulrichotripella oculata MOR., Stenopora thuliformis MOR., S. timanensis 

MOR., Tabulipora ordinata MOR., Rhombotripella superanguata MOR., Dyscritella spinigeriformis MOR., Pinegopora delicata SHISHOVA, 

Streblascopora fasciculata (BASSLER), Neorhombopora polita SHISHOVA, Fabifenestella shurae (MOR.), Rectifenestella microretiformis (MOR.), R. 

permulta (MOR.), Parafenestralia arborencens (NETSCH.), Triznella vjatkensis (NIKIF.), Wjatkella wjatkensis (NETSCH.), W. permiana MOR., 

Polyporella magnicava (MOR.), P. soyanensis (MOR.), Polypora baytuganenais SOSK., P. sakanensis MOR., Reteporidra soloduchoi MOR., R. 

quasimicropora MOR., Kingspora parvifenestrata MOR., Timanodictya nikiforovae MOR., T. borealis MOR., Girtyoporina applicata MOR. 

• Brachiopods: 17. Orbiculoidea conincki (GEIN.), Aulosteges fragilis (NETSCH.), A. wangenheimi (VERN.), Cancrinella cancrini (VERN.), Odontospirifer 

subcristatus (NETSCH.), Cleiothyridina pectinifera (SOW.), Beecheria angusta (NETSCH.). 18. Lingula orientalis GOL., Aulosteges 

horrescens (VERN.), A. gigas NETSCH., Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.), Licharewia rugulata (KUT.), L. stuckenbergi 

(NETSCH.), Tumarinia latiareata (NETSCH.), Pinegathris royssiana (KEYS.), Cleiothyridina pectinifera (SOW.), Beecheria netschajewi GRIG., 

Licharewia grewingki (NETSCH.), Permospirifer keyserlingi (NETSCH.), Blasispirifer blasii (VERN.). 

• Vertebrates: 19. Chroniosuchus uralensis TVERD., Ch. paradoxus VJUSCH., Jgosuchus mirabilis (VJUSCH.), J. licharevi RIAB., Dvinosaurus primus 

AMAL., D. purlensis SHISH., Kotlasnia prima AMAL., Karpinskiosaurus secundus (AMAL.), Raphanodon ultimus (TCHUD. and VJUSCH.), 

Chthonosaurus velocidens VJUSCH., Pravoslavlevia parva (PRAVOSL.), Inostrancevia alexandri AMAL., I. uralensis TAT., I. latifrons PRAVOSL., 

Hexacynodon purlensis TAT., Anna<strong>the</strong>rapsidus petri (AMAL.), Nanocynodon seductus TAT., Dvinia prima AMAL., Scutosaurus karpinski (AMAL.), 

Dicynodon trautscholdi AMAL., D. annae (AMAL.), D. amalitzkii SUSH. 20. Chroniosaurus dongusensis TVERD., Raphanodon tverdochlebovae 

IVACH., Microphon exiguus IVACH., Jugosuchus boreus IVACH., J. hartmanni IVACH., Bystrowiana permira VJUSCH., Dvinosaurus primus AMAL., 

D. egregius SHISH., Proburnetia viatkensis TAT., Niukenitia sukhonensis TAT., Sauroctonus progressus (HART.–WEIN.), Noschowhaitsia vjuschkovi 

TAT., Scutosaurus permianus (HART. -WEIN), S. rossicus (HART.–WEIN.), S. utilensis IVACH. and LEB., Deltavjatia vjatkensis (HART.–WEIN.), 

Archosaurus rossicus TAT., Dicynodon scholdi AMAL. 21. Platyops rickardi TWELV., P. stuckenbergi TRAUT., P. watsoni EFR., Melosaurus kinelensis 

VJUSCH., M. paucidens (KONZH.), M. uralensis MEYER, M. vetustus KONZH., Zygosaurus lucius EICHW., Chalcosaurus rossicus MEYER, 

Permotriturus herrei TAT., Brithopus bashkyricus EFR., B. priscus KUT., B. ponderus EFR., Syodon biarmicus KUT., Titanophoneus potens EFR., 

Dolionauriscus adamanteus ORL., D. janshinovi ORL., Notosvodon gusevi TCHUD., Mnemeiosaurus jubilaevi NOP., Admetophoneus kargalensis 

EFR., Deuterosaurus biarmicus EICH., D. gigas EFR., Rhapalodon wangenheimi FISCH., Molybdopygus arcanus TCHUD., Moschops svijagensis 

(RIAB.), Venjukovia prima AMAL., Phreatosaurus bazhovi EFR., Ph. menneri EFR., Phthinoeuchus discors EFR., Porosteognathus efremovi VJUSCH., 

Ennatosaurus tecton EFR., Gecatogomphius kavejevi VJUSCH. and TCHUD., Lanthanosuchus lukjanovae JVACH., L. watsoni EFR., Lanthaniscus 

efremovi JVACH., Macroleter poesicus TVERD. and JVACH., Nycteroleter inaptus EFR., N. kassini TCHUD., Discoaauricus netschajeri RIJAB., 

Nyctibeotus liteus TCHUD., Enosuchus breviceps KONZH., Nyctiphruretus asudens EFR., Biarmica tchudinovi JVACH., Leptoropha talanophora 

(TCHUD.), Mesenosaurus romeri EFR. 22. Platyops stuckenbergi TRAUT., Phthinosaurus borisiaki EFR., Nycteroleter bashkyricus EFR., Rhipaeosaurus 

tricuspidens EFR., Tokosaurus perforatus TVERD. and IVACH., Belebey vegrandis IVACH., B. maxomovi IVACH. and TVERD., Gecatogomphius 

kavejevi VJUSCH. and TVERD., Brithopus priscus KUT., Archaeosydon praeventor TCHUD., Biarmosuchus tener TCHUD., Eotitanosuchus olsoni 

TCHUD., Ivantosaurus ensifer TCHUD., Chthomaloporus lenocinator TCHUD., Estemmenosuchus mirabilis TCHUD., B. uralensis TCHUD., 

Anoplonuchus tenuirostris TCHUD., Parabradysaurus udmurticus EFR., Zopherosuchus luceus TCHUD., Otsheria netzvetaevi TCHUD. 23. 

Phreatophasma aenigmatum EFR., Phreatosuchus qualeni EFR. 

• Flora: 24. Pelliothallites tataricus MEYEN, Protosphagum nervatum NEUB., Rhizinigerites neuburgae MEYEN, Fasciostomia delicata GOM., 

Fefilopteris pilosa GOM., Peltaspermopsis buevichiae GOM. and MEYEN, Tatarina olferievii MEYEN, T. conspiqua MEYEN, T. pinnata MEYEN and 

GOM, T. mira GOM., Stiphorus biceriatus MEYEN, Glossophyllum permiense MEYEN, Salpingocarpus bicornutus MEYEN, S. variabilis MEYEN, 

Phylladoderma aequalis MEYEN, P. anullata MEYEN, P. rastorguevii MEYEN, P. trichophora MEYEN, P. tatarica MEYEN, Doliostomia krassilovii 

MEYEN, Rhaphidopteris kiuntzeliae MEYEN, R. antiqua MEYEN, Permo<strong>the</strong>ca sardykense ZAL., P. striatifera MEYEN and GOM., P. vesicasporoides 

MEYEN, ESAUL. and GOM., P. vittatinifera MEYEN and GOM., Pursongia amalitzkii ZAL., P. angustifolia ZAL., P. beloussovae (RADCZ.) GOM. 

and MEYEN, Cordaites clercii ZAL., Crassinervia subpentegonata KONYKOV, Samaropsis irregularis ZAL., Sashinia aristovensis MEYEN, S. 

borealis MEYEN, Dvinostrobus sagittalis GOM. and MEYEN, Quadrocladus dvinensis MEYEN, Q. borealis MEYEN, Q. schweitzerii MEYEN. 25. 

Sphenophyllum stoukenbergii SCHMALH., Phylladoderma muravievii BOGOV, Nucicarpus piniformis NEUB. 26. Signacularia noinskii ZAL., 

Sphenophyllum stoukenbergii SCHMALH., Paracalamites striatus (SCHMALH.) ZAL., Pecopteris helenaeana ZAL., Odontopteris rossica ZAL., 

Syniopteris nesteronkoi ZAL., Rhaphidopteris praecursoria MEYEN, Cordaites candalepensis (ZAL.) RADCZ., Phylladoderma meridionalis MEYEN, 

P. fefiloviana BOGOV, Permo<strong>the</strong>ca sardykensis ZAL., Nucicarpus piniformis NEUB. 27. Paracalamites striatus (SCHMALH.) ZAL., Callipteris erosa 

(MORRIS) ZAL., Psygmophyllum (Syniopteris) expansum BRONGN., P. cuneifolium KUTORGA, Odontopteris rossica ZAL., Phylladoderma spinosa 

BOGOV, Permo<strong>the</strong>ca sardykensis ZAL. 28. Intia variabilis NEUB., Viatcheslavia vorcutensis ZAL., Sylvella alata ZAL., Samaropsis vorcutana 

TSCHIRK. 

• Miospores and Acritarchs: 29. Vitreisporites pallidus Zone (J 2 ). Calamospora hartungiana BALME, Punctatisporites sp., Apiculatisporites hispidus 

ANDR., Granizonospora granifera LUBER, Osmundacidites sp., Nevesisporites sp., Lycospora papullata (SMIRN.) WAR., Cirratriradites sp.,


Azonaletes sp., Ginkgocycadophytus sp., Vittatina cincinneta LUBER, V. costabilis WILS., Cladaitina sp., Vesicaspora schemeli KL., Platysaccus major 

(LUBER) WAR., Lueckisporites virkkiae POT. and KL., Vitreisporites pallidus (REISSINGER) NILSSON, Vitreisporites sp., Gnetaceaepollenites sp., 

Limitisporites ambiguus (ZAUER) WAR., Striatopodocarpites tojmensis SED., S. rarus BHARD. and SAL., Striatopiceites suchonensis SAD., 

Striatoabieites sp., Striatolebacheites sp., Vesicaspora ex gr. magnalis ANDREYEVA, Protohaploxypinus limpidus BALME. 30. Taxodiacites permianus 

Zone (J 1 ). Calamospora sp., Trachytriletes sp., Cyclogranisporites sp., Apiculatisporites hispidus ANDR., Cirratriradites sp., Lycospora papullata 

(SMIRN.) WAR., Azonomonoletes vulgaris IBR. f. minor LOOSE, Densoisporites sp., Osmundacidites sp., Raistrickia sp., Azonaletes sp., Vittatina 

costabilis WILS., V. elegans ZAUER, V. striata f. major LUBER, V. lata WILS., V. crassa ZAUER, Florinites sp., Cordaitina sp., Falcisporites sp., 

Caytoniapollenites pallidus LESCHIK, Vesicaspora sp., Taxodiacites permianus ZAUER, Striatopodocarpites tojmensis SED., S. rarus (BHARD. and 

SAL.), Striatopiceites suchonensis SED., Taeniasporites sp., Hamiapollenites sp. 31. Lueckisporites virkkiae Zone (I 2 ). Calamospora sp., Apiculatisporites 

sp., Trachytriletes sp., Granulatisporites graniferus (K.-M.) WAR., Osmundacidites sp., Raistrickia cardi<strong>of</strong>ormis (K.-M.) WAR., Neoraistrickia 

selaginelliformis SAMOIL., Leiotriletes inderjensis (MAL.) WAR., Kraeuselisporites sp., Azonaletes sp., Ginkgocycadophytus sp., Vittatina 

cincinnata LUBER, V. persecta ZAUER, Florinites sp., Cordaitina sp., Lueckisporites virkkiae POT. and KL., Taxodiacites sp., Vesicaspora ovata 

BALME and HEN., Vestigisporites sp., Falcisporites zapfei (POT. and KL.), Limitisporites sp., Striatopodocarpites tojmensis SEd., Striatopinites 

dvinensis SED., S. suchonensis SED., Striatohaplopinites perfectus NAUM., Striatodiplopinites lata SCHATK., Taeniasporites hexagonalis JANS. 32. 

Striatopodocarpites tojmensis Zone (I 1 ). Trachytriletes punctalatus NAUM., Granulatisporites microgranifer IBRAH., Cyclogranisporites osmundae 

SAMOIL., Spinosisporites spinosus WALTZ, Raistrickia cardi<strong>of</strong>ormis (K.-M.) WAR., R. cornuta (ANDR.) DR., Granulatisporites parviverrucosus 

WALTZ, Azonaletes sp., Vittatina striata LUBER, V. vittifer LUBER, V. persecta ZAUER, Cordaitina sp., Platysaccus sp., Vesicaspora schemeli KL., V. 

ovata BALME and HEN., Falcisporites zapfei POT. and KL., Alisporites sp., Platysaccus major LUBER, Limitisporites sp., Chordasporites sp., 

Striatopodocarpites tojmensis SED., Striatopinites dvinensis SED., Striatolebacheites sp., Lueckisporites sp., Taxodiacidites sp., Hamiapollenites sp., 

Vitreisporites sp., Taeniasporites hexagonalis JANS. 33. Granizonospora vulgaris Zone (H 2 ). Calamospora sp., Spinosisporites parvispinus LUBER, 

Granulatisporites parviverrucosus WALTZ, Granizonospora vulgaris NAUM., Verrucosisporites sp., Cirratriradites procumbens LUBER, C. ornatus 

LUBER, Vittatina sp., Cordaitina sp., Vesicaspora sp., Platysaccus sp., Florinites luberae SAMOIL., Crusisaccites sp., Hamiapollenites tractiferinus 

SAMOIL., Striatodiplopinites elongatus LUBER, Taeniasporites sp. 34. Cirratriradites ornatus Zone (H 1 ). Calamospora sp., Cyclogranisporites 

gibberuliformis K.-M., C. polypyrenus LUBER, Granulatisporites gibbosus IBR., G. parviverrucosus WALTZ, Granizonospora vulgaris NAUM., G. 

granifera LUBER, Spinosisporites rectispinus LUB., Acanthotriletes osmundae SAMOIL., Cirratriradites ornatus LUBER, C. procumbens LUBER, 

Lycospora variabilis JANS., Remysporites psilopterus LUBER, Ginkgocycadophytus sp., Vittatina striata LUBER, V. vittifer LUBER, Florinites luberae 

SAMOIL., Crucisaccites ornatus SAMOIL., Alisporites sp., Limitisporites monstruosus LUBER, Striatohaplopinites latissimus LUBER, Hamiapollenites 

sp., Striatolebachiites sp. 

Table 3 

Eastern European basin, Arkhangelsk belt, part 1 

Bassin Est-Européen, ceinture d’Arkhangelsk, première partie


541 

35. Bivalves: Oligodontella geinitzi (AMAL.), Anthraconaia(?) verneuili AMAL.; Ostracods: Paleodarwinula chramovi (GLEB.), P. elongata (LUN.), 

Suchonellina futschiki (KASH.), S. inornata SPIZH., S. inornata var. macra (LUN.), S. parallela SPIZH., S. impostor (MISH.), Prasuchonella nasalis 

(SHAR.), P. stelmachovi (SPIZH.); Flora s.l.: Ginkgocycadophytus sp., Vittatina costabilis WILS., V. cincinnata LUBER, V. striata LUBER, Cordaitina sp., 

Podocarpites major (LUBER) WAR., Limitisporites monstruosus (LUBER) HART., Piceites sublevis (LUBER) HART., Vesicaspora schemeli KL., 

Vitreisporites sp., Striatopodocarpites nudus (LUBER) ZAUER, S. tojmensis SED., S. rarus (BHARD. and SAL.) WAR., Striatolebachiites sp. 

36. Aleurolites, red marls – Aleurolites et marnes rouges: Ostracods: Paleodarwinula spizharskyi (POSNER), P. teodorovichi (BEL.), P. fragiliformis 

(KASH.). 

37. Aleurolites, marls, dolomitised marls – Aleurolites, marnes, marnes dolomitisées; aleurolites, marnes, marnes dolomitisées. Ostracods: Paleodarwinula 

aff. perlonga (SHAR.), <strong>Permian</strong>a elongata (POSNER), P. oblonga (POSNER); Bivalves: Prilukiella(?) mirabilis GUS. 

38. Aleurolites, sandstones, red marls – Aleurolites, grès marnes rouges: Ostracods: Amphissites tscherdynzevi POSNER, Moorea facilis SOHN, Cavellina 

edminstonae (HAR. and LALL.). 

39. Western part: red and thin aleurolites with gypsum; eastern part: grey thin aleurolites – Partie occidentale: aleurolites rouges et peu épaisses avec du 

gypse; partie orientale: aleurolites grises et peu épaisses. Vertebrates: Nycteroletes inertus EFR., Nyctiphuretus asudens EFR., Mesenosaurus romeri EFR., 

Microphon poesicus TVERD. and IVACH., Lanthaniscus efremovi IVACH. 

40. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Trachytriletes sp., Granulatisporites sp., Cyclogranisporites sp., Acanthotriletes 

sp., Cirratriradites ornatus (LUBER) HART., Azonomonoletes vulgaris IBR., Azonaletes microdyctius LUBER, Vittatina striata LUBER, V. cincinnata 

LUBER, V. costabilis WILS., V. lata WILS., V. crassa ZAUER, Alisporites ovatus JANS., Protopodocarpus alatus (LUBER) SAM., Protohaploxypinus sp., 

Striatopodocarpites nudus (LUBER) ZAUER, S. tojmensis SED., S. rarus BHARD. and SAL., Striatopinites trivialis (SCHATK.) SIV., Taeniaesporites tenor 

(ZAUER) WAR. 

41. Aleurolites, calcareous sandstones, marls, argillites, limestones – Aleurolites, grès calcaires, marnes, argilites, calcaires. Bivalves: Netschajewia 

pallasi VERN., Schizodus sp., Plurophoria simplex REIS., Nuculana wimensis KUL., Myonia komiensis (MASL.), Oriocrasatella plana (GOLOV.); 

Ostracods: Fascianella plotnikovi MAND., Pseudobythocypris pediformis KNIGHT, Ps. cf. osagensis KELLETT, Moorea sp., Cavellina sp. 

42. Foraminifera: Pseudonodosaria lata K.M. MACL., Cornuspira microsphaerica GERKE, Memigordius planispiralis K.M. MACL.; Bivalves: 

Pseudomonotis permianus MASL., Pseudobakewellia antiquaeformis NOIN., Nuculavus wymensis (KEYS.); Bryozoans: Dyscritella spinigeriformis MOR., 

Rhombotrypella summa MOR.; Ostracods: Healdia pseudosimplex KOTSCH., Cavellina grandis SCHNEID., Actuaria seconda KOTSCH.; Brachiopods: 

Lingula orientalis GOL., Rhynchopora geinitziana VERN., Pinegathyris royssiana (KEYS.), Dielasma elongeta (SCHLOTH.), Amblypterus tuberculatus 

(EICHW.); Flora s.l.: Calamospora rotunda (NAUM.) WAR., Trachytriletes punctatus (WALTZ) LUBER, Granulatisporites granifer (K.-M.) WAR., 

Cyclogranisporites osmundae (SAMOIL.) WAR., Raistrickia cardi<strong>of</strong>ormis (LUBER) WAR., Azonaletes plicatus WALTZ, Ginkgocycadophytus sp., Vittatina 

vittifer LUBER, V. persecta ZAUER, Cordaitina sp. 

43. Upper part: marls, limestones, aleurolites – Partie supérieure : marnes, calcaires, aleurolites. Foraminifera: Hyperammina delicatula GERKE, 

Nodosaria suchonensis K.M. MACL., Pseudonodosaria nodosariaeformis K.M. MACL.; Ostracods: Moorea facilis SCHNEID., Healdia subtriangula 

KOTSCH., Healdianella vulgaris KOTSCH. Flora s.l.: Aulosteges horrescens (VERN.), Rhynchopora variabilis STUCK., Cleiothyridina pectinifera (SOW.). 

Lower part: argillaceous biogenic limestones, aleurolites, marls, argillites; in western part, biohermes – Partie inférieure : calcaires biogéniques argileux, 

aleurolites, marnes, argilites ; biohermes dans la partie occidentale. Foraminifera: Verneuilloides andrei UCH., Nodosaria hexagona TSCHERD., N. 

pseudoconcinna K.M. MACL., Pseudonodosaria lata K.M. MACL.; Ostracods: Kirkbya ingeniosa SCHN., Amphissites tscherdynzevi POSNER, Healdia 

postcornuta SCHN. Brachiopods: Licharewia stuckenbergi (NETSCH.), L. latiareata (NETSCH.), Cancrinella cancrini VERN. 

44. Foraminifera: Saccamina rotunda K.M. MACL., Nodosaria hexagona TSCHERD., N. elabugae TSCHERD., Lingulonodosaria kamensis K.M. 

MACL., Frondicularia longissima (K.M. MACL.). Ostracods: Healdia postcornuta SOHN, H. pseudosimplex KOSTCH. Brachiopods: Lingula orientalis 

GOL., Globiella tschernyschevi (NETSCH.), Schisodus rossicus VERN. Flora s.l.: Phylladoderma arberi ZAL., Pecopteris synica ZAL., Meristophyllum 

sojamaeanum ZAL. 

45. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Cyclogranisporites osmundae (WALTZ) WAR., Spinosisporites spinosus 

(WALTZ), Raistrickia cardi<strong>of</strong>ormis K.M., Lycospora variabilis JANS., Entylissa capereta LUBER, Vittatina cincinnata LUBER, V. striata LUBER, 

Cordaitina rugulifera (LUBER) SAMOIL. 

46a. Flora s.l.: Netschajewia tschernyschewi LICH., Moorea facilis SOHN, Paleodarwinula jatskovae (KASH.), P. fainae (BEL.), P. trita (PAL.), P. irenae 

(BEL.), Vymella dobryinini (KASH.), Trachytriletes punctulatus NAUM., Cyclogranisporites osmundae (SAMOIL.) WAR., C. gibberuliformis (K.-M.) WAR. 

46b. Ostracods: Paleodarwinula procera (MAND.), P. Angusta (MAND.), P. Abunda (MAND.), P. inerta (KASH.), P. sobela (KASH.), P. sinedorensis 

(KASH.), P. pergusta (KASH.). Brachiopods; Lioes<strong>the</strong>ria simplex MOL., Pseudoes<strong>the</strong>ria mezeniana MOL., Ps. borealis MOL., Notocrypta timanica MOL. 

Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Trachytriletes punctalatus NAUM., Cyclogranisporites gibberuliformis K.-M., 

Granulatisporites gibbosus IBR., Spinosisporites rectispinus LUBER, Remysporites psilopterus LUBER, Cirratriradites ornatus (LUBER) HART., C. 

procumbens (LUBER) SIV., Lycospora variabilis JANS., Vittatina vittifer LUBER, V. striata LUBER, Florinites luberae SAMOIL., Alisporites ovatus JANS. 

46c. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Trachytriletes asperatus (LUBER) ISCH., Cyclogranisporites gibberuliferus 

K.-M., Raistrickia cardi<strong>of</strong>ormis (K.-M.) WAR., Azonaletes levis LUBER, Vittatina vittifer LUBER, V. striata LUBER, Florinites luberae SAMOIL., 

Cordaitina vera (SADK.) SIV., C. subrotata (LUBER) SAMOIL., Protopodocarpites alatus (LUBER) SAMOIL., Striatodiploxypinites striatus (LUBER) 

WAR., Striatopinites trivialis (SCHATK.) SIV. 

47. Ostracods: Paleodarwinula perlonga (SHARAP.), Suchonellina inornata (SPIZH.), S. parallela (SPIZH.) 

48a. Brachiopods: Cancrinalia cancrini (VERN.). Bivalves: Promytilus retusus CHRON., Permophorus angustus MUROM., Stutchburia pallasi (VERN.), 

Myonia komiensis (MASL.), Praeunduloyva leonidi MUROM. 

48b. Bivalves: Aviculopecten wymensis LICH., Anisotrypella borealis MOR., Discritella spinigeriformis MOR., Paralioclema multispinosum MOR., 

Wyatkella permiana MOR.; Brachiopods: Aulosteges longus NETSCH., Cancrinella cancrini (VERN.), Ulugkemia leonidi NOV., U. petri NOV., Limnadia 

kastiatum NOV.


49. Foraminifera: Nodosaria elabugae TSCHERD., N. suchonensis K.M. MACL. Brachiopods: Discritella spinigeriformis MOR., Anisotrypella borealis 

MOR., Fistulipora dybowskyi GOR., Cancrinella cancrini (VERN.), Permospirifer keyserlingi (NETSCH.), Licharewia sp., Rhynchopora geinitziana 

(VERN.), Beecheria netschajewi GRIG.; Flora s.l.: Stepanovites meyeni KOZ. and MOV. 

50. Bivalves: Prilukiella mirabilis (GUS.), Maturipina kholunicensis GUS. Ostracods: Paleodarwinula fragiliformis KASH., Sinusuella ignota SPIZH., 

Placidea lutkevichi (SPIZH.), Suchonellina futschiki (KASH.). 

51. Bivalves: Prilukiella mirabilis (GUS.), P. nitida GUS., Palaeanodonta subcastor AMAL. Ostracods: Paleodarwinula belousovae (KASH.), Vymella 

dobrynini (KASH.), Placidea lutkevichi (SPIZH.). Flora s.l.: Callipteris (Feonia) aequabilis MEYEN, Phylladoderma meridionalis MEYEN, Rhaphidopteris 

sp., Slivkovia sp., Aequistomia sp.; Miospore complex I 2 (see Table 2). 

52. Green to grey argillites, marls, aleurolites with coal levels; eastern part: red aleurolites – Argilites vertes à grises, marnes, aleurolites avec des niveaux 

de charbon; dans la partie orientale: aleurolites rouges. Foraminifera: Pseudonodosaria ventrosa SCHL., Frondicularia pseudotriangularis GERKE. 

Bivalves: Abiella subovata (JON.), Anthraconauta pseudophillipsi FED., A. iljinnklensis FED., A. globosa FED., Syniella (Pseudosinomya) bella GUS.. 

Ostracods: Paleodarwinula jatskovae (KASH.), P. irinae (BEL.), P. pergusta (KASH.), P. belousovae (KASH.), P. borealis (KASH.), Prasuchonella 

tichwinzkajae (BEL.), Placidea petchorica (SCHN.), Volganella laevigata SCHN. 

53. Red argillites with intercalations <strong>of</strong> aleurolites and sandstones – Argilites rouges avec intercalations d’aleurolites et grès. Ostr Paleodarwinula abunda 

(MAND.), P. angusta (MAND.), P. sobela (KASH.), Prasuchonella scolia (MAND.), Sinusuella pergraphica MAND. 

54. Red argillites, marls, aleurolites – Argilites rouges, marnes et aleurolites. Ostracods: Paleodarwinula angusta (MAND.), P. inerta (KASH.), P. procera 

(MAND.), P. burajevoensis (PAL.) 

55. Argillites and sandstones with intercalations <strong>of</strong> limestones – Argilites et grès à intercalations de calcaires. Ostracods: Paleodarwinula sobela (KASH.), 

P. lancetiformis (KASH.), Praesuchonella stelmarta KASH. 

56. Ostracods: Paleodarwinula belousovae (KASH.), Vymella dobrinini KACH., Placidea lutkovichi (SPIZH.). Brachiopods: Lingula orientalis GOL.; 

Flora s.l.: Bardocarpus superus NEUB., Miospore complex I 2 (see Table 2). 

57. Bivalves: Astartella permocarbonica TSCH.. Brachiopods: Leaia cargalensis NETSCH., Lingula orientalis GOL., L. hyperborea IFAN., L. credneri 

GEIN.. Ostracods: Paleodarwinula pergusta (KASH.), P. jatskovae (KASH.), Placidea mesotuberculata SCHN., Volganella laevigata SCHN., Cavellina 

edmistonae HARRIS and LALICKER, C. grandis KOTSCH.; Miospore complex I 1 (see Table 2). 

58. Foraminifera: Frondicularia tsaregradski K.M. MACL., F. dilema GERKW, F. geinitziniformis (K.M. MACL.), F. longissima (K.M. MACL.), 

Lingulonodosaria cf. farcimeniformis (K.M.–MACL.), Anthraconaia cf. castor (EICHW.), Nuculavus cf. wymensis KEYS.. Bivalves: Astartella sp. Brachiopods: 

Lingula orientalis GOL., L. credneri GOIN., Cancrinella cancriniformis TSCHERN.. Ostracods: Healdia postcornuta SCHN., H. enervis MART., H. 

kozhimica GUSS., Cavellina permiana KOTSCH., Graphiodactylus uchtensis MART., Monoceratina cf. kungurica (MART.), Bairdia sp., Placidea lutkevichi 

(SPIZH.); in <strong>the</strong> lower part, miospore complex H 2 and H 1 (see Table 2). 

Table 4 

Eastern European basin, Arkhangelsk belt, part 2 

Bassin Est-Européen, ceinture d’Arkhangelsk, deuxième partie


543 

59. Bivalves: Anthraconauta moltschanovi BETEKHT., A. degeniaensis BETEKHT., Concinnella gravis PAP, C. cf. concinna (JONS.). Flora s.l.: 

Protosphagum nervatum NEUB., Crassinervia ovifolia NEUB., Rhipidopsis ginkgoides SCHMALH., Phylladoderma arberi ZAL., Bardocarpus superus 

NEUB., Nucicarpus piniformis NEUB.; Miospore complex I 2 (see Table 2) 

60. Bivalves: Palaeomutela visenda GUS., P. monstruosa GUS., P. krotowi NETSCH., Anthraconauta volgensis GUS., A. gibbosa BETEKHT., Peraborella 

fedotovi (KAN.), Syniella bella (GUS.), S. angusta KAN. Ostracods: Paleodarwinula irinae (BEL.), P. chramovi (GLEB.), P. fainae (BEL.), Placidea 

lutkevichi (SPIZH.), P. petschorica (SCHN.). Flora s.l.: Intia vermicularia NEUB., Odontopteris rossica ZAL., Callipteris adzvensis ZAL., Cordaites clercii 

(ZAL.) RADCZ., C. candalepensis ZAL., Rufloria synensis (NEUB.) MEYEN, Rhaphidopteris praecursoria MEYEN, Phylladoderma arberi ZAL., 

Samaropsis extensa NEUB., S. postfrigida DOMBR., Bardocarpus superus NEUB., Permo<strong>the</strong>ca sardykensis ZAL., Nucicarpus piniformis NEUB.; Miospore 

complex I 2 . (see Table 2) 

61. Bivalves: Anthraconaia rhomboida (NETSCH.), A. oblonga (BEL.), Mrassiella komiensis KAN., Concinnella angulata POG., Abiella kolvae KAN., 

Palaeomutela cf. alta POG. Ostracods: Paleodarwinula cf. martjevi (PAL.), P. cf. lubimovae (KASH.), Vymella timanica (KASH.), Sinusuella pergraphica 

MAHD. Flora s.l.: Intia variabilis NEUB., Kosjunia polyedra FEF., Rufloria synensis (NEUB.) MEYEN, Phylladoderma sp., Zamiopteris neuburgiana 

MEYEN, Samaropsis niamdensis NEUB., Bardocarpus superus NEUB.; Miospore complex I 1 – H 2 (see Table 2) 

62. In <strong>the</strong> upper part – Dans la partie supérieure. Bivalves: Wilkingia komiensis MASL., Nuculavus wymenzis (KSYS). Brachiopods: Lingula ex gr. 

orientalis GOL. Ostracods: Vymella timanica (KASH.), V. petschorica (KASH.), Placidea cf. mesotuberculata SCHN. In <strong>the</strong> lower part – Dans la partie 

inférieure. Foraminifera: Protonodosaria praecursor RAUS., Nodosaria lata SOSSIP., Pseudonodosaria nodosariaeformis K.M. MACL., Frondicularia 

planilata GERKE, F. bella GERKE. Brachiopods: Dentalina praenuntia GERKE, Rhynchopora cf. nikitini TSCH., Lissochonetes sp. Flora s.l.: Streblopteris 

ex gr. pusilla SCHLF., S. sericea VERN., Laevidentalium sp. In all <strong>the</strong> series – Dans toute la série. Bivalves: Astartella permocarbonica TACHERN., 

Nuculopsis ellipticus GUS. and POG., Intia variabilis NEUB., Kosjunia retusa FEF., Junjagia glottophylla NEUB. Flora s.l.: Vorcutannularia plicata (POG.) 

NEUB., Sphenophyllum comiense ZAL., Rufloria recta (NEUB.) MEYEN, Miospore complex H 1 (see Table 2) 

63. Ostracods: Paleodarwinula borealis (KASH.), P. procera (MAND.), P. pseudopetschorica (KASH.), Sinusuella pergraphica MAND. Foraminifera: 

Palaeomutela sp., Palaeanodonta vorcutica POG., P.exgr.castor (EICHW.), Concinnella ex gr. concinna JONES. Flora s.l.: Cordaites clercii (ZAL.) RAD., 

Rhipidopsis sp., Phylladoderma arberi ZAL., Bardocarpus superus NEUB., B. synensis DOMB.; in <strong>the</strong> upper part: Miospore complex I; in <strong>the</strong> middle part: 

Miospore complex I 2 (see Table 2) 

64. In <strong>the</strong> upper and middle parts, dans les parties supérieure et moyenne: Bivalves: Anthraconaia longissima (NETSCH.), A. pseudocastor (BETEKHT.), 

Abiella subovata (JON.), Mransiellina sp. Foraminifera: Concinnella solita PAP. Brachiopods: Lingula sp., Aulosteges sp., Cancrinella ex gr. cantrini VERN., 

Waagenoconcha cf. purdoni DAV., Lissochonetes aff. rotundatus (TOULA). Flora s.l.: Intia sp., Comia peraborensis ZAL., Lepeophyllum trigonum NEUB., 

Cordaites clercii (ZAL.) RADCZ., Rufloria sp., Miospore complex I 1 (see Table 2) 

65. In <strong>the</strong> lower part – Dans la partie inférieure. Bivalves: Anthraconaia oblonga (BEL.), Mrassiella trapezoidalis (KROT.). Ostracods: Graphiodactylus 

uchtensis Mart.; Brachiopods – brachiopodes: Swalbardia capitolina TOULA, Rhynchopora geinitziana VERN., Lissochonetes sulaensis V. BARCH., 

Megousia kulikii FRED. Flora s.l.: Jakovlevia mammata KEYS., Viatscheslavia sp., Viatscheslaviophyllum vorcutense NEUB., Sphenophyllum comiense 

TSCHIRK., Samaropsis vorcutana TSCHIRK., S. striata NEUB.; Miospore complex H 2 (see Table 2) 

66. Foraminifera: Protonodosaria proceraformis GERKE, Nododosaria lata SOSSIP., N. cassiaformis IGON., Rectoglandulina pseudoborealis SOSSIP., 

Frondicularia sectorialis VOR., F. subtilistriata GORKEL, F. bajcurica SOSSIP. Bivalves: Polidevcia cf. formosa DEMB., Nuculopsis wymensis (KEYS.), 

Nuculana sinuata (DEMB.). Ostracods: Healdia ovataeformis SCHN. Brachiopods: Megousia kulikii FRED. Flora s.l.: Vorcutannularia plicata (POG.) 

NEUB., Sphenophyllum thonii MAHR., Samaropsis subelegans NEUB., Sylvella aleta ZAL., Miospore complex H 1 (see Table 2) 

67. Flora s.l.: Protosphagum nervatum NEUB., Crassinervia ovifolia NEUB., Rhipidopsis ginkgoides SCHMALH., Phylladoderma arberi ZAL., 

Bardocarpus superus NEUB. 

68. Foraminifera: Palaeomutela visenda GUS., P. petschorica KAN. Bivalves: Syniella bella (GUS.), Anthraconauta volgensis GUS. Ostracods: 

Paleodarvinula irenae (BEL.), P. fainae (BEL.), P. chramovi (GLEB.), Placidea lutkevichi (SPIZH.), P. petchorica (SCHN.). Flora s.l.: Intia vermicularis 

NEUB., Comia pereborensis ZAL., Callipteris adzvensis ZAL., Cordaites clercii (ZAL.) RADCZ., Rufloria synensis (NEUB.) MEYEN, Rhaphidopteris 

praecursoria MEYEN, Phylladoderma arberi ZAL., Samaropsis extensa NEUB., S. postfrigida DOMB., S. pajchoica DOMB., Bardocarpus superus NEUB.; 

Miospore complex I 2 (see Table 2) 

69. In <strong>the</strong> lower part: Bivalves: Khosedaella alta POG., Concinnella concinnaeformis POG., C. exgr.concinna (JONES), Palaeomutela cf. alata POG., 

Oligodontella ex gr. geinitzi AMAL. Ostracods: Paleodarwinula cf. martjevi (PAL.), P. cf. lubimovae (KASH.), Vymella timanica (KASH.), Sinusuella 

pergraphica MAND. Flora s.l.: Intia variabilis NEUB., Kosjunia polyedra FEF., Todites sp., Oligocarpia permiana FET., Rufloria synensis (NEUB.) MEYEN, 

Phylladoderma sp., Zamiopteris neuburgiana MEYEN, Samaropteris niamdensis NEUB., Bardocarpus superus NEUB.; Miospore complex H 2 –I (see Table 2) 

70. In <strong>the</strong> upper part; dans la partie supérieure: Bivalves: Wilkingia komiensis MASL., Nuculavus wymensis (KEYS.). Ostracods: Vymella timanica KASH., 

Sinusuella pergraphica MAND. In <strong>the</strong> middle part – Dans la partie moyenne. Foraminifera: Protonodosaria praecursor RAUS., Nodosaria lata SOSS., 

Frondicularia planilata GERKE. Bryozoans: Rhinchopora nikitini TSCH. In <strong>the</strong> lower part – Dans la partie inférieure: FLORA s.l.: Intia variabilis NEUB., 

Junjagia glottophylla NEUB., Vorcutannularia plicata (POG.) NEUB., Rufloria recta (NEUB.) MEYEN. 

71. Foraminifera: Palaeomutela talbeica POG., Anthraconauta moltachanovi BETCKHT., Adzvaella tatarica (KAN.), Concinnella buredanica POG. 

Brachiopods: Lioes<strong>the</strong>ria raslovensis NOVOJ., Pseudoes<strong>the</strong>ria dvinaensis MOLIN., Ps. nordvikensis (NOVOJ.), Ps. shujaensis NOVOJ., Eulimnadia lima 

NOVOJ., E. pausa NOVOJ. Ostracods: Paleodarwinula chramovi (GLEB.), P. fragiliformis (KASH.), P. <strong>the</strong>odorovichi (BEL.), Prasuchonella stelmachovi 

(SPIZH.). Flora s.l.: Polyssaievia deflexa NEUB., Viatscheslavia chejagensis NEUB., Cladophlebis aff. nystroromii HALLE, Pecopteris anthriscifolia 

(GOEPP.) ZAL. f. adzvensis FET., Callipteris elegans (SCHMALH) MEYEN, Phylladoderma arberi ZAL., Sphenobaiera sp., Glossophyllum sp., Sporophyllites 

petschorensis (SCHMALH.) Meyen; in <strong>the</strong> upper part, Miospore complex J; in <strong>the</strong> lower part Miospore complex I (see Table 2). In <strong>the</strong> lower part; dans la 

partie inférieure: Bivalves: Palaeomutela krotowi NETSCH., P. visenda GUS., P. petschorica KAN., Pereborella fedotovi (KAN.), Anthraconauta gibbosa 

BETEKHT., Abiella subovata (JON.) 

72. Bivalves: Palaeomutela celebrata GUS., Concinnella pajchoica POG. Ostracods: Paleodarwinula belousovae (KASH.). In <strong>the</strong> upper part; dans la 

partie supérieure. Flora s.l.: Paikhoia tschernovii ZAL., Astero<strong>the</strong>ca? pluriseriata FEF., Pecopteris obtusa RADCZ., Syniopteris nesterenkoi ZAL., 

Phylladoderma arbori ZAL., Pereborites rarinervis ZAL., Bardocarpus superus NEUB., Nucicarpus piniformis NEUB.; Miospore complex I (see Table 2). In


<strong>the</strong> lower part; dans la partie inférieure. Bivalves: Anthraconauta uralica KAN., Abiella kolvae KAN., Concinnella anomala SIB. and PAP. Flora s.l.: 

Viatscheslavia vorcutensis ZAL., Sphenophyllum comiense TSCHIRK., Pecopteris compta RADCZ., Compsopteris adzvensis ZAL., Callipteris adzvensis 

ZAL., Rufloria synensis (ZAL.) MEYEN, Bardocarpus superus NEUB., Samaropsis vorcutana TSCHIRK.; Miospore complex H 2 (see Table 2) 

73. Bivalves: Anthraconauta subovalis POG., Concinnella angulata POG., Mranciella trapezoidalis (KROT.). Ostracods: Paleodarwinula angusta 

(MAND.), Sinusuella pergraphica MAND. In <strong>the</strong> upper part – Dans la partie supérieure: Compsopteris adzvensis ZAL., Callipteris sp., Samaropsis 

postfrigida DOMBR.; Miospore complex H 2 (see Table 2). In lower part – Dans la partie inférieure: Lingula hyperborea IFAN., L.? miloradovichi IFAN., 

Astartella raricostata DEMB., Hemicycloleaia elata (ZASP.), Intia variabilis NEUB., Viatscheslavia vorcutensis ZAL., Samaropsis extensa NEUB.; Miospore 

complex H 1 (see Table 2) 

74. Foraminifera: Nodosaria netschajewi TSCHERD. Bivalves: Aviculopecten frederixi LUTK. and LOB., Astartella permocarbonica TSCHERN., 

Schizodus subobscurus LICH.; Brachiopods: Lingula? miloradovichi IFAN., Megousia kuliki (FRCKS.), Waagenowoncha irginae (STUCK.) 

Table 5 

Eastern European basin, Syktyokar belt 

Bassin Est-Européen, Ceinture de Syktyokar 

75. Poldarskaia formation: aleurolites, sandstones with intercalations <strong>of</strong> limestones and marls; in <strong>the</strong> lower part, sands with intercalations <strong>of</strong> gravels and 

conglomerates – Formation de Poldarskaia: aleurolites, grès à intercalations de calcaires et de marnes ; dans la partie inférieure, sables à intercalations 

de graviers et de conglomérats. Ostracods: Prasuchonella stelmachovi (SPIZH.), Suchonellina inornata SPIZH., S. fragilis (SCHN.). Conchostraca: 

Palaeortho<strong>the</strong>mos lochtaensis NOVOJ., Sphaerortho<strong>the</strong>mos cellelatus (LUTK.) 

76. Nuksenitskaia member: aleurolites, argillites, marls – Membre Nuksenitskaia: aleurolites, argilites, marnes. Ostracods: Suchonellina inornata SPIZH., 

S. parallela SPIZH., Prasuchonella stelmachovi (SPIZH.), Sinusuella ignota SPIZH. Vertebrates s.l.: Acrolepis macroderma EICHW. Dmitrievskaia member: 

dolomites, dolomitic marls, sometimes silicified – Membre Dmitrievskaia: dolomites, marnes dolomitiques, quelquefois silicifiées. Verkhnetozmenskaia 

member: marls, aleurolites and argillites, sandstones <strong>of</strong>ten gypseous with lenses <strong>of</strong> white sands – Membre Verkhnetozmenskaia: marnes, aleurolites et argilites, 

grès souvent gypseux à lentilles de sables blancs. 

77. Nijneonstinskaia formation; Formation de Nijneonstinskaia. Ostracods: Paleodarwinula sp., <strong>Permian</strong>a sp. Shardinskaia member: grey marls gypseous 

with intercalations <strong>of</strong> limestones and dolomites – Membre Shardinskaia: marnes grises gypseuses avec des intercalations de calcaires et de dolomies. 

Karpogorskaia member: brown red aleurolites, sandstones strongly gypseous with intercalations <strong>of</strong> argillites – Membre Karpogorskaia: aleurolites rouge-brun, 

grès fortement gypseux avec des intercalations d’argilites. Maregorskaia member: brown aleurolites and argillites with gypsum – Membre Maregorskaia: 

aleurolites brunes et argilites avec gypse.


545 

78. Foraminifera: Tetrataxis lata SPAND., Geinitzina postcarbonica SPAND., Glomospira pusilla GEIN., Netschajewia striata SCHLOTH. Bivalves: 

Pseudomonotis permianus MASL., Aviculopecten mutabilis LICH., Streblochondria sericea VERN., S. pusilla SCHLOTH. Gasteropods: Straparollus rossicus 

LICH. 

79. Foraminifera: Hemigordius discoides K.M. MACL., H. schlumbergeri (HOWCH.), Nodosaria hexagona TSCHERD., N. noinskii TSCHERD., N. 

farcimeniformis K.M. MACL., N. krotowi TSCHERD., N. cushmani PAALZ., Pseudonodosaria lata K.M. MACL., Lingulonodosaria kamensis K.M. MACL., 

Cornuspira megasphaerica GERKE, Nuculavus trivialis EICHW., Pleurophorus subocostatus BROWN., Polidevcia kazanensis (VERN.), Schizodus rossicus 

VERN. Ostracods: Pseudoparaparchites furmidabilis SCHN., Cavellina grandis SCHN., Healdianella vulgaris KOTSCH., Amphissites tscherdynzevi POSN., 

Kirkbya ingeniosa SCHN. Brachiopods: Neochonetes pinegenis KUL., Globiella tschernyschewi (NETSCH.), Licharewia rugulata (KUT.), Aulosteges 

horrescens (VERN.), Dielasma elongata SCHLOTH., Cancrinella cancrinoides (KUT.), Rhynchopora geinitziana (VERN.), Stepanoviella hemisphaerium 

(KUT.), Stepanovites meyeni KOZ. and MOV. 

80. Komaritskaia member: red aleurolites and sandstones with carbonated nodules and lenses <strong>of</strong> sandstone and sand at <strong>the</strong> bottom – Membre Komaritskaia: 

aleurolites et grès rouges avec des nodules de carbonates et des lentilles de grès et sables à la base. Foraminifera: Palaeomutela curiosa AMAL., P. 

keyserlingi AMAL. Bivalves: Opokiella tschernysohewi PLOTN. Ostracods: Suchonellina parallela SPIZH., S. trapezoida (SHAR.), S. fragiloides (ZEK.), S. 

fragilina (BEL.), Suchonella typica SPIZH., S. cyrta ZEK. Flora s.l.: Tatarina conspicua MEYEN, Peltaspermopsis sp., Permo<strong>the</strong>ca sp. Tsherevkovskaia 

member: red massive aleurolites, marls with intercalations <strong>of</strong> argillites, grey marls and limestones with lenses <strong>of</strong> sandstone and sand – Membre Tsherevkovskaia: 

aleurolites rouges massives, marnes avec intercalations d’argilites, marnes et calcaires gris avec des lentilles de grès et de sables. Ostracods: Suchonellina 

inornata SPIZH., S. parallela SPIZH., S. fragilina (BEL.), S. vladimirinae (BEL.), S. ignatjevi ZEK., Suchonella cyrta ZEK., S. typica SPIZH., S. auriculata 

(SHAR.), Darwinuloides svijazhicus (SHAR.) 

81. Vertebrates s.l.: Acrolepis macroderma EICHW., Dvinosaurus primus AMAL., Kotlassia prima (AMAL.), Karpinskiosaurus secundus (AMAL.), 

Jugosuchus licharevi RIAB., Inostrancevia alexandri AMAL., In. latifrons PRAVOSL., Ana<strong>the</strong>rapsidus petri (AMAL.), Dvinia prima AMAL., Scutosaurus 

karpinzkii AMAL., Pravoslavlevia parva (PRAVOSL.), Dicynodon amalitzkii SUSH., D. trautscholdi AMAL., D. annae (AMAL.) 

82. Poldarskaia formation: marls, dolomitic limestones, argillites, aleurolites, some conglomerates, sandstones, lenses <strong>of</strong> sandstone and sand with bones – 

Formation Poldarskaia: marnes, calcaires dolomitiques, argilites, aleurolites, quelques conglomérats, grès, lentilles de grès et de sables avec des fragments 

d’os. Bivalves – bivalves: Palaeomutela ovalis AMAL., P. oleniana GUS., Prilukiella janischewskia PLOT. Ostracods: Suchonellina inornata SPIZH., S. 

parallela SPIZH., Prasuchonella stelmachovi (SPIZH.). Vertebrates s.l.: Isadia suchonensis A. MIN., Platysomus biarmicus EICHW., Kotlassia prima 

(AMAL.), Dvinosaurus primus AMAL., Jugosuchus boreus IVACH., Niuksenitia sukhonensis TAT., Stinamara abrasa SAID. Flora s.l.: Tatarina sp. In <strong>the</strong> 

upper part – Dans la partie supérieure: Ostracods: Suchonellina vladimirinae (BEL.), S. fragilina (BEL.), S. fragilis SCHN., Suchonella cyrta ZEK., S. 

auriculata (SHAR.), <strong>Permian</strong>a tuberculata KASH. 

83. Nuksenitskaia member: alternation <strong>of</strong> brown, grey dolomites, marls and argillites with rare intercalations <strong>of</strong> white sands – Membre Nuksenitskaia: 

alternance de dolomies brunes, grises, de marnes et d’argilites avec de rares intercalations de sables blancs. Ostracods: Prasuchonella stelmachovi 

(SPIZH.), Suchonellina inornata SPIZH., S. futschiki KASH. 

84. Dmitrievskaia member: light grey and white dolomites with silex concretions – Membre Dmitrievskaia: dolomies gris clair et blanches avec des 

concrétions de silex. Ostracods: Prasuchonella stelmachovi (SPIZH.), Suchonellina inornata SPIZH., <strong>Permian</strong>a oblonga POSN. Verkhnetoz’menskaia 

member: dolomitic grey aleurolites and marls, grey and brown dolomites and argillites – Membre Verkhnetoz’menskaia: aleurolites et marnes grises 

dolomitiques, dolomies et argilites brunes et grises. Ostracods: Suchonellina inornata SPIZH., S. futschiki (KASH.). Vertebrates s.l.: Platysomus biarmicus 

EICHW., Amblypterus cf. tuberculatus EICHW., Paraamblypterus sp., Acrolepis cf. rhombifera EICHW. 

85. Aleurolites, argillites, marls, limestones and dolomites red and grey, with intercalations and lenses <strong>of</strong> sands and sandstone – Aleurolites, argilites, 

marnes, calcaires et dolomies rouges et grises, avec des intercalations et des lentilles de sables et de grès. Ostracods: Suchonellina inornata SPIZH., S. 

parallela SPIZH., S. futschiki (KASH.), S. undulata (MISH.), Prasuchonella stelmachovi (SPIZH.), <strong>Permian</strong>a elongata (POSN.), Sinusuella ignota SPIZH. 

Flora s.l.: Calamospora sp., Vittatina vittifer LUBER, Cordaitina sp., Podocarpites alatus LUBER, Piceites sublevis LUBER, Lebachiites sp., Striatohaplopinites 

elongatus LUBER, S. perfectum NAUM., Hamiapollenites tractiferinus (SAMOIL.) JANS, Vitreisporites sp. In upper part – Dans la partie supérieure: 

Ostracods: Suchonellina fragilis (SCHN.), S. ignatjevi ZEK., Suchonella cyrta ZEK., S. auriculata SHAR., S. typica SPIZH. 

86. Sandstones, sands, aleurolites, marls with thin levels <strong>of</strong> dolomitic limestones and argillites – Grès, sables, aleurolites, marnes avec des lentilles peu 

épaisses de calcaires dolomitiques et d’argilites. Ostracods: Paleodarwinula sp., Prasuchonella sp. Vertebrates s.l.: Palaeoniscidae gen. ind., Slivkovia sp.; 

Flora s.l.: Leiotriletes rotundum NAUM., Azonaletes levis LUBER, Ginkgocycadophytus tunguskensis (ANDR.) LUBER, Vittatina striata LUBER, Cordaitina 

sp., Podocarpites sp., Protodiploxypinus silvestritypus SAMOIL. 

87. Sand and sandstones with dolomitic cement with intercalations <strong>of</strong> argillites, marls, dolomites and gypsum – Sables et grès avec ciment dolomitique et 

avec des intercalations d’argilites, de marnes de dolomies et de gypse. Vertebrates s.l.: Ennatosaurus tecton VJUSCH. Mar’egorskaia member: argillites and 

gypseous aleurolites with intercalations <strong>of</strong> marls, dolomites and sandstones – Membre Mar’egorskaia: argilites et aleurolites gypseuses avec des intercalations 

de marnes, dolomies et grès. 

88. Upper part: anhydrites, in alternation with gypsum, dolomites, marls – Partie supérieure: anhydrites en alternance avec du gypse, des dolomies et des 

marnes. <strong>Middle</strong> part: dolomites, dolomitic marls, argillites with intercalations <strong>of</strong> gypsum – Partie moyenne: dolomies, marnes dolomitiques, argilites ave des 

intercalations de gypse. Lower part: dolomitic limestones, silicified, with gypsum – Partie inférieure: calcaires dolomitiques, silicifiés avec gypse. Flora s.l.: 

Trachytriletes sp., Raistrickia cardi<strong>of</strong>ormis (K.M.) WAR., Acanthotriletes gracilis (K.M.) WAR., Leiotriletes inderjensis (MAL.) WAR., Azonaletes tenuis 

LUBER, Vittatina sp., Cordaitina stenolimbata SAMOIL., C. rugulifera LUBER, Striatopodocarpites tojmensis SED., Striatodiplopinites lata (SCHATK.) 

WAR., Vitreisporites sp. 

89. argillaceous and biogenic limestones with intercalations <strong>of</strong> argillites and marls – Calcaires biogéniques et argileux avec des intercalations d’argilites 

et de marnes. Foraminifera: Nodosaria suchonensis K.M. MACL., N. hexagona TSCHERD., N. krotowi TSCHERD., Cornuspira microsphaerica K.M. 

MACL.; Ostracods: Healdia pseudosimplex KOTSCH., Moorea facilis SCHN., Amphissites tscherdynzevi POSN.; Brachiopods – brachiopodes: Lingula 

orientalis GOL., Stepanoviella tachernyschewi LICH., Licharewia rugulata (KUT.), L. keyserlingi (NESCH.), Aulosteges horrescens (VERN.), Spiriferella 

blasii (VERN.). Vertebrates s.l.: Anodontacanthus ru<strong>the</strong>norum CHAB., Ctenacanthus sp.; Flora s.l.– flore s.l.: Stepanovites meyeni KOZ. and MOV. 

90. Vagorskaia formation: argillaceous calcareous sandstones, in alternation with aleurolites and argillites, with intercalations <strong>of</strong> marls and biogenic 

limestones with inclusions <strong>of</strong> gypsum – Formation d’Ivagorskaia: grès calcaires argileux, en alternance avec des aleurolites et argilites, intercalation de 

marnes et de calcaires biogéniques à inclusions de gypse. Brachiopods: Lingula credneri GEIN., Licharewia stuckenbergi (NETSCH), Cleiothyridina 

semiconcava (WAAG.). Vertebrates s.l.: Janassa bituminosa (SCHLOTH.)


91. Flora s.l.: Trachytrilites punctatus (WALTZ), T. typicus NAUM., Acanthotriletes spinosellus WALTZ., A. gracilis (K.M.) WAR., Raistrickia 

cardi<strong>of</strong>ormis (K.M.) WAR., Azonaletes sp., Vittatina sp., Podocarpites major (LUBER) WAR., Protohaploxypinites tecturatus (LUBER) SAMOIL., 

Striatopodocarpites tojmensis SED., Striatodiplopinites elongatus (LUBER) ZAUER. 

92. Red marls and argillites with levels <strong>of</strong> sandstones and nodules <strong>of</strong> limestones and dolomites; marnes et argilites rouges avec niveaux de grès et nodules 

de calcaires et dolomies. Ostracods: Paleodarwinula elongata (LUN.), P. fragiliformis (KASH.), P. perlonga (SHARAP.), P. chramovi (GLEB.), 

Darwinuloides? tscherdynzevi KASH., <strong>Permian</strong>a elongata (POSN.) 

93. Elvinsky levels: marls and limestones with intercalations <strong>of</strong> argillites and dark grey limestones – Niveaux Elvinsky: marnes et calcaires avec 

intercalations d’argilites et de calcaires gris foncé. Glotovsky levels: limestones and marls light grey, sandy, gypsum, intercalations <strong>of</strong> limestones and 

aleurolites – Niveaux Glotovsky: calcaires et marnes gris clair, sableux, gypseux, intercalations de calcaires et d’aleurolites. Bivalves: Prilukiella? lata 

NETSCH. Ostracods: Paleodarwinula elongata (LUN.), P. fragiliformis (KASH.), P. chramovi (GLEB.), P. kassini (BEL.); Miospore complex J 1 (see Table 2) 

+ Striatohaplopinites amplus BALME and HEN., S. ellipticus ZAUER, Vittatina subsaccata SAMOIL., f. connectivalis ZAUER. 

94. Foraminifera: Nodosaria elabugae TSCHERD. Bivalves: Anthraconaia gapeevi (FED.), Palaeomutela attenuata GUS., Pseudomonotis permianus 

MASL., Promytilus retusus CHRON., Pteronites timanicus (MASL.), Pseudobakewellia cerathophagaeformis NOIN. Ostracods: Cavellina grandis SCHN., 

Healdia pseudosimplex KOTSCH. Brachiopods: Aulosteges fragilis (NETSCH.), Cancrinella cancrini VERN. Conchostraca: Ulugkemia borisi NOVOJ., 

Hemicycloleaia raymondi MOL. Flora s.l.: Viatscheslaviophyllum expansum BRONGN., Walchia sp.; Miospore complex I 2 (see Table 2) +Leiotriletes sp., 

Caytonipollenites sp., Acanthotriletes sp., Libumella rugulifera LUBER, Striatohaplopinites perfectus NAUM., Striatolebachiites varius ZAUER, Hamiapollenites 

tractiferinus SAMOIL., Alisporites tenuicorpus KL., Vesicaspora schemeli KL., Vittatina striata LUBER, Stepanovites meyeni KOZ. and MOV. 

95. Ostracods: Paleodarwinula borealis (KASH.), P. fainae (BEL.), Vyaella dobrynini KASH., <strong>Permian</strong>a posneri KASH. 

96. Sandy limestones and argillites with intercalations <strong>of</strong> marls and limestones – Calcaires sableux et argilites avec intercalations de marnes et de 

calcaires. Foraminifera: Nodosaria vjatkensis TSCHERD., N. pseudoconcinna K.M. MACL., Lingulonodosaria kamensis K.L. MACL. Bivalves: Nuculopsis 

wymensis (KEYS.), Cyrtorostra laticostata (NETSCH.), Pseudomonotis permianus MACL., Schizodus netschajewi LICH., Oriocrassatella plana (GOL.). 

Ostracods: Cavellina unica KOTSCH., Amphissites tscherdynzevi POSN., Healdia pseudosimplex KOTSCH. Brachiopods: Licharewia stuckenbergi 

(NETSCH.), Cancrinella cancrini VERN.; Flora s.l.: Callipteris adzvensis ZAL., Syniopteris nesterenkoi ZAL., Rufloria synensis (ZAL.); Miospore complex 

I 1 (see Table 2)+Granizonospora vulgaris (NAUM.), Laevigatosporites vulgaris IBR., f. gigantea LUBER, Florinites luberae SAMOIL., Lebachiacites 

pulcherrimus (ZAUER), Striatohaplopinites productus ABR., S. perfectus NAUM. 

97. Ostracods: Paleodarwinula ironae (BEL.), P. belousovae (KASH.), P. jatskovae (KASH.), Iniella oilmensis KASH. 

98. Ostracods: Paleodarwinula lubimovae (KASH.), P. biriensis (PAL.), Sinusuella ingloria KASH.; Miospore complex H 2 (see Table 2)+Remysporites 

psilopterus LUBER, Laevigatosporites vulgaris IBR. f. gigantea LUBER, Lebachiacites pulcherrimus ZAUER, Vesicaspora sp. 

99. Vitshegodskaia formation: reddish brown aleurolites, argillites, gypseous marls, with intercalations <strong>of</strong> limestones – Formation de Vitshegodskaia: 

aleurolites brun-rougâtre, argilites, marnes gypseuses; intercalations de calcaires. Bivalves: Notocrypta timanica MOL., Limnadea rossica MOL.; Miospore 

complex H 1 (see Table 2)+Cirratriradites procumbens LUBER, Granizonospora vulgaris NAUM., Florinites luberae SAMOIL., Lebachiacites pulcherrimus 

ZAUER, Striatohaplopinites perfectus NAUM., Vittatina striata LUBER. 

100. Ziminskaia formation: brown sandstones, aleurolites and marls in alternation with argillites and anhydrites – Formation de Ziminskaia: grès bruns, 

aleurolites et marnes en alternance avec des argilites et des anhydrites. Miospore complex H 1 (see Table 2) +Cordaitina uralensis LUBER, C. rugulifera 

LUBER, Klausipollenites sp., Scheuringipollenites ovatus BALME and HEN., Striatohaplopinites perfectus NAUM., Striatolebachiites varius ZAUER, 

Hamiapollenites tractiforinus SAMOIL., Vittatina subsaccata SAMOIL., Azonaletes irregulariplicatus SAMOIL. 

101. Oust’koulomsky levels: red argillites with intercalations <strong>of</strong> limestones and sandstones – Niveaux Oust’koulomsky: argilites rouges avec des 

intercalations de calcaires et de grès. Ostracods: Paleodarwinula abunda (MAND.), P. angusta (MAND.), Prasuchonella scolia (MAND.), Vymeolla 

timanica (KASH). 

102. Limestones and argillites; calcaires et argilites. Ostracods: Paleodarwinula angusta (MAND.), Prasuchonella scolia (MAND.), Sinusuella 

pergraphica MAND. 

103. Kirdorsky levels: argillites with intercalations <strong>of</strong> limestones and sandstones – Niveaux Kirdorsky: argilites avec intercalations de calcaires et de grès. 

Ostracods: Paleodarwinula angusta (MAND.), P. abunda (MAND.). 

104. Sindorsky levels: limestones, dolomites, argillites and sandstones – Niveaux Sindorsky: calcaires, dolomies, argilites et grès. Bivalves: Anthraconala 

castor (EICHW.), Nuculopsis celebrosus (DEMB.). Ostracods: Bairdia plebeia REUS, Paleodarwinula parphenovae (BEL.), P. angusta (MAND.), P. abunda 

(MAND.), Volganella spizharskii MAND.; Vertebrates s.l.: Palaeolimnadiopsis eichwaldi (NETSCH.); Miospore complex H 1 (see Table 2) +Granulatisporites 

larvatus LUBER. 

105. Vertebrates s.l.: Clatysomus biarmicus EICHW., Acrolepis cf. rhombifera EICHW., A. cf. murchisonia EICHW., A<strong>the</strong>rstonia sp., Amblypterus sp. Flora 

s.l.: Callipteris adzvensis ZAL., Rhipidopsis ginkgoides SCHMALH., Phylladoderma arberi ZAL., Glostophyllus sp.; Miospore complex J (see Table 2). 

106. Ostracods: Paleodarwinula abunda (MAND.), Sinusuella pergraphica MAND. Bivalves: Anthraconauta tschernyschewi KHALF., A. gibbosa 

BETEKHT., Abiella subovata (JONES), Syniella bella (GUS.). 

107. Bivalves: Concinnella angulata POG., Anthraconaia(?) rhomboides (NETSCH.), Mrassiella(?) trapezoidalis (KROT.). Ostracods: Volganella objecta 

KASH., Iniella intaensis KASH., Placidea cf. petschorica SCHN., Sinusuella pergraphica MAND. Flora s.l.: Intia vermicularis NEUB., Viatscheslavia 

vorkoutensis ZAL., Knorriopsis uralica ZAL.; Miospore complex H (see Table 2) 

108. Alternation <strong>of</strong> grey argillites, aleurolites and marls – Alternance d’argilites grises, aleurolites et marnes. Bivalves: Concinnella concinnaeformis 

POG., Lithophaga consobrina EICHW., Netschajewia globosa (NETSCH.), Pseudobakewellia ceratophagaeformis NOIN. Flora s.l.: Bardocarpus aliger 

ZAL., Samaropsis triquetra ZAL.; Miospore complex H 1 (see Table 2).


547 

Table 6 

Eastern European Basin, Perm belt 

Bassin Est-Européen, Ceinture de Perm 

109. Molomskaia member – Membre Molomskaia. Ostracods: Darwinuloides tataricus POSN., D. swijazhicus (SHARAP.). Loutugskaia member – Membre 

Loutugskaia. Bivalves: Palaeanodonta okensis AMAL., P. segmentata GUS., P. solemvaetormis (NETSCH.), Maturipupa tolstichinae (REV.). Ostracods: 

Suchonella typica SPIZH., S. cornuta KASH. Conchostraca: Pseudes<strong>the</strong>ria olferievi NOVOJ. Flora s.l.: Cyclotungusites vokhmaensis NOVOJ., Tatarina 

olferievi MEYEN, Nahocynodon seductus TAT.; Miospore complex J 2 (see Table 2). Zamochnikovskaia member. Bivalves: Palaeanodonta segmentata GUS., 

P. solemyaeformis (NETSCH.), Surella blomi GUS., Vetlugaia schmidti (REV.). Ostracods: Suchonella typica SPIZH., S. cornuta KASH. Conchostraca: 

Leptes<strong>the</strong>ria convexa NOVOJ. 

110. Bivalves: Palaeomutela orthodonta AMAL., P. keyserlingi (KEYS.), P. ovalis KEYS., P. inostranzevi AMAL., P. curiosa AMAL., Palaeanodonta 

okensis AMAL., P. fischeri AMAL. Ostracods: Suchonellina parallela SPIZH., S. inornata var. macra LUN., S. spizharskyi POSNER, S. futschiki KASH. 

Conchostraca: Limnadia jeskinoica NOV. Vertebrates s.l.: Dvinosaurus egregius SHISHK., Bystrowiana permira VJUSCH., Jugosuchus mirabilis (VJUSCH.). 

111. Ostracods: Suchonellina parallela SPIZH., S. inornata SPIZH., S. daedala MISH., Prasuchonella stelmachovi (SPIZH.). Conchostraca: Limnadia 

kastiana NOV.; Miospore complex J 2 (see Table 2). 

112. Ostracods: Paleodarwinula perlonga (SHARAP.), P. chramovi (GLEB.), P. kassini (BEL.), P. elongata (LUN.), P. impostor (MISH.), Darwinuloides 

sentjakensis SHARAP., Prasuchonella nasalis SHARAP. 

113. Member 2: Miospore complex I 2 (see Table 2). Member 1: Miospore complex I 1 (see Table 2). 

114. Member 2: alternation <strong>of</strong> dolomites and dolomitised limestones, grey with nodules <strong>of</strong> gypsum, with organic remains – Alternance de dolomies et de 

calcaires dolomitiques gris à nodules de gypse et restes organiques. Member 1: light grey biogenic limestones sometimes with gypsum, sands, shales, with 

concretions <strong>of</strong> flint, organic remains; in <strong>the</strong> lower part, gravels and pieces <strong>of</strong> dolomitic limestones – Calcaires biogéniques gris clair, quelquefois avec du gypse, 

des sables, des shales, à concrétions siliceuses et débris organiques; dans la partie inférieure, graviers et fragments de calcaires dolomitiques. Bivalves: 

Pseudomonotis permianus MASL., Netschajewia globosa (NETSCH.), Streblochonaria sericea (VERN.), Aviculopecten cf. russiensis NETZCH. Brachiopods: 

Cancrinella cancrini (VERN.). 

115. Bivalves: Streblochonaria sericea (VERN.). Brachiopods: Stenoscisma superstes (VERN.), Cleiothyridinna pectinifera (SOW.), Pinegathyris 

royssiana (KEYS.), Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.), Licharewia rugulata (KUT.), Blasispirifer blasii (VERN.). Ostracods: 

Amphissites tscherdynzevi POSNER, Healdia oblonga KOTSCH., H. simplex ROUNDY, H. reniformis GUSS., H. postcornuta SCHN., Healdianella vulgaris 

KOTSCH. 

116. Nefedovskaia member: yellow-brown sandstones, oblique stratifications, with conglomerates with pebbles coming from local and metamorphic rocks 

(up to 20 m), in <strong>the</strong> upper part brownish red shales, limestones, with limestone concretions – Grès brun-jaunâtre, stratifications obliques, avec des 

conglomérats à galets provenant des roches locales et métamorphiques (jusqu’à 20 m), dans la partie supérieure des shales rouge-brunâtre, des calcaires 

et des concrétions calcaires. Ostracods: Suchonella inornata SPIZH.; Vertebrates s.l.: Nanocynodon seductus TAT. In western part: brown and reddish brown


shales, friable, greenish grey – Dans la partie occidentale: shales brun-rougeâtre et bruns, friables, gris-verdâtre. Ostracods: Darwinuloides tataricus 

(POSNER), D. svijazhicus (SHARAP.). Bikovskaia member: brownish grey sandstones, large, oblique stratifications, with conglomerates and gravels <strong>of</strong> local 

rocks and metamorphic rocks from Ural; higher, shales and aleurolites reddish brown, limestones with intercalations <strong>of</strong> variegated and dark grey limestones – 

Membre Bikovskaia: grès gris-brunâtre, stratifications larges et obliques, avec conglomérats et graviers d’origine locale et métamorphique provenant de 

l’Oural; plus haut, shales et aleurolites brun-rougeâtre, calcaires avec intercalations de calcaires gris foncé et bariolé. Ostracods: Suchonella typica SPIZH. 

Westwards, reddish brown shales, with intercalations <strong>of</strong> friable aleurolites and coarse sandstones, at <strong>the</strong> bottom gravels – Vers l’Ouest, shales brun-rougeâtre, 

avec des intercalations d’aleurolites friables et de grès grossiers, graviers à la base. Ostracods: Suchonella parvaeformis (KASH.), S. typica SPIZH., 

Darwinuloides tataricus (POSNER), D. svijazhicus (SHARAP.). Kalininskaia member: shales and brown aleurolites, limestones, porous with roots – Membre 

Kalininskaia: shale et aleurolites brunes, calcaires poreux avec des racines. 

117. Poutaninskaia member: shales and aleurolites brownish grey, limestones, variegated marls, dark grey shaly limestones with gastropods – Membre 

Poutaninskaia: shales et aleurolites gris-brun, calcaires, marnes bariolées, calcaires détritiques gris foncéàgastéropodes. Ostracods: Suchonellina inornata 

SPIZH., Darwinuloides avijazhicus (SHAR.), <strong>Permian</strong>a oblonga (POSNER). Bivalves: Palaeomutela verneuili AMAL., P. subparallela AMAL. 

118. Urpalovkaia member: at <strong>the</strong> bottom coarse sandstones with oblique stratifications; higher, red aleurolites and shales with concretions <strong>of</strong> limestones – 

Membre Urpalovkaia: à la base, grès grossiers avec stratifications entrecroissées; plus haut, aleurolites et shales rouges à concrétions de calcaires. Rare 

Paleaomutela trapezoidalis AMAL. (Bivalves – bivalves) and ostracods. 

119. Vertebrates s.l.: Scutosaurus rossicus (HART.–WEIN.), Deltovjatia vjatkensis (HART.–WEIN.). 

120. Filinskaia member: grey limestones, marls pink, grey, green, calcareous grey marls – Membre Filinskaia: calcaires gris, marnes roses, grises, vertes, 

marnes calcaires grises. Ostracods: Suchonellina parallela SPIZH. Slobodskaia member: in <strong>the</strong> lower part, sandstones and conglomerates with oblique 

stratifications; in <strong>the</strong> upper part shales and red aleurolites – Membre Slobodskaia: partie inférieure, conglomérats et grès à stratifications obliques; partie 

supérieure, shales et aleurolites rouges. Bivalves: Palaeomutela ulemaensis GUS. Ostracods: Suchonellina paralella SPIZH. 

121. Siramskaia member – Membre Siramskaia: Bivalves: Prilukiella nitida GUS., Palaeomutela vjatkensis GUS., P. ovalis AMAL. Ostracods: 

Paleodarwinula fragiliformis (KASH.), <strong>Permian</strong>a oblinga (POSNER). 

122. Belokholomnizkaia member: white fine grain sandstones, thin bedded, with quartz and limestones, shales, grey yellowish aleurolites, variegated marls 

with intercalations <strong>of</strong> limestones – Membre Belokholomnizkaia: grès blancs à grains fins, finement lités, avec quartz et calcaires, argillites, aleurolites 

gris-jaunâtre, marnes bariolées avec intercalations de calcaires. Ostracods: <strong>Permian</strong>a tuberculata KASH. 

123. Arthropods: Platyops stuckenbergi TRAUT., P. watsoni EFR. Vertebrates s.l.: Melosourus uralensis MEYER, Nycteroleter kassini TCHUD., 

Nyctiboetus liteus TCHUD., Leptoropha talanophora (TCHUD.), Gecatogomphius kavejevi VJUSCH., Molybdopygus arcanus TCHUD. 

124. Ieinskaia member: brownish red calcareous shales and aleurolites, with grains <strong>of</strong> quartz and limestones, intercalations <strong>of</strong> poorly sandy marls and 

limestones – Membre Ieinskaia: aleurolites et shales calcaires rouge-brunâtre, avec des grains de quartz et de calcaires, intercalations de marnes sableuses 

et de calcaires. Rare <strong>Permian</strong>a tuberculata KASH. (Ostracods). Makisinovskaia member: alternation <strong>of</strong> feldspatho-quartzic sandstones, thin grains, with 

gypsum, aleurolites, shales, variegated marls and limestones – Membre Makisinovskaia: alternance de grès feldspatho– quartzeux, à grain fin avec du gypse, 

shales, aleurolites, marnes et calcaires bariolés. Ostracods: Darwinuloides edmistonus (BEL.) 

125. Member 3: in <strong>the</strong> lower part, greenish grey sandstones mainly quartzitic; higher, alternation <strong>of</strong> sandstones with gypsum, aleurolites, shales, marls, 

dolomites, limestones with pelecypods – Membre3:partie inférieure, grès gris-verdâtre principalement quartizitiques ; puis, alternance de grès avec gypse, 

aleurolites, shales, marnes, dolomites, calcaires avec pélécypodes. Arthropods: Platyops stuceubergi TRAUT. Member 2: in <strong>the</strong> lower part, green-grey 

sandstones with quartz, locally gypsum, in alternation with shales and grey aleurolites, higher, limestones, dolomites, grey marls with foraminifers, brachiopods 

and bivalves – Membre2:partie inférieure, grès gris-vert à quartz, localement du gypse, en alternance avec des shales et des aleurolites grises ; plus haut, 

calcaires, dolomites, marnes grises à foraminifères, brachiopodes et bivalves. Member 1: in <strong>the</strong> lower part, green-grey and brown sandstones, with gypsum 

nodules; higher, grey shales, dolomites, thinly bedded, limestones sometimes oolithic, bryozoans, marine ostracods – Membre1:partie inférieure, grès bruns 

et gris-vert, avec nodules de gypse ; plus haut, shales gris, dolomies, finement litées, calcaires quelquefois oolithiques, bryozoaires, ostracodes marins. 

Brachiopods: Cancrinella cancrini (VERN.) 

126. Calcareous member: biogenic limestones, reefal with brachiopods and crinoids, algae and bryozoans, ooli<strong>the</strong>s, with intercalations <strong>of</strong> marls, shales, 

sandstones. Membre calcaire : calcaire biogénique, récifal avec brachiopodes et crinoïdes, algues et bryozoaires, avec intercalations de marnes, shales, grès. 

Foraminifera: Nodosaria elabugae TSCHERD., N. pugioidea ZOL. and IGON., N. netschajewi TSCHERD., N. permiana SPAND., N. krotowi TSCHERD., 

Frondicularia sp. Brachiopods: Cleiothyridina pectinifera (SOW.), Cancrinella sp. Marly calcareous member: biogenic limestones with stratifications, light 

marls, dark grey shales. Membre marno-calcaire : calcaires biogéniques stratifiés, marnes claires, shales gris foncé. Foraminifera: Nodosaria elabugae 

TSCHERD., N. netachajewi TSCHERD., N. krotowi TSCHERD. Bivalves: Dentalina sp. Brachiopods: Cancrinella cancrini (VERN.), Globiella 

hemisphaerium (KUT.). Sandy calcareous member: greenish grey sandstones, glauconitic limestones, dark grey shales, limestones; eastwards, terrigenous rocks 

with few intercalations <strong>of</strong> limestones. Membre calcaro-sableux: grès gris-verdâtre, calcaires glauconitiques, shales gris foncé, calcaires; vers l’Est, roches 

terrigènes avec quelques intercalations de calcaires. Foraminifera: Cornuspira kamae TSCHERD., Nodosaria netachajewi TSCHERD., Prondicularia 

longissima sintasica IGON. Brachiopods: Cleiothyridina pectinitera (SOW.), Rhynchopora sp. 

127. Foraminifera: Palaeomutela oleniana GUS., P. plana AMAL. Ostracods: Suchonellina parallela SPIZH., S. futschiki (KASH.). Vertebrates s.l.: 

Pareiasauria sp., Gorgonopsia sp., Inostranzevia sp. 

128. Ostracods: Suchonellina parallela SPIZH. 

129. Upper part: alternation <strong>of</strong> conglomerate with pebbles <strong>of</strong> flint and quartz, sandstones, brown shales, grey marls, algae limestones. Partie supérieure: 

alternance de conglomérats à galets de silice et de quartz, grès, shales bruns, marnes grises, calcaires algaires. Bivalves: Prilukiella nitida GUS., 

Anthraconauta volgensis GUS., A. uslonensis GUS. Ostracods: Paleodarwinula fragiliformis (KASH.), P. elongata (LUN.). Vertebrates s.l.: Biarmica 

tchudinovi IVACH., Brithopus priscus KUT. Lower part: alternation <strong>of</strong> conglomerate, gravels with flint and quartz, with fossil lignite, grey sandstones with 

oblique stratifications, shales, calcareous brown red aleurolites, variegated marls, dark grey limestones. Partie inférieure : alternance de conglomérats, 

graviers avec silice et quartz, lignite fossile, grès gris à stratifications obliques, shales, aleurolites calcaires rouge-brunâtre, marnes bariolées, calcaires gris 

sombre. Bivalves: Prilukiella lata (NETSCH.), Palaeomutela doratiformis GUS., P. celebrata GUS., Prilukiella fastigata GUS., P. pugnatoria GUS. 

Ostracods: Paleodarwinula kassini (BEL.)


549 

130. Ostracods: Prasuchonella serpula BEL., Paleodarwinula varsan<strong>of</strong>ievae (BEL.), P. alexandrinae (BEL.), Darwinuloides edmistonus (BEL.), D. 

sentjakensis (SHARAP.). Vertebrates s.l.: Parabradysaurus udmurticus EFR., Archaesyodon praeventor TCHUD., Eotitanosuchus olsoni TCHUD., Ivantosaurus 

ensifer TCHUD., Chthomaloporus lenocinator TCHUD., Estemmenosuchus mirabilis TCHUD., Est. uralensis TCHUD., Anoplosuchus tenuirostris 

TCHUD., Zopherosuchus luceus TCHUD., Otsheria netzvetaevi TCHUD., Brithopus priscus KUT. 

131. Ostracods: Paleodarwinula parphonovae (BEL.), P. abunda (MAND.), Vymella timanica (KASH.), V. petschorica (KASH.). Flora s.l.: Comia 

biarmica (ZAL.) VLAD., Sphenopteris bifida SCHMALH., Syniopteris cuneifolia KUTORGA, Callipteris erosa (MORRIS) ZAL., Paracalamites striatus 

SCHMALH., Odontopteris rossica (ZAL.) VLAD., Syniopteris cuneifolia KUTORGA, Permo<strong>the</strong>ca sardykensis ZAL. 

132. To <strong>the</strong> East, limestones, dolomites, grey shales, marls, in <strong>the</strong> lower part levels with bivalves and brachiopods. Vers l’Est : calcaires, dolomies, shales 

gris, marnes; dans la partie inférieure, niveaux à bivalves et brachiopodes. Ostracods: Paleodarwinula monitoria (MAND.), Kemeroviana angulata MAND.; 

in <strong>the</strong> lower part Brachiopods: Cancrinella cancrini (VERN.), C. koninekiana (KEY.). Flora s.l.: Intia variabilis NEUB., Viatscheslavia vorcutensis ZAL. To 

<strong>the</strong> West, variegated shales with gypsum, marls, limestones, dolomites, with rare ostracods. Vers l’Ouest, shales bariolés à gypse, marnes, calcaires, dolomies, 

rares ostracodes. Flora s.l.: Odontopteris artipinnata ZAL., Entsovia rarisulcata MEYEN, Samaropsis frigida NEUB., S. vorcutana TSCHIRK. 

Table 7 

Eastern European basin, Kazan belt 

Bassin Est-Européen, Ceinture de Kazan 

133. Vertebrates s.l.: Chroniosuchus sp. Bivalves: Palaeomutela oleniana GUS., P. plana AMAL., Palaeanodonta okensis AMAL., P. abunda NETSCH. 

Conchostraca: Pseudoes<strong>the</strong>ria suchonensis (NOVOJ.), Ps. nordwikensis (NOVOJ.) 

134. Conchostraca: Pseudoes<strong>the</strong>ria suchonensis (NOV.), Ps. nordwikensis (NOV.), Ps. lineata (LUTK.), Asmussia petasa (NOVOJ.), Concherisma 

trapezoidalis (NETSCH.), Brachyes<strong>the</strong>ria ovalis MOL., Leptes<strong>the</strong>ria uralensis MOL. Bivalves: Palaeomutela keyserlingi AMAL. Ostracods: Suchonellina 

parallela SPIZH., S. futechiki KASH. 

135. Brown red argillites and aleurolites, dolomitised with intercalations <strong>of</strong> pink and grey marls, argillites with palygorskite. In <strong>the</strong> lower part, level <strong>of</strong> 

calcareous argillites. Argilites et aleurolites rouge-brun, dolimitisées avec intercalations de marnes roses et grises à palygorskite ; dans la partie inférieure, 

niveau d’argilites calcaires. 

136. Dolomites and dolomitised limestones, grey, silicified with bivalve, brachiopod, gastropod debris. Dolomies et calcaires dolomitisés, gris, silicifiés 

avec débris de bivalves, brachiopodes, gastéropodes. 

137. Yellow grey clastic dolomitised limestones with Licharewia rugulata (KUT.) (Brachiopods). Calcaires dolomitisés clastiques gris–jaune avec 

Licharewia rugulata (KUT.). 

138. Light grey dolomitised limestones, oolitic with Pseudomonototis garfor<strong>the</strong>nsis (KING.) (Bivalves). Calcaires dolomitisés gris clair, oolithiques avec 

Pseudomonototis garfor<strong>the</strong>nsis (KING.).


139. Dolomitised biogenic limestones with Lingula orientalis GOL. (Brachiopods). Calcaires biogéniques dolomitisés avec Lingula orientalis GOL. 

140. Vertebrates s.l.: Platysomus biarmicus EICHW., Jugosuchus hartmanni IVACH., Sauroctonus progressus (HAR.–WEIN.), Scutosaurus permianus 

(HART.–WEIN.), Sc. itilenis IVACH. and LAB., Dicynodon trautsholdi ANAL. 

141. In <strong>the</strong> lower part, brown-grey sandstones and conglomerates with pebbles <strong>of</strong> metamorphic rocks; in <strong>the</strong> upper part red-brown argillites and aleurolites, 

variegated marls, algal stratified limestones. Dans la partie inférieure, grès gris-brun et conglomérats à galets de roches métamorphiques ; dans la partie 

supérieure, argilites et aleurolites brun-rouge, marnes bariolées, calcaires algaires stratifiés. Bivalves: Palaeomutela doratifomis GUS., Antraconauta 

volgensis GUS., Palaeanodonta (Kuthreconia) castor (EICHW.). Ostracods: Paleodarwinula <strong>the</strong>odorovichi (BEL.), Prasuchonella nasalis SHARAP. 

Vertebrates s.l.: Xenosynechodus egloni GLÙK., Melosaurus psucidens (KONZH.), Lanthanosuchus watsoni EFR., Enosuchus breviceps KONZH., 

Titanophoneus potens EFR., Syodon biarmicum KUT., Doliosauriscus janshinovi ORL. 

142. In <strong>the</strong> lower part, brown sandstones, polymictites with conglomerates (pebbles <strong>of</strong> chert and marly limestones); in <strong>the</strong> upper part, brown argillite and 

aleurolites, variegated marls, stratified platy and algal limestones. Dans la partie inférieure, grès bruns, polymictites avec conglomérats (galets de cherts et 

de calcaires marneux) ; dans la partie supérieure, argilites et aleurolites brunes, marnes bariolées, calcaires algaires et en plaquettes bien stratifiés. 

Bivalves: Anthraconauta uslonensis GUS. 

143. In <strong>the</strong> lower part, greenish grey sandstones, in places, very coarse pebbles mainly quartzose; in <strong>the</strong> upper part variegated marls and argillites. Dans la 

partie inférieure, grès gris-verdâtre, localement, galets très grossiers principalement quartzeux ; dans la partie supérieure, marnes et argilites, bariolées. 

Bivalves: Nuculavus trivialis (EICHW.), Schizodus rossicus VERN., Pseudomonotis permianus MASL., Streblochondria sericea (VERN.), Pseudobakewellia 

ceratophagaeformis NOIN., Lithophaga consobrina (EICHW.); Brachiopods: Cancrinella cancrini (VERN.). Arthropods : Platyops stuckenbergi TRAUT. 

Vertebrates s.l.: Nyateroleter barshkyricus EFR., Rhipaeosaurus tricuspidens EFR., Phthinosaurus borissiaki EFR., Belebey vegrandis IVACH. Flora s.l.: 

Signacularia noinskyi ZAL., Sphenophyllum stueckenbergii SCHMALH., Phyllododerma volgensis ESAUL., Permo<strong>the</strong>ca sardykensis ZAL. 

144. Light grey cavernous oolithic limestones and sandstones, thin stratifications with in some places gypsum; at <strong>the</strong> base, brown greyish sandstones, grey 

yellowish argillites. Grès et calcaires oolithiques caverneux gris clair, fines stratifications avec quelques niveaux de gypse ; à la base, grès brun-grisâtre, 

argilites gris-jaunâtre. Bivalves: Alula kutorgai (VERN.), Pseudobakewellia ceratophagaeformis NOIN., Pseudomonotis pemianus MASL. Bryozoans: 

Permonantis cornutus (GOL.), Paralioclemeta multispinosum MOROZ., Fenestella schurae MOROZ., Polypora kasanensis MOROZ., Dyscritella incrustata 

MOROZ. Brachiopods: Cancrinella cancrini (VERN.), Odontospirifer subcostatus (NETSCH.). 

145. Greyish brown limestones, compact and oolithic, dolomies with gypsum. Calcaires brun-grisâtre, compacts et oolithiques, dolomies avec gypse. 

Foraminifera: Nodosaria krotowi TSCHERD. Cnidaria: Conularia hollebeni GEIN. Bryozoans: Fistulipora dybowskii GOR., Tabulipora ordinata MOROZ., 

Discritella incrustata MOROZ. Brachiopods: Aulosteges wangenheimi (VERN.), Stenoscisma superates (VERN.), Orbiculoidea konincki (GEIN.) 

146. In <strong>the</strong> lower part, grey argillites and sandstones with gypseous cement; in <strong>the</strong> upper part, stratified and oolithic limestones and dolomites locally with 

gypsum. Dans la partie inférieure, argilites grises et grès à ciment gypseux ; dans la partie supérieure, dolomies et calcaires oolithiques et stratifiés, gypse 

de place en place. Foraminifera: Nodosaria elabugae TSCHERD., N. hexagona TSCHERD., N. suchonensis K.M. MACL.; Brachiopods: Aulosteges fragilis 

(NETSCH.), A. horrescens (VERN.). Bryozoans: Pseudobatostomella decora MOROZ., Fenestella permulta MOROZ. Bivalves: Pseudomonotis permianus 

MASL., Pseudobakewellia certophagaefomis NOIN. 

147. Bioclastic, reefal limestones with brachiopods and crinoids, with intercalations <strong>of</strong> argillites, marls and sandstones. Calcaires récifaux, bioclastiques à 

brachiopodes et crinoïdes, avec intercalations d’argilites, marnes et grès. Foraminifera: Nodosaria elabugae TSCHERD., N. pugioides ZOL. and IGON., N. 

netschajewi TSCHERD., N. permiana SPAND., N. krotowi TSCHERD., Fronticularia sp. Brachiopods: Cleiothyridina pectinifera (SOW.), Cancrinella sp. 

148. Bioclastic limestones with medium and thin foliaceous stratifications, calcareous marls, dark grey argillites. Calcaires bioclastiques avec stratifications 

en feuillets fins ou moyens, marnes calcaires, argilites gris foncé. Foraminifera: Nodosaria elabugae TSCHERD., N. netschajewi TSCHERD., N. krotowi 

TSCHERD., Dentalina sp. Brachiopods: Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.). 

149. Greenish grey sandstones, limestones with glauconite, mainly quartzose, dark grey argillites sometimes calcareous, dolomites. Grès gris-verdâtre, 

calcaires à glauconite, principalement quartzeux, argilites gris foncé, quelquefois calcaires, dolomies. Foraminifera: Cornuspira kamae TSCHERD., 

Nodosaria netschajewi TSCHERD., Frondicularia longissima sintasica IGON. Brachiopods: Cleiothyridina pectinifera (SOW.), Rhynchopora sp. 

150. Westwards, sandstones, argillites, grey aleurolites and limestones with brachiopods; eastwards, variegated argillites, aleurolites. Vers l’Ouest, grès, 

argilites, calcaires et aleurolites grises à brachiopodes ; vers l’Est, argilites bariolées, aleurolites. Gasteropods: Goniasma subangulata (GOL.), Polidevcia 

kazanensis (VERN.). Bivalves: Pseudobakewellia ceratophagaeformis NOIN., Netschajewia globosa (NSTACH.), Lithophaga consobrina (EICHW.). 

Brachiopods: Cancrinella cancrini (VERN.), Licharewia rugulata (KUT.), Cleiothyridina pectinifera (SOW.). Ostracods: Amphissites tscherdynzevi 

POSNER, Healdia subtriangula KOTSCH., Cavellina unica KOTSCH. 

151. Sandstones, aleurolites, grey argillites with intercalations <strong>of</strong> marls and limestones. Grès, aleurolites, argilites vertes avec intercalations de marnes et 

calcaires. Foraminifera: Nodosaria hexagona (TSCHERD.), N. urmarensis K.M. MACL. Brachiopods: Lingulina semivelata TSCHERD, Lingula orientalis 

GOL., Globiella hemisphaerium (KUT.), Licharewia stuckenbergi (NETSCH.), Pinegathyris royssiana (KEYS.), Beecheria netschajewi GRIG. Bivalves: 

Pseudobakewellia ceratophagaeformis NOIN. Bryozoans: Wjatkella wjatkensis (NESCH.). Ostracods: Amphissites tscherdynzevi POSN., Healdia subtriangula 

KOTSCH. Flora s.l.: Signacularia noinskii ZAL., Odontopteris rossica ZAL., Phylladoderma meridionalis NEUB., Nucicarpus tarlovskiensis ESAUL. 

152. Argillites with lignite, limestones. Argilites avec lignite, calcaires. Foraminifera: Nodosaria hexagona (TSCHERD.), N. urmarensis K.M. MACL. 

Tabulates: Tabulata ordinata MOROZ. Bryozoans: Fenestella permulta MOROZ. Brachiopods: Cancrinella cancrini (VERN.), Globiella hemisphaerium 

(KUT.), Licharewia rugulata (KUT.), Pinegathyris royssiana (KEYS.), Cleiothyridina pectinifera (SOW.), Baitugania netschajewi GRUNT, Beecheria 

netschajewi GRIG. Ostracods: Amphissites tscherdynzevi POSN., Cavellina unica KOTSCH. Flora s.l.: Phylladoderma fefiloviana BOGOV, Sylvella alata 

ZAL. 

153. Grey sandstones with charred vegetal, with intercalations <strong>of</strong> argillites. Grès gris avec végétaux carbonisés, intercalations d’argilites. Brachiopods: 

Lingula orientalis GOL. 

154. Ostracods: Paleodarwinula ufimica (PAL.), P. biriensis (PAL.), P. burajavoensis (PAL.). Flora s.l.: Paracalamites striatus ZAL., Odontopteris rossica 

ZAL., Phylladoderma septata BOGOV. 

155. Ostracods: Suchonella inornata (SPIZH.), Suchonella typica (SPIZH.), Tscherdynzeviana buslukensis KASH. 

156. Ostracods: Suchonella stelmachovi SPIZH., S. nasalis SHARAP., Paleodarwinula fragiliformis (KASH.), P. polenovae (BEL.). 

157. Ostracods: Paleodarwinula fragiliformis (KASH.), Darwinuloides tscherdynzevi KASH., D. dobrjankaensis KASH. 

158. Variegated argillites, aleurolites, sandstones, grey marls, limestones, dolomites with undetermined fresh water ostracods. Argilites bariolées, 

aleurolites, marnes grises, calcaires, dolomites avec des ostracodes d’eau douce indéterminés. 

159. Red conglomerates, sandstones, light grey limestones, marls, dolomites. Conglomérats rouges, grès, calcaires gris clair, marnes et dolomies. 

Ostracods: Prasuchonella nasalis (SHARAP.), P. belebeica BEL., D. fainae BEL., Darwinuloides edmistonus BEL., D. sentjakensis (SHARAP.)


551 

160. Light grey limestones foliated and gritty, dolomites, red conglomerates, sandstones, aleurolites, argillites. Calcaires gris clair graveleux et en 

plaquettes, dolomies, conglomérats rouges, grès, aleurolites et argilites. Ostracods: Schneideria kazanica KOTSCH., Paleodarwinula alexandrinae (BEL.), 

P. fainae (BEL.), P. tuimazensis (KOTSCH.). Flora s.l.: Signacularia noinskii ZAL., Odontopteris rossica ZAL. 

161. Greenish grey sandstones, conglomerates, red aleurolites, dolomites cavernous and porous. Grès gris-verdâtre, conglomérats, aleurolites rouges, 

dolomies caverneuses et poreuses. Ostracods: Darwinuloides sentjakensis (SHARAP.), D. edmistonus BEL. 

162. Foliated limestones, light and dark grey dolomites with intercalations <strong>of</strong> grey argillites, marls, lignite, gypsum. Calcaires feuilletés, dolomies gris clair 

à foncé avec intercalations d’argilites, marnes, lignite et gypse. Ostracods: Cavellina unica KOTSCH., Healdia simplex ROUNDY, Schneideria kazanica 

KOTSCH.; Bivalves: Liebea hausmanni (GOLDFUSS), Pseudomonotis garfor<strong>the</strong>nis KING. 

163. Light grey and greenish sandstones with lenses <strong>of</strong> conglomerates. Grès gris clair et verdâtres avec lentilles de conglomérats. Foraminifera: 

Cornuspira megasphaerica GERKE. Ostracods: Cavellina grandis SCHN. 

164. Argillites, marls, bioclastic limestones, greenish grey sandstones. Argilites, marnes, calcaires bioclastiques, grès gris-verdâtre. Foraminifera: 

Nodosaria hexagona (TSCHERD.), Frondicularia longissima (K.M. MACL.), Tetrataxis sp. Ostracods: Cavellina unica KOTSCH., Healdia simplex 

ROUNDY, Bairdia beedei ULR. and B. Brachiopods: Globiella hemisphaerium (KUT.), Licharewia regulata (KUT.), Dielasma elongata SCHLOTH. 

165. Greenish grey argillites, grey platy marls. Argilites gris-verdâtre, marnes grises en plaquettes. Foraminifera: Cornuspira megasphaerica GERKE, C. 

microsphaerica K.M. MACL., Calcivertella sp., Frondicularia longissima (K.M. MACL.). Ostracods: Graphiodactylus sp., Moorea facilis SCHN., Healdia 

pseudosimplex KOTSCH., H. postcornuta SCHN., H. subtriangula KOTSCH., Bairdia beedei ULR. and B., Acratia baschkirica KOTSCH. Brachiopods: 

Licharewia rugulata (KUT.), Cleiothyridina pectinifera SOW., Globiella hemisphaerium (KUT.) 

166. Grey and red tarry sandstones with charred vegetal, with intercalations <strong>of</strong> argillites. Grès gris et rouges bitumineux à végétaux fusinisés, avec 

intercalations d’argilites. 

167. Conglomerates, sandstones, argillites, aleurolites gypseous; in <strong>the</strong> upper part, limestones and cavernous dolomites. Conglomérats, grès, argilites, 

aleurolites gypseuses ; dans la partie supérieure, calcaires et dolomies caverneuses. Ostracods: Paleodarwinula martjevi (PAL.), P. sobela (KASH.), 

Prasuchonella scolia (MAND.), P. stelmarta (KASH.) 

168. Limestones, dolomites, marls, argillites, sandstones, conglomerates with gypsum lenses. Calcaires, dolomies, marnes, argilites, grès, conglomérats 

avec lentilles de gypse. Ostracods: Paleodarwinula burajevoensis (PAL.), P. trita (PAL.), Prasuchonella nasuta KOTSCH., Darwinuloides djurtjuliensis PAL., 

Sinusuella pergraphica MAND. 

169. Conglomerates, sandstones, argillites, aleurolites, dolomites, limestones. Conglomérats, grès, argilites, aleurolites, dolomies et calcaires. Ostracods: 

Paleodarwinula biriensis (PAL.), P. parphenovi (BEL.), P. forschi (PAL.), Prasuchonella kamyschincaensis PAL. 

170. Dolomites, limestones, marls, argillites, sandstones with gypseous cement, aleurolites. Dolomies, calcaires, marnes, argilites, grès à ciment gypseux, 

aleurolites. Ostracods: Paleodarwinula angusta (MAND.), P. biriensis (PAL.), P. forschi (PAL.) 

Table 8 

Eastern European basin, Samara and Volgograd (first part) belts 

Bassin Est-Européen, Ceintures de Samara et de Volgograd (première partie)


171. Bivalves: Palaeomutela ovalis AMAL., P. curiosa AMAL., P. plana AMAL., P. obloqua (AMAL.), Palaeanodonta fischeri AMAL.; Ostracods: 

Suchonellina trapezoida (SHARAP.), S. inornata SPIZH., S. fragilis (SCHNEID.), S.(?) praelongata (ZEK.), S. vladimirinae (BEL.), Suchonella auriculata 

(SHARAP.), S. grandis (BEL.), Darwinuloidea svijazhicus (SHARAP.), Sinusuella vjatkensis POSNER; Conchostraca: Pseudes<strong>the</strong>ria cf. cicatriosa (NOV.), 

Ps. cf. elongata (NETSCH.), Ps. cf. lineata (LUTK.), Limnadia cf. maitchatica NOV. Vertebrates s.l.: Chroniosuchus uralensis TVERD., Ch. paradoxus 

VJUSCH., Dvinosaurus sp., Inostrancevia uralensis TAT., Scutosaurus sp., Chthonosaurus velocidaens VJUSCH., Rhaphanodon ultimus (TCHUD. and 

VJUSCH.), Rh. tverdochlebovae IVACH., Kotlassia prima AMAL. 

172. Bivalves: Palaeomutella obliqua AMAL., P. keyserlingi AMAL., P. ovalis AMAL. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH., S. 

parallela var. typica (LUN.), S. undulata (MISH.), S. spizharskyi (POSNER), S. cultella (MISH.), Prasuchonella stelmachovi (SPIZH.), P. sulacensis 

(STAROZH.), Suchonella grandis BEL., Darwinuloides buguruslanicus KASH., <strong>Permian</strong>a oblonga (POSNER). Conchostraca: Limnadia turaica NOV., L. 

uralensis NOV., Pseudes<strong>the</strong>ria nordvikensis (NOV.), Ps. lineata (LUTK.). Arthropods: Polygrapta aff. chatangensis NOV. Vertebrates s.l.: Chroniosaurus 

dongusensis TVERD., Dvinosaurus sp., Scutosaurus sp., Microphon exigus IVACH. 

173. Amanak formation: red or variegated aleurolites, clays, sandstones, limestones and marls. Formation d’Amanak: aleurolites, argiles, grès, calcaires 

et marnes rouges ou bariolés. Ostracods: Paleodarwinula elongata (LUN.), P. chramovi (GLEB.), P. torensis (KOTSCH.), Prasuchonella nasalis (SHAR.). 

174. Paleodarwinula fragiliformis (KASH.), Placidea lutkevichi (SPIZH.). 

175. Bivalves: Palaeomutela extensiva GUS., P. krotovi NETESCH. Conchostraca: Ulugkemia petri NOV., Rossoes<strong>the</strong>ria acutangulata NOV. Vertebrates 

s.l.: Platyops stuckenbergi TRAUT., Melosaurus kinelensis VJUSCH. 

176. Ostracods: Paleodarwinula torensis (KOTSCH)., P. kassini (BEL.), P. perlonga (SHAR.), P. fragilis var. angusta (LUN.), P. chramovi (GLEB.), P. 

elongata (LUN.), P. fragiliformis (KASH.), P. faba (MISH.), P. imposter (MISH.), Volganella magna SPIZH., Prasuchonella nasalis (SHAR.). Vertebrates s.l.: 

Platysomus biarmicus EICHW., Kargalichthis efremovi MINICH, Melosaurus vetustus KONZH., Platyops stuckenbergi TRAUT., Doliosauriscus admanteum 

ORL., Syodon biarmicum KUT., Titanophoneus potens EFR., Zygosaurus lucius EICHW., Discosauricus netscha jewi RJAB. 

177. Soksk formation: aleurolites with gypsum, sandstones. Formation de Soksk: aleurolites avec gypse, grès. Ostracods: Paleodarwinula elongata 

(LUN.), P. tichonovichi (BEL.), Darwinuloides tscherdynzevi KASH.; Miospore complex I 2 (see Table 2). 

178. Sosnovsk formation: anhydrite, dolomites, limestones, thin beds <strong>of</strong> clays, marls, aleurolites; inclusions <strong>of</strong> salt, gypsum. Formation de Sosnovsk: 

anhydrite, dolomies, calcaires, lits fins d’argiles, marnes, aleurolites, inclusions de sel et de gypse. Ostracods: Paleodarwinula fainae (BEL.), P. alexandrinae 

(BEL.), P. varsan<strong>of</strong>ievae (BEL.), Prasuchonella onega (BEL.), P. belebeica (BEL.), P. tichwinskaja (BEL.), Darwinuloides sentjakensis (SHARAP.) 

179. “hydrochemistry” formation: anhydrites with rare beds <strong>of</strong> clayed dolomites, salts, aleurolites, sandstones. Formation “ hydrochimique ”: anhydrites 

avec rares lits de dolomies argileuses, sels, aleurolites, grès. 

180. Ostracods: Paleodarwinula fainae (BEL.), P. alexandrae (BEL.), P. justa (SPIR. and MOLOST.), P. cuneata (KOTSCH.), P. lira (SPIR. and 

MOLOST.), Prasuchonella belebeica (BEL.), P. tichwinskaja (BEL.). Vertebrates s.l.: Platyops stuckenbergi TRAUT., Tokosaurus perforatus TYERD. and 

IVACH., Belebey maximovi TVERD., B. vergrandis IVACH.; Miospore complex I 2 (see Table 2) +Granizonospora vulgaris LUBER f. elegans STAN., 

Vittatina elegans ZAUER, V. scutata STAN., Cordaitina rugulifera (LUBER), C. stenolimbata WAR., Protohaploxypinus perfectus (NAUM.) SAMOIL. 

181. Barbashinsk unit: light grey pelitic dolomites; dark grey clays, green-grey sandstones, lenses <strong>of</strong> gravelles. Unité Barbashinsk : dolomies pélitiques gris 

clair, argiles gris foncé, grès gris-vert, lentilles de gravelles. Foraminifera: Hyperammina borealis GERKE, Nodosaria noinskyi TSCHERD. Ostracods: 

Healdia simplex ROUNDY, H. subtriangula KOTSCH. 

182. Kamyshlinsk unit: light grey organodetritic limestones; dark grey to black clays; thin bedded marls, rare thin beds <strong>of</strong> aleurolites and sandstones; some 

inclusions <strong>of</strong> anhydrite and salt. Unité Kamyshlinsk : calcaires organodétritiques gris clair, argiles noires à gris foncé, marnes finement litées, rares bancs 

fins d’aleurolites et de grès, quelques inclusions d’anhydrite et de sel. Foraminifera: Saccammina parvula GERKE, Nodosaria pseudoconcinna K.M. MACL. 

Brachiopods: Lingula semivelata TSCHERD. Ostracods: Pseudoparaparchites furmidabilis SCHN., Amphissites tscherdyncevi POSNER, Healdia oblonga 

KOTSCH., Bairdia sobolevensis KHIV. 

183. Baytugansk unit: light grey thin grained limestones, clays, marls, dolomites; calcaires à grain fin, argiles, marnes et dolomies gris clair. Foraminifera: 

Cornuspira microsphaerica GERKE. Ostracods: Amphissites tscherdyncevi POSNER, Actuaria diffusa SCHN., Monoceratina fasgiata KOSCH. 

184. Miospore complex I 1 (see Table 2)+Remysporites psilopterus LUBER, Azonaletes levis LUBER, Vittatina persecta ZAUER, Cordaitina rotata 

(LUBER) MEDV., Striatolebachites varius (ZAUER) WAR., Striatohaplopinites prolixus LUBER, Protohaploxypinus tecturatus LUBER 

185. Kalinovsk formation: interbedded <strong>of</strong> light grey limestones, marls, clays; rare beds <strong>of</strong> aleurolites, sandstones. Formation Kalinovsk : calcaires, marnes 

et argiles gris clair interstratifiées ; rares lits d’aleurolites et de grès. Brachiopods: Lingula orientalis GOL., Dielasma clongata SCHLOTH. Ostracods: 

Cavellina unica KOTSCH., Healdia simplex ROUNDY, H. postcornuta SCHN., H. subtriangula KOTSCH., Bairdia beedei ULRICH and BASSLER, Acratia 

stricta SCHN. Vertebrates s.l.: Phreatophasma aenigmatum EFR., Phreatosuchus qualeni EFR. 

186. Ostracods: Paleodarwinula procera (MAND.), P. parphenovae (BEL.), P. abunda (MAND.), P. angusta (MAND.), P. pyriformis (KASH.), 

Prasuchonella stelmarta (KASH.), Sinusuella pergraphica (MAND.); Miospore complex H (see Table 2)+Acanthotriletes osmundae SAMOIL., Granizonospora 

vulgaris WAR., Cirratriradites procumbens (LUBER) WAR. 

187. Ostracods: Suchonellina trapezoidea (SHAR.), S. parallela SPIZH., S. undulata MISHINA, Wjatkellina fragilina (BEL.), W. fragilis (SCHN.), W. 

vladimirina (BEL.), Dvinella ex gr. cyrta (ZEKINA), Suchonella typica SPIZH., Tatariella angulata MISHINA, Placidea lutkevichi (SPIZH.), Tscherdynzeviana 

pulchra (BEL.); in eastern zone, endemic sculptured cy<strong>the</strong>rides, Tscherdynzeviana pennata KOTSCH., Netschajewiana costata KOTSCH., Kashevaroviana 

delicata MOLOST., Pamilio primaris delicata MOLOST. Bivalves: Palaeomutela inostranzevi AMAL., P. orthodonta AMAL., P. ovalis AMAL., P. plana 

AMAL., Palaeanodonta fisheri AMAL., P.okensis AMAL., Opokiella tschernyschevi PLOTN. Conchostraca: Limnadia angulidorsa NOVOJ., L. maitschatica 

NOVOJ., L. (Falsisca) furaica NOVOJ., Megasitum kaljugense NOVOJ., M. vanum NOVOJ. Vertebrates s.l.: Chroniosuchus uralensis TVERD., Kotlassia sp., 

Raphanodon sp., Scutosaurus sp., Inostrancevia uralensis TAT., Scylacosuchus orenburgensis TAT., Dicynodon cf. amalitzkii SUSCH.; Isadia aristoviensis A. 

MINICH, Wodnika invicta A. MINICH. 

188. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH, S. undulata (MISH.), S. daedala (MISH.), S. cultella (MISH.), S. degitalis (MISH.), 

S. futschiki (KASH.), S. sizharskyi (POSNER), Prasuchonella stelmachovi (SPIZH), P. sulacensis (STAROZ.), Darwinuloides buguruslanicus KASH., 

<strong>Permian</strong>a elongata POSNER, P. oblonga POSNER, Tscherdynzeviana pylchra BEL., Tsch. buzulukensis KASH.; endemic cy<strong>the</strong>rides (ostracods) in <strong>the</strong> eastern 

zone – Tscherdynzeviana busulucensis KASH., Tsch. pennata KOTSCH., Tsch. abscondita KOTSCH., Netschajewiana costata KOTSCH., Nuguschia sinizyni 

KOTSCH., Kaschevaroviana delicata MOLOST., K. pectunata MOLOST., Jungumulus mirus MOLOST., <strong>Permian</strong>ella inilaterata MOLOST., Pamilio 

primaris MOLOST. Bivalves: Palaeomutela inostranzevi AMAL., P. keyserlingi AMAL., Palaenodonta fischeri AMAL., Opokiella ignatjevi GUSEV.


553 

Conchostraca: Siberioleaia oblonga (MITSCH.), Hemicycloleaia intermediata (MITSCH.), Pseudos<strong>the</strong>ria cicatricosa (NOJ.). Vertebrates s.l.: Chroniosaurus 

dongusensis TVERD., Rophanodon tverdochlebovae IVACH., Microphon exiquus IVACH.; Isadia aristoviensis A. MINICH, I. suchonensis A. MINICH, 

Toyemia tverdochlebovi MINICH; Flora s.l.: Pursongia amalitzkii ZAL., Cordaites clercii ZAL.; Limitisporites sp., Jugasporites sp., Lueckisporites virkkiae 

POT. and KREMP, Taeniaesporites albertae JANS. 

189. Western part: lower half, obliquely layered red sandstones, aleurolites, clays and fine-pebble conglomerates; upper half, red aleurolites, clays, fine and 

more seldom coarse-grained sandstones; central part: high thickness, aleurolite-clayey composition, conglomerate lens occurrence and widely spread nodular 

limestones; eastern part: lower half, conglomerate/sandstone sequence with rare interlayers <strong>of</strong> red-brown clays and marls; upper half, marl-clay-sandstone 

sequence with interlayers <strong>of</strong> grey clayey limestones and aleurolites. Partie occidentale : moitié inférieure, grès à stratifications obliques, aleurolites, argiles 

et conglomérats à galets fins ; dans la moitié supérieure, aleurolites rouges, argiles, grès à grain fin et plus rarement à grain grossier ; partie centrale : 

grande épaisseur d’aleurolites argileuses, lentilles de conglomérats et calcaires noduleux largement répandus ; partie orientale : moitié inférieure, séquence 

de conglomérats – grès avec rares niveaux de marnes et argiles rouge-brun ; moitié supérieure, séquence, marnes – argiles – grès avec niveaux de calcaires 

argileux gris et aleurolites. Ostracods: Paleodarwinula fragiliformis (KASCH.), P. elongata (LUNJAK), P. fragilis var. angusta (SCHN.), P. inornatinae 

(LUNJAK), P. chramovi (GLEB.), P. torensis (KOTSCH.), P. faba (MISCH.), P. impostor (MISCH.), P. defluxa (MISCH.), P. arida (MOLOST.), P. obvia 

(MOLOST.), P. gariainovi (MOLOST.), Prasuchonella nasalis (SHARAP.), Darwinuloides buguruslanicus KASCH., Sinusuella ignota SPIZH., Placidea 

lutkevichi (SPIZH.). Bivalves: Abiella subovata (JONES), Microdantella microdonta (KHALFIN), M. plotnikovskiensis (FED.), Palaemutela doratiformis 

GUSEV, P. parallelogramma KULEVA, P. gusevi KULEVA, Anthraconaia verneuili (AMAL.), A. aktubensis GUSEV, Anthraconauta (Prokopievskia) 

pseudophillipsi FEDOTOV. Conchostraca: Pseudos<strong>the</strong>ria angulata (LUTK.), Sphaeres<strong>the</strong>ria belorussica NOVOJ., Triedrolophus tverdochlebovi MOLIN, 

Pseudos<strong>the</strong>ria otchevi MOLIN, Megasitum jaroslavlense NOVOJ. Vertebrates s.l.: Mnemeiosaurus jubilaevi NOVI., Melosaurus sp., Deuterosaurus sp., 

Platyops sp., Notosyodon sp. Flora s.l. : Lebachiacites pulcherrimus SAUER, Striatolebachites, Jugasporites sp., Faunipollenites sp., Taeniaesporites albertae 

JANS., Vittatina striata LUBER f. angusticostata SAUER, Platysaccus?, Alisporites striatohaplopinites. The upper Urzhum horizon is characterized in some 

localities by endemic Cy<strong>the</strong>rides (ostracods). La partie supérieure de l’horizon Urzhum est caractérisé par des Cythéridés (ostracodes) endémiques : 

Tscherdynzeviana squamosa KOTSCH., Wetluginella (?) tatarica MOLOST., Kaschevaroviana delicata MOLOST., K. pectunata MOLOST. 

190. Western part: lagoonal-continental formations; red, less frequently grey clays, aleurolites, sandstones, and, to a smaller extent, limestones, marls, 

dolomites; central part: clays, aleurolites and sandstones with rare interlayers <strong>of</strong> limestones and marls. Eastern part : carbonate-terrigenous sequence composed 

<strong>of</strong> interlaid brown, lilac and grey clays, aleurolites, marls, limestones, sandstones / sandstones, gravelstones and conglomerates / monotonous interlayers <strong>of</strong> 

concretionary and bedded brown limestones. Partie occidentale : formations lagunaires – continentales ; argiles, aleurolites, grès rouges ou plus rarement 

gris, plus rarement calcaires, marnes et dolomies ; partie centrale : argiles, aleurolites et grès avec rares niveaux de calcaires et de marnes ; partie 

orientale : séquence carbonatée – terrigène, composée deséries interstratifiées grises, brunes et lilas, d’aleurolites – marnes – calcaires – grès /grès – 

graviers – conglomérats / séries monotones de calcaires bruns lités ou en concrétions. Ostracods: Paleodarwinula fainae (BEL.), P. alexandrinae (BEL.), P. 

irenae (BEL.), P. cuneata (KOTSCH.), P. tichonovichi (BEL.), P. tuimazensis (KOTSCH.), P. varsan<strong>of</strong>ievae (BEL.), P. inornatinaeformis (BEL.), P. 

chramovella (BEL.), P. aronovae (BEL.), P. persimplex (KOTSCH.), P. procurva (KOTSCH.), Garjainowula lija (SPIR. and MOLOST.), G. ex. gr. gracilis 

(SPIR. and MOLOST.), Prasuchonella tichwinskaja (BEL.), P. belebeica (BEL.), Darwinuloides sentjakensis (SCHAR.), Placidea ex gr. lutkevichi (SPIZH.), 

Schneideria kazanica KOTSCH. Bivalves: Palaeanodonta rhomboidea (NETSCH.), Palaemutela celebrata GUSEV, P. quadrata KULEVA, P. novalis 

NETSCH., P. umbonata (FISCHER), P. krotovi NETSCH, P. olgae GUSEV, P. doratiformis GUSEV. Conchostraca: Palaeolimnadiopsis eichwaldi 

(NETSCH.), Clyptoasmussia exiqua (EICHW.), Cyclotungusites kazanensis MOL., Ulugkemia borisi NOVOJ., U. orengurgiana MOLIN, U. petri NOVOJ., 

Sphaeres<strong>the</strong>ria ikensis MOLIN, Pseudes<strong>the</strong>ria kaslanica MOLIN. Vertebrates s.l.: Platyops sp., Deuterosaurus sp., Tapinocephalidae. Flora s.l.: Cordaitina 

sp., Limitisporites leschikii KLAUS, Gigantosporites hallsta<strong>the</strong>nsis KLAUS, Striatites marginalis KLAUS, Striatohaplopinites latissimus LUBER, Striatopiceites 

suchinensis SED., Vittatina striata LUBER, Cyclogranisporites, Acanthotriletes, Azonaletes; Carpoli<strong>the</strong>s sp., Phylladoderma sp., Noeggerathiopsis sp., 

Lepeophyllum sp., Samaropsis sp., Paracalamites sp., Odontopteris sp., Equisetina sp., Rufloria sp., Taeniopteris sp., Pecopteris sp., Voltzia mamadyschensis 

ZAL., Samaropsis irregularis NEUB., Psygmophyllum expansum BRONGN., Odontopteris cf. rossica ZAL., Ullmannia biarmica EICHW. 

191. Western part: grey sandstones, aleurolites, clays and limestones. Partie occidentale: grès gris, aleurolites, argiles et calcaires. Ostracods: 

Pseudoparaparchites formidabilis SCHN., Aecuminella bella KHIV., Cornigella valosa KHIV., Kirkbya rara KHIV., Amphissites tscherdynzevi POSNER, 

Moorea facilis SCHN., Cavellina grandis SCHN., Cavellina unica KOTSCH., Healdia postcornuta SCHN., H. oblonga KOTSCH., Healdia simplex 

ROUNDY, H. pseudosimplex KOTSCH., H. subtriangula KOTSCH., H. reniformis SCHN., Healdianella vulgata KOTSCH., Cribroconcha urschakensis 

KOTSCH., Bairdia beedei ULRICH and BASSLER, B. garrisonensis UPSON, B. pompiliodes HARLTON, Acratia baschkirica KOTSCH., Fabalicypris 

crepidalis KOTSCH., Actuaria diffusa SCHNEIDER, A. secunda KOTSCH., Monoceratina fastigiata KOTSCH., M. faveolata KOTSCH., M. parvula 

KOTSCH., M. exilis SCHNEIDER, Fascianella notabilis SCHN. Brachiopods: Glomospira sp., Ammodiscus cf. bradynus SPANDEL, Verneuilinoides 

kasanica UCHARS. Foraminifera: Cornuspira megasphaerica GERKE, C. microsphaerica K.M. MACLAY, Calcivertella kasanica K.M. MACLAY, 

Nodosaria cushmani PAALZOV, N. forcimeniformis K.M.–MACLAY, N. hexagona (TSCHERD.), N. aff.noinskyi TSCHERD., N. krotovi TSCHERD., 

Pseudonodosaria lata K.M. MACLAY, Fronticularia longissima (K.M. MACLAY), Fr. geinitzinaeformis (K.M. MACLAY), Geinitzina spandeli TSCHERD., 

Tetrataxis sp., Tristis permiana GERKE. Brachiopods: Cleiothyridina pectinifera SOW, Cl. semiconcava WAAGEN, Cl. reyssiana KEYS., Stepanoviella 

hemisphaerium KUTOR., Pr. (Cancrinella) cancrini VERNEUIL, Pr. koninckianus KEYS., Licharewia rugulatia KUTOR., L. curvirostris VERNEUIL, 

Dielasma elongata SCHLOT., D. elliptica NETSCH., D. angusta NETSCH., Lingula orientalis GOLOW., Aulosteges horrescens VERNEUIL. Bivalves: 

Leibea hausmanni GOLDF., Allorisma elegans KING, Pseudobakewellia sulcata (GEIN.), Ps. ceratophragaeformis NOINSKI, S. tutchburia tschernyschevi 

LICH. Central and eastern parts: lagoonal-deltaic and littoral-marine grey-coloured sandstones, aleurolites, clays, marls, limestones and dolomites. Parties 

centrale et orientale : série grise, laguno-deltaïque et littorale composée de grès, aleurolites, argiles, marnes, calcaires et dolomies. Ostracods: 

Paleodarwinula cuneata (KOTSCH.), P. fainae (BEL.), P. irinae (BEL.), Prasuchonella belebeica (BEL.), Darwinuloides cf. sentjakensis (SHARAP.). 

Bivalves: Paleomutela sintasensis GUSEV, P. ovataeformis GUSEV. Flora s.l.: Pecopteris anthriscifolia ZAL., Sphenophyllum sp., Asterodiscus; Cordaitina 

uralensis (LUBER) SAMOIL., C. cubrotata (LUBER) SAMOIL.; Ginkgocycadophytus erosus (LUBER) SAMOIL.; Protodiploxypinus elongatus SAMOIL., 

Protopodocarpus alatus SAMOIL.; Azonalites (Rugosina) tenuis LUBER, A. (Tenuella) levis LUBER; Azonotriletes cf. resistens LUBER, A. osmundes 

SAMOIL.


192. Western part: red terrigenous rocks: clays, aleurolites, sandstones, rare marls, limestones and dolomites interlayers; intensive gypsification in <strong>the</strong> lower 

part. Partie occidentale : roches terrigènes rouges : argiles, aleurolites, grès, rares marnes, calcaires et dolomies interstratifiées ; gypsification intense dans la 

partie inférieure. Ostracods: Paleodarwinula abunda (MAND.), P. angusta (MAND.), P. parphenovae (BEL.), P. procera (MAND.), P. lancetiformis (KASH.), P. 

trita (PALANT), P. burjevoensis (PALANT), P. lubimovae (KASH.), Garjainovula lija (SPIR. and MOLOST.), G. kulevae (SPIR. and MOLOST.), G. gracilis 

(SPIR. and MOLOST.), Prasuchonella stelmarta (KASH.), Pr. kargalensis (KOTSCH.), Pr. kamyschencaensis (PALANT), Darwinuloides djurtjuliensis 

PALANT, Sinusuella pergraphica MAND. Bivalves: Paleomutela attenuata GUSEV, P. ovataeformis GUSEV, P. sintasensis GUSEV. Miospores complex H 1 + 

H 2 (see Table 2). Central part: red clays, aleurolites and sandstones; eastern part: red conglomerates with flint pebbles, sandstones, aleurolites and marls with rare 

thin limestone interlayers; reddish-brown argillites with interlayers <strong>of</strong> brownish-grey sandstones and grey limestones. Partie centrale : argiles, aleurolites et grès 

rouges ; partie orientale : conglomérats rouges avec galets de silice, grès, aleurolites et marnes avec rare niveaux fins de calcaires ; argilites brun-rougeâtre avec 

niveauxdegrès gris-brunâtre et de calcaires gris. Ostracods: Paleodarwinula abunda (MAND.), P. angusta (MAND.), P. lancetiformis (KASH.). 

193. Ostracods: Suchonellina inornata SPIZH., Suchonellina parallela SPIZH., S. cf. futschiki (KASH.), S. spizharskyi (POSNER), Prasuchonella 

stelmachovi (SPIZH.), Darwinuloides svijaazhicus SHARAP., Paleodarwinula elongata (LUN.), Paleodarwinula fragiliformis (KASH.), Suchonella nasalis 

SHAR., 

194. Ostracods: Amphissites tscherdynzevi POSNER, Bairdia beedei ULRICH and BASSLER, Bairdianella semilunulata KOTSCH., Actuaria diffusa 

SCHN.; Miospore complex I 1 (see Table 2) +Calamospora sp., Cyclogranisporites osmundae SAMOIL., Apiculatisporites tenuispinosus (WALTZ.) SIV., 

Cirratriradites procumbens LUBER, Azonaletes tenuis LUBER, Vittatina cicinnata LUBER, V. striata LUBER, Florinites luberae SAMOIL., Cordaitina sp., 

Platysaccus latus LUBER, Vesicaspora schemeli KL., Falcisporites zapfei POT. and KL., Caytonispollenites sp., Striatohaplopinites perfectus NAUM., 

Striatopodocarpites tojmensis SED., Lueckisporites sp., Taeniaesporites sp. 

195. Miospore complex H (see Table 2) +Punctatostorites sp., Cyclogranisporites polypyranus (LUBER) SIV., Spinosisporites cornutus K.-M., 

Verrucosisporites grumosus IBR., Cirratriradites procumbens LUBER, Granulatisporites sp., Azonaletes irregulariplicatus SAMOIL., Ginkgocycadophytus 

sp., Vittatina crassa ZAUER, V. striata LUBER, V. laxa ZAUER, Cordaitina sp., Vesicaspora sp., Alisporites sp., Striatohaplopinites glomus HART., 

Protohaploxypinus latissimus (LUBER) SAMOIL., Hamiapollenites bullaeformis (SAMOIL.) JANS. 

196. Stomochara abrasa SAID., Stm. cybaea KIS., Stm. diserta SAID., Horniella concinna KIS.; Miospore complex J 2 (see Table 2) +Caytonipollenites 

sp., Calamospora sp. Cyclogranisporites polypyrens LUBER, Apiculatisporites sphaericus VIRB., Granulatisporites dubia PRESL., Falcisporites sp., 

Fusacolpites permicus DIUP., Platysaccus sp., Striatopiceites sp., Lueckisporites virkkiae POT. and KR., Taeniaesporites sp., Vittatina vittifer LUBER. 

197. Ostracods: Suchonellina inornata (SPIZH.), S. parallela (SPIZH.), S. stelmachovi (SPIZH.), Darwinuloides tataricus POSNER, D. svijazhicus SHAR. 

198. Ostracods: Paleodarwinula fragiliformis (KASH.), P. elongata (LUN.), Suchonellina nasalis (SCHAR.). 

199. Ostracods: Amphissites tscherdynzevi POSNER, Healdia simplex ROUNDY, Healdia subtriangula KOTSCH., Healdianella ex gr. osagensis 

KELLETT, Healdianella ex gr. panda KOTSCH., Healdianella parallela KNIGHT, Healdianella vulgata KOTSCH., Cavellina? cf. unica KOTSCH., 

Cavellina ex gr. edmistonae (HAR. and LAI.). Brachiopods: Cancrinella cancrini VERN., Beecheria netschaevi GRIG., Cleiothyridina reussiana (KEIS.), C. 

pectinifera (SOW.). Bivalves: Pseudomonotis elengatula NETSCH., Parallelodon kingi VERN., P. cf. striatus (SCHLOTH.), Edmondia elongata HOUSE, 

Bakevellia (Bakevellia) ceratophaga (SCHLOTH.). Bryozoans: Dyscritella incrustata MOR. Foraminifera: Palaeonubecularia uniserialis REITL., P. fluxa 

REITL., Geinitzina pusilla GROZD., G. spandeli TSCHERD., G. cf. postcarbonica SPAND., Nodosarina elabugae TSCHERD., N. netschevi TSCHERD., 

Spandellina ex gr. cordiformis GERCE; Miospore complex I (see Table 2) +Leiotriletes osmundae SAMOIL., Azonamonoletes marattiformis SAMOIL., 

Azonaletes tenuis LUBER, Vittatina vittifer LUBER, Ginkgocycadophytus sp., Vitreisporites sp., Florinites sp., Striatopinites dvinensis SED., Protohaplopinites 

tecturatus LUBER, Protodiplopinites silvestritypus SAMOIL., Protocedrus parviextensisaccus SAMOIL. 

200. Miospore complex J (see Table 2)+Granulatisporites sp., Punctatisporites sp., Verrucosisporites vetlugencis (MAL.) WAR., Densoisporites nejburgii 

(SCHULTZ) BALME, Spinosisporites (MAL.) WAR., Vittatina striata LUBER, V. costabilis WILS., Cordaitina sp., Podocarpites alatus LUBER, Gardenasporites 

heisseli KL., Striatopodocarpus sp., Hamiapollenites sp., Striatolebacheites sp. 

201. Miospore complex J (see Table 2) +Vittatina costabilis WILS., V. vittifer LUBER, V. striata LUBER, Vesicaspora schemeli KL., Vestigisporites 

minutus KL., Gardenasporites heisseli KL., G. leonardii KL., Falcisporites sp., Podocarpites alatus (LUBER), Striatopodocarpites cancellatus KL., 

Striatolebacheites sp., Lueckisporites virkkiae POT. and KL., Vitreisporites sp. 

202. Miospore complex I 2 (see Table 2)+Punctatisporites sp., Raistrickia cardi<strong>of</strong>ormis (LUBER), Entylissa sp., Vittatina sp., Lueckisporites virkkiae POT. 

and KL., Vitreisporites sp., Striatolebacheites sp., Striatopodocarpites ellipticus ZAUER, Acritarches. 

203. Ostracods: Paleodarwinula sp., Moorea cf. facilis SCHN.; Miospore complex I 1 (see Table 2) +Vittatina costabilis WILS., Entylissa sp., Cordaitina 

sp., Gardenasporites sp., Platysaccus papilionis POT. and KL., Alisporites sp., Vitreisporites sp., Lueckisporites virkkiae POT. and KL., Striatolebacheites sp., 

Taeniaesporites noviaulensis LESCHIK, Striatopodocarpites ellipticus ZAUER, S. cancellatus KL., Acritarches 

204. Miospore complex H (see Table 2)+Leiotriletes sp., Lycospora sp., Zonomonotriletes granulatus (SAMOIL.) DIBN., Vittatina cincinnata LUBER, V. 

vittifer LUBER, V. striata LUBER, V. costabilis WILS., Platysaccus sp., Limitisporites sp., Striatohaplopinites micros HART., S. verus EFR., Striatites minor 

KL., Striatodiplopinites elongatus (LUBER) SAMOIL., Acritarches.


555 

Table 9 

Eastern European basin, Volgograd (second part) belt 

Bassin Est-Européen, Ceinture de Volgograd (deuxième partie) 

205. Ostracods: Suchonella futschiki (KASH.), Darwinuloides tataricus POSNER, D. swijazhicus SHARAP. 

206. Ostracods: Suchonellina futschiki (KASH.) 

207. Ostracods: Darwinuloides edmistnus (BEL.) 

208. Brachiopods: Cleiothyridina royssiana (KEYS.), Beecheria netschajewi GRIG. Bryozoans: Dyscritella incustata MOROZ. 

209. Ostracods: Darwinuloides edmistonae BEL., Darwinuloides sentjakensis (SHAR.), Darwinuloides triangula BEL., Paleodarwinula accomo<strong>data</strong> 

(STAR.), Paleodarwinula alexandrinae (BEL.), Paleodarwinula inornatina (BEL.), Paleodarwinula varsan<strong>of</strong>ievae (BEL.), Paleodarwinula vinocurovi 

(BEL.), Sinusuella ignota (SPIZH.), Suchonella belebeica BEL., Prasuchonella onega (BEL.), Suchonella serpula BEL., Palaeanodonta rhomboidea 

(NECH.), Palaeomutela cf. umbonata (FISCH.), Palaeomutela cf. vjatkensis GUS., Palaeomutela ex gr. dorati<strong>of</strong>ormis GUS., Palaeomutela ex gr. krotovi 

NECH. 

210. Paleodarwinula irinae (BEL.), P. varsan<strong>of</strong>ievae (BEL.), Paleodarwinula accomo<strong>data</strong> (STAR.), Paleodarwinula ex gr. tichwinskaje (BEL.), 

Paleodarwinula isetica (STAR.), Paleodarwinula varsan<strong>of</strong>ievae (BEL.), Paleodarwinula. irinae (BEL.), Prasuchonella onega (BEL.), Darwinuloides 

sentjakensis (SHARAP.), Darwinuloides edmistonae BEL., Darwinuloides triangula BEL., Sinusuella cf. ignota SPRZH., Suchonella belebeica BEL., 

Suchonella sacmarensis STAR., pelecypods, miospores 

211. Ostracods: Paleodarwinula angusta (MAND.), P. procera (MAND.), P. abunda (MAND.), P. sobela (KASH.) 

212. Ostracods: Paleodarwinula cf. angusta (MAND.), P. abunda (MAND.), Prasuchonella ex gr. scolia (MAND.) 

211 – 212. At Lugovskaya 1 well: Ostracods: Paleodarwinula angusta (MAND.), Paleodarwinula burajevoensis (PAL.), Paleodarwinula cf. pyriformis 

(KASH.), Paleodarwinula cf. trita (PAL.), Paleodarwinula fainae (BEL.), Paleodarwinula lanzetiformis (KASH.), Paleodarwinula lubimovae (KASH.). 

Bivalves: Palaeomutela cf. stegocephalum NECH., miospores and conchostraca. 

213. Bivalves: Palaeomutela verneuili (AMAL.), P. ovalis AMAL. Ostracods: Suchonellina parallela SPIZH. Flora s.l.: Cladophlebis kamiensis VLAD. 

214. Ostracods: Paleodarwinula fragiliformis (KASH.). Flora s.l.: Cladophlebis kamiensis VLAD., Voltzia mamadyschensis ZAL., Cordaicarpus 

czakaensis ZAL., C. burensis ZAL. 

215. Bivalves: Antraconauta uslonensis GUS. Ostracods: Darwinuloides elegantella BEL. 

216. Bivalves: Palaeomutela umbonata (FISCHER), P. attenuata GUS., P. pseudoumbonota GUS. Ostracods: Paleodarwinula inornatinaeformis (BEL.), 

Prasuchonella onega (BEL.), Darwinuloides sentjakensis SHARAP. Flora s.l.: Cladophlebis kamiensis VLAD., Phylladoderma sp. 

217. Bivalves: Palaeomutela stegocephalum NETSCH., F. sintasensis GUS., P. ovatiformis GUS., Palaeanodonta (Antraconaia) rhomdoidea (NETSCH.). 

Ostracods: Paleodarwinula abunda (MAND.), P. sobela (KASH.), P. pergusta (KASH.). Flora s.l.: Knorriopsis sp., Viatcheslavia sp.


Table 10 

Eastern European Basin, Pripyat depression. Characteristic fossils <strong>of</strong> Western European Basin 

Bassin Est-Européen, dépression de Pripyat. Fossiles caractéristiques du Bassin Ouest-Européen 

218. Foraminifera: Nodosaria sp. 

219. Foraminifera: Haplophragmoides probata SCHERP., H. lithuanicus UCHAR., Ammobaculites rectus UCHAR., Trochammina aff. pulvilla CRESP. 

and PARR., Verneuilinoides andrei UCHAR., V. distinctus UCHAR. 

220. Foraminifera: Ammodicus bradynius (SPAND.), Haplophragmoides probata SCHERP., H. lithuanicus UCHAR., Trochammina aff. pulvilla CRESP. 

and PARR., Nodosaria permiana (SPAND.), N. conicodensestriata PAALZ., N. scharpi K.M. MACL., Dentalina permiana JONES, Lunucammina hastata 

SCHERP., Spanaelina delicatula (JURK.). 

221. Bivalves: Liebea squamosa (SOW.), L. Septifer (KING), Schizodus obscurus SOW., Sch. truncatus KING, Sch. schlo<strong>the</strong>imi (GEIN.) 

222. Bivalves: Pseudobakewellia ceratophagaeformis NOIN., Ps. antiquaeformis NOIN., Schizodus obscurus SOW., Sch. schlo<strong>the</strong>imi (GEIN.), Liebea 

squamosa (SOW.), Astartella vallisneriana (KING.), Permophorus costatus (BROWN). 

223. Bivalves: Pseudobakewellina ceratophagaeformis NOIN. 

224. Ostracods: Dorsoobliquella pulchra KNUEPF., Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM. 

225. Ostracods: Dorsoobliquella pulchra KNUEPF., Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Bairdia plebeia REUSS., B. hisingeri 

(MUENST.). 

226. Ostracods: Polycope perminuta KELLETT, Kirkbya permiana (JONES), Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Microcheilinella 

artiensis GUSS., Bairdia plebeia REUSS, Basslerella regularis (RICHTER), Monoceratina parvula KOTSCH. 

227. Bryozoans: Acanthocladia anceps (SCHLOTH.), Fenestella retiformis (SCHLOTH.), Kingopora ehrenbergi (GEIN.). 

228. Brachiopods: Lingula credneri GEIN., Horridonia horrida (SOW.), Strophalosia morrisiana KING, Pterospirifer alatus (SCHLOTH.), Dielasma 

elongata (SCHLOTH.). 

229. Brachiopods: Lingula credneri GEIN. 

230. Flora s.l.: Klausipollenites schaubergeri (POT. and KLAUS.) JANS., Gardenasporites tonsus INOS., KRUZ. and SCHWARZ., Lueckisporites virkkiae 

POT. and KLAUS., Striatopodocarpites tojmensis SED. 

231. Flora s.l.: Lueckisporites virkkiae POT. and KLAUS., Kausipollenites schaubergeri (POT. and KLAUS.) JANS.


557 

Table 11 

Western European Basin, Brest and Kaliningrad belts 

Bassin Ouest-Européen, Ceintures de Brest et Kaliningrad 

232. Foraminifera: Nodosaria permiana (SPANDEL), N. noinskyi TSCHERD., N. geinitzi REUSS, N.exgr.netschajewi TSCHERD., Geinitzina spandeli 

TSCHORD., Dentalina ex gr. bradyi SPANDEL, Ammodiscus ex gr. semiconstrictus WATH. Bivalves: Schizodus ex gr. obscurus SOW., Pseudobakewellia ex 

gr. ceratophagaeformis NOIN. Bryozoans: Fenestella sp. Ostracods: Bairdia sp., Acratia sp. Brachiopods: Horridonia ex gr. horrida (SOW.), Licharevia ex gr. 

rugulata (KUT.) 

233. Foraminifera: Nodosaria sp. 

234. Foraminifera: Nodosaria sp. Bivalves: Liebea squamosa (SOW.). Ostracods: Dorsoobliquella pulchra KNUEPF., Cavellina permiana KOTSCH., 

Healdia dahilgrueni KROEM. 

235. Foraminifera: Haplophragmoides probata SCHAR., H. lithuanicus UCHAR., Ammobaculites rectus UCHAR., Trochaminna aff. pulvilla CRESPIN 

and PARR., Verneuilinoides andrei UCHAR., V. distinctus UCHAR. Bivalves: Liebea squamosa (SOW.), L. septifer (KING.), Schizodus truncatus KING., Sch. 

obscurus SOW., Sch. schlo<strong>the</strong>imi (GEIN.). Ostracods: Dorsoobliquella pulchra KNUEP., Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., 

Bairdia plebeia REUSS, B. hisingeri (MUNST.) 

236. Foraminifera: Ammodiscus bradyinus (SPAND.), Haplophragmoides probata SCHERP., H. lithuanicus UCH., Trochammina aff. pulvilla CRESPIN 

and PARR., Nodosaria permiana (SPANDEL), N. conicodensestriata PAAL., N. scharpi K.M.–MACL., Dentalina permiana JONES, Lunucammina hastata 

SCHERP., Spandolina delicatula (JURK.). Bivalves: Pseudobakewallia ceratophagaeformis NOIN., Ps. antiquaeformis NOIN., Schizodus obscurus SOW., 

Sch. schlo<strong>the</strong>imi (GOIN.), Lisbea squamosa (SOW.), Astartella vallisneriana (KING.), Permophorus costatus (BROWN.). Bryozoans: Acanthocladia anceps 

(SCHLOTH.), Fenestella retiformis (SCHLOTH.), Kingopora ehrenbergi (GEIN.). Brachiopods: Lingula credneri GEIN., Horridonia horrida (SOW.), 

Strophalosia morrisiana KING, Pterospirifer alatus (SCHLOTH.), Dielasma elongata (SCHLOTH.). Ostracods: Polycope perminuta KELLETT, Kirkbya 

permiana (JONES), Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Microchellinella artiensis GUSS., Bairdia plebeia REUSS, Basslerella 

regularis (RICHTER), Monoceratina parvula KOTSCH. Flora s.l.: Klausipollenites schaubergeri (POT. and KLAUS.) JANS., Gardenasporites tonsus INOS., 

KRUZ. and SCHWARZ., Lueckisporites virkkiae POT. and KL., Striatopodocarpites tojmensis SOD., Stepanovites inflatus (BEND. and STOP.) 

237. Bivalves: Pseudobakewellia ceratophagaeformis NOIN. Brachiopods: Lingula credneri GOIN. Flora s.l.: Lueckisporites virkkiae POT. and KL., 

Klausipollenites schaubergeri (POT. and KL.) JANS.


Acknowledgements 

We greatly acknowledge Dr. Maria St Germes for her help 

in <strong>the</strong> Russian translation. We are indebted to Dr. Brigitte 

Meyer-Berthaud (Montpellier University, France) and Dr. 

Denis Vaslet (BRGM, France) for <strong>the</strong>ir critical and constructive 

remarks. 

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