CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834 ...

CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 

1834 (COLEOPTERA: TENEBRIONIDAE: OPATRINI). 

PART 2. COMMENTS, ADDITIONS AND REFERENCES 

INTRODUCTION 

This catalogue should be treated as a summary of 

the nomenclatural and partly faunistic studies on the 

genus Gonocephalum. We hope that the paper will be 

useful for the continuation of the systematic studies on 

this widespread genus. 

Previous revisions related to Gonocephalum species 

occurring in separate geographic regions 

were published over a long period of time (Gridelli 

1945, 1948, Kaszab 1952, Ferrer 1993a, 1995, 

2000). This makes it difficult to compare all of the 

information contained therein. Hence, the purpose of 

our study was to collect the largest possible number of 

published information, both original and secondary 

(quoted by other authors). These data should be 

reviewed during the preparation of a new revision of 

the genus Gonocephalum covering all known species. 

In the first part of the catalogue (Iwan et al. 2010) 

information on the types (including depository places) 

A N N A L E S Z O O L O G I C I (Warszawa), 2011, 61(2): 259-276 

DARIUSZ IWAN 1 , OTTÓ MERKL 2 and JULIO FERRER 3 

1Museum and Institute of Zoology Polish Academy of Sciences, Wilcza 64, 

00-679 Warszawa, Poland; e-mail: darek@miiz.waw.pl 

2Department of Zoology, Hungarian Natural History Museum, H-1088 Budapest, 

Baross utca 13, Hungary; e-mail: merkl@nhmus.hu 

3Department of Entomology. The Swedish Museum of Natural History, 

S-10405 Stockholm, Sweden; e-mail: julio_ferrer@hotmail.com 

Abstract.— The second part of the world catalogue of Gonocephalum Solier, 1834 

contains the list of the type species preserved in the Hungarian Natural History Museum 

(Hungary, Budapest), the list of the papers on Gonocephalum species and additional data 

for 95 species and subspecies. At present, 417 species and subspecies belong to the genus 

Gonocephalum. 

� 

Key words.— Entomology, taxonomy, catalogue, World, Coleoptera, Tenebrionidae, 

Gonocephalum. 

PL ISSN 0003-4541 © Fundacja Natura optima dux 

doi: 10.3161/000345411X584762 

based on published data (both original and secondary 

citations) was published. Information related to data 

from the labels or quotes from the publications were 

preserved into the language in which the text was written 

(eg. French, German, English); such data were published 

in quotation marks. We wanted to allow the 

demonstration of any inaccuracies that may affect the 

nomenclature decisions, and perhaps also the taxonomic 

interpretations (eg interpretation of species, 

subspecies). 

A number of differences and/or incomplete data 

between published information and actual state (storage, 

data labels) must be corrected and supplemented 

by direct study of museum collections. Ottó Merkl verified 

data on the type specimen of Gonocephalum 

species which are deposited in the Hungarian Natural 

History Museum, HNHM (Hungary, Budapest). This is 

presented in this publication. 

At present, 417 species and subspecies belong to 

the genus Gonocephalum, including G. tuberipenne 

and G. tibiale described by Grimm (2011).

260 D. IWAN, O. MERKL and J. FERRER 

CORRECTIONS, ADDITIONAL CITATIONS 

AND FAUNISTICS DATA 

The paper by Iwan et al. (2010) contains catalogue 

of the species belonging to Gonocephalum. The following 

authors’ citations and distributional data are 

presented in addition to the previous paper: 

aequatoriale (Blanchard, 1853). – Ren 2002b: 390; Yang and 

Ren 2005a: 187. 

annamita Chatanay, 1917. – Ren 2002a: 140, 2002b: 390; Ren 

and Yang 2004a: 311; Yang and Ren 2005a: 186; [China: 

Guanxi (Kwangsi)]. 

angolense escalerai Ferrer, 2000 (1993a: 99). – In the 

description erroneously G. subtilistriatum escalerai, 

but in the catalogue of the same paper (Ferrer 2000: 144) 

correctly G. angolense escalerai. 

angusticolle parvithorax Gridelli, 1948. – Medvedev and 

Abdurakhmanov 1988: 89; Abdurakhmanov and Medvedev 

1994: 58. 

arisi Reitter, 1905. – Yakobson 1913: tab. 78, fig. 13; 

Kryzhanovsky 1965: 176; Nepesova 1980: 29; Medvedev 

and Abdurakhmanov 1988: 90; Abdurakhmanov and 

Medvedev 1994: 58. 

assimile (Küster, 1849). – Medvedev and Abdurakhmanov 

1988: 89; Abdurakhmanov and Medvedev 1994: 58. 

besnardi besnardi Kaszab, 1982. – Medvedev and Abdurakhmanov 


bilineatum (Walker, 1858). – Ren and Saihong 2001: 348; Ren 

2001: 89, 2002a: 141, 2002b: 390; Ren and Yang 2004a: 311; 

Yang and Ren 2005a: 187; [China: Zhejiang (Chekiang)]. 

bimaculatum Ferrer, 1995. – Drinkwater 1999: 183. 

buitenzorgense Kaszab, 1952. – The specimen in the HNHM 

has a holotype label. However, holotype (or “type”) was 

inadvertently not designated in the description (“Typus 

befindet sich in der Sammlung des Museums G. Frey in 

München (Männchen) und im Ungarischen Naturwissenschaftlichen 

Museum in Budapest (Weibchen).”) So, 

these two mentioned specimens should be regarded as 

syntypes. 

brachelytra Kaszab, 1952. – Medvedev and Abdurakhmanov 


coenosum Kaszab, 1952. – Ren 1998b: 91, 2002a: 141; Yang 

and Ren 2005a: 187. 

consobrinum Blair, 1923. – Medvedev and Abdurakhmanov 


controversum Gridelli, 1948. – Medvedev and Abdurakhmanov 

1988: 89; Abdurakhmanov and Medvedev 1994: 

58. 

coriaceum Motschulsky, 1854. – Medvedev 1992: 638; Ren 

and Saihong 2001: 348; Ren 2002a: 143; Ren and Shen 

2002: 384; Ren and Yang 2004a: 312; Ren and Ming 

2005: 382; Yang and Ren 2005a: 186, 2005b: 227; Ren and 

Hua 2006: 267; [China: Gansu (Kansu), Guizhou (Kweichow), 

Henan (Honana), Shaanxi (Shensi), Zhejiang 

(Chekiang)]. 

costatum costatum (Brullé, 1832). – Abdurakhmanov 1983: 

83 (Gonocephalum costatum); Medvedev and Abdurakhmanov 

1988: 89 (Gonocephalum costatum); Abdurakhmanov 

and Medvedev 1994: 58, 158 (Gonocephalum 

costatum); Abdurakhmanov et al. 1995: 201 (Gonocephalum 

costatum); Tezcan et al. 2000: 245–247; Abdurakhmanov 

and Abdulmuslimova 2002: 44 (Gonocephalum 

costatum); Tezcan et al. 2004: 167 (Gonocephalum 

costatum). 

costatum rugulosum (Küster, 1849). – Medvedev and Abdurakhmanov 

1988: 89 (Gonocephalum rugulosum); Abdurakhmanov 

and Medvedev 1994: 58, 158 (Gonocephalum 

rugulosum); Abdurakhmanov et al. 1995: 200 (Gonocephalum 

rugulosum); Abdurakhmanov and Abdulmuslimova 

2002: 44 (Gonocephalum rugulosum). 

curvicolle Reitter, 1889. – Ren and Ming 2005: 382; Yang 

and Ren 2005a: 186; [China: Gansu (Kansu), Shaanxi 

(Shensi)]. 

deliense Kaszab, 1952. – Ren and Yang 2004b: 306; Yang and 

Ren 2005a: 187. 

depressum (Fabricius, 1801). – Ren 2002b: 390. 

dorsogranosum (Fairmaire, 1896). – Medvedev and Abdurakhmanov 

1988: 90 (Gonocephalum dorsagralosum, 

misprint); Abdurakhmanov and Medvedev 1994: 58 (Gonocephalum 

dorsagranosum, misprint). 

ermischi Kaszab, 1960. – Medvedev and Abdurakhmanov 


formosanum Gebien, 1913. – Yang and Ren 2005a: 187. 

freyi Kaszab, 1952. – This was proposed as nomen novum 

for G. spinicolle Chatanay, 1917, not Fairmaire, 1896, so 

type designation was not necessary. 

freudei Kaszab, 1960. – Medvedev and Abdurakhmanov 1988: 

90; Abdurakhmanov and Medvedev 1994: 58. 

geneirotundum Ren, 1998. – Ren 2001: 89; Yang and Ren 

2005a: 187. 

germari germari Ferrer, 2000. – Drinkwater 1999: 183 

(G. germari, nomen nudum). 

gracile (Bates, 1879). – Yang and Ren 2005a: 187. 

granulatum nigrum (Küster, 1849). – Schneider and Leder, 

1877: 225 (cited for many records from Transcaucasus, 

probably G. granulatum pusillum, M. Nabozhenko pers. 

com.); Koenig, 1899: 379 (Gonocephalum pusillum var. 

nigrum, probably G. granulatum pussilum, M. 

Nabozhenko pers. com.) 

granulatum pusillum (Fabricius, 1791). – Schneider and 

Leder 1877: 225 (Gonocephalum viennense); Seidlitz 

1891b: 551 (Gonocephalum pusillum); Kulikovsky 1897: 

183 (Gonocephalum prolixum Er.); Kulikovsky 1897: 

183 (Gonocephalum pusillum); Yakobson 1927: 266 

(Dasus pusillus); Ogloblin and Kolobova 1927: 38; Yakobson 

1931: 209 (Dasus pusillus); Bogachev 1938: 140 

(Gonocephalum pusillum); Gilyarov 1964: 481 (Gonocephalum 

pusillum, larva); Dobrovolskiy 1951: 57 

(Gonocephalum pusillum); S. Medvedev and Shapiro 

1957: 196 (Gonocephalum pusillum); Kühnelt 1957: 

70 (Gonocephalum pusillum); Dobrovolskiy 1959: 

105 (Gonocephalum pusillum); Kryzhanovsky 1965: 

176 (Gonocephalum pusillum); Medvedev 1965: 370

(Gonocephalum pusillum); Kelejnikova 1968: 228 

(Gonocephalum pusillum); Protsenko and Kaltaev 1974 

(Gonocephalum pusillum, cited on many pages); 

Nepesova 1980: 29 (Gonocephalum pusillum); Abdurakhmanov 

1983: 83 (Gonocephalum pusillum); Medvedev 

and Abdurakhmanov 1988: 89 (Gonocephalum 

pusillum); Abdurakhmanov and Medvedev 1994: 58, 157 

(Gonocephalum pusillum); Abdurakhmanov et al. 1995: 

200 (Gonocephalum pusillum); Ovchinnikov 1996: 155 

(Gonocephalum pusillum); Nabozhenko 1999: 43 

(Gonocephalum pusillum); Jambazishvili 2000: 187 

(Gonocephalum pusillum); Tezcan et al. 2000: 245–247; 

Abdurakhmanov and Abdulmuslimova 2002: 44 (Gonocephalum 

pusillum); Tezcan et al. 2004: 168; Mordkovich 

2002: 60, 61 (Gonocephalum pusillum); Medvedev 

2004: 167 (Gonocephalum pusillum); Abdurakhmanov 

et al. 2005: 49 (Gonocephalum pusillum); 

Cherney 2005: 180, 182 (Gonocephalum pusillum); Yang 

and Ren 2005a: 186 (Gonocephalum pusillum); Cherney 

and Fedorenko 2006: 56, 144 (Gonocephalum pusillum). 

gridellii Koch, 1953. – Drinkwater 1999: 183. 

griseovittatum griseovittatum Gridelli, 1940. – Medvedev 

and Abdurakhmanov 1988: 90 (Gonocephalum griseovittatum); 

Abdurakhmanov and Medvedev 1994: 58 

(Gonocephalum griseovittatum); [Saudi Arabia]. 

guerryi Chatanay, 1917. – Medvedev and Abdurakhmanov 

1988: 90; Abdurakhmanov and Medvedev 1994: 58; Yang 

and Ren 2005a: 187. 

guignoti Kaszab, 1960. – Medvedev and Abdurakhmanov 


hadroide (Fairmaire, 1888). – Yang and Ren 2005a: 187 

(Gonocephalum reichardti). 

hauschildi Kaszab, 1952. – Ren and Yang 2004b: 306; Yang 

and Ren 2005a: 187. 

hintoni Kaszab, 1952. – Yang and Ren 2005a: 187. 

hispidulum Kaszab, 1952. – Ren 2002a: 142; Yang and Ren 

2005a: 186. 

inquinatum inquinatum (Sahlberg, 1923). – Medvedev and 

Abdurakhmanov 1988: 90 (Gonocephalum inquinatum); 

Abdurakhmanov and Medvedev 1994: 58 (Gonocephalum 

inquinatum). 

inquinatum parcepilosum Gridelli, 1940. – Medvedev and 

Abdurakhmanov 1988: 90 (Gonocephalum parcepilosum); 


parcepilosum). 

japanum Motschulsky, 1860. – Yang and Ren 2005a: 187. 

joliveti Ardoin, 1965. – Medvedev and Abdurakhmanov 1988: 


kandaharicum Kaszab, 1960. – Medvedev and Abdurakhmanov 


58. 

klapperichi Kaszab, 1952. – Ren 2002a: 139; Yang and Ren 

2005a: 187. 

kochi Kaszab, 1952. – Ren 2002a: 140; Ren and Yang 2004a: 

311; Yang and Ren 2005a: 186, 2005b: 227; Ren and Hua 

2006: 267; [China: Guizhou (Kweichow)]. 

konoi Kaszab, 1952. – Yang and Ren 2005a: 186. 

CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. 2 PART 261 

kuluanum Kaszab, 1952. – Medvedev and Abdurakhmanov 


kuntzeni Kaszab, 1952. – Ren 2002b: 390; Yang and Ren 

2005a: 186. 

lefranci (Fairmaire, 1863). – Medvedev and Abdurakhmanov 

1988: 89 (Gonocephalum lefrancei, misprint); Abdurakhmanov 

and Medvedev 1994: 58 (Gonocephalum 

lefrancei, misprint). 

lewisi Blair, 1922. – Kulzer 1954: 20. 

malayanum Gebien, 1935. – Ren 2002b: 390; Yang and Ren 

2005a: 187. 

massauense Gridelli, 1945. – Medvedev and Abdurakhmanov 


miyakense Nakane, 1963. – Ren 2001: 89; [China: Zhejiang]. 

moluccanum (Blanchard, 1853). – Kulzer 1954: 20; Ren 

2002a: 142, 2002b: 390; Yang and Ren 2005a: 186. 

muelleri muelleri Gridelli, 1945. – Medvedev and Abdurakhmanov 

1988: 89 (Gonocephalum muelleri); Abdurakhmanov 


muelleri). 

niloticum Ardoin, 1965. – Medvedev and Abdurakhmanov 


outreyi Chatanay, 1917. – Yang and Ren 2005a: 186. 

patrizii patrizii Gridelli, 1948. – Medvedev and Abdurakhmanov 

1988: 89 (Gonocephalum patrizii); Abdurakhmanov 


patrizii). 

patruele patruele (Erichson, 1843). – Kühnelt 1957: 70 (Gonocephalum 

rusticum patruele); Medvedev and Abdurakhmanov 


58 (Gonocephalum patruele). 

patruele turchestanicum Gridelli, 1948. – Kryzhanovsky 

1965: 176 (Gonocephalum turchestanicum); Nepesova 

1980: 29 (Gonocephalum turchestanicum); Medvedev 

and Abdurakhmanov 1988: 90 (Gonocephalum turchestanicum); 

Abdurakhmanov and Medvedev 1994: 58 

(Gonocephalum turchestanicum); Ren, Dai and Yu 

2000: 3 (Gonocephalum turchestanicum); Yang and 

Ren 2005a: 187 (Gonocephalum turchestanicum); 

[Kyrgyzstan, type locality: Issyk-Kul; China: Xinjiang 

(Sinkiang)]. 

perplexum (Lucas, 1849). – Medvedev and Abdurakhmanov 


persimile (Lewis, 1894). – Medvedev 1992: 638; Yang and Ren 

2005a: 186. 

platipenne Kaszab, 1960. – Medvedev and Abdurakhmanov 


prolixum prolixum (Erichson, 1843). – Medvedev and Abdurakhmanov 

1988: 89 (Gonocephalum prolixum); Abdurakhmanov 

and Medvedev 1994: 58 (Gonocephalum prolixum). 

pseudopubens Kaszab, 1952. – Ren 2002b: 390; Yang and Ren 

2005a: 186. 

pubens (Marseul, 1876). – Yang and Ren 2005a: 186. 

pubiferum Reitter, 1904. – Kühnelt 1957: 70 (Gonocephalum 

setulosum v. pubiferum); Kryzhanovsky 1965: 176;


Medvedev, 1965: 370; Davletshina, 1967: 530 (Gonocephalum 

pubeferum, misprint); Pirnazarov 1970: 25; 

Medvedev and Kaszab 1973: 93 (not Medvedev 1973: 93); 

Nepesova 1980: 29; Abdurakhmanov 1983: 83; Medvedev 


Medvedev 1994: 58, 157; Ovchinnikov 1996: 155; Ren, Dai 

and Yu 2000: 3; Yang and Ren 2005a: 186; [China:Xinjiang 

(Sinkiang)]. 

pygmaeum (Steven, 1829). – Kulikovsky 1897: 183 (Gonocephalum 

pygmaeum var. pedestre); Bogdanov-Katjkov 

1914: 474 (Gonocephalum setulosum); Yakobson 1927: 

266 (Dasus pygmaeus); Yakobson 1931: 209 (Dasus 

pygmaeus); Medvedev 1965: 370; Medvedev and Abdurakhmanov 


58; Cherney 2005: 179, 180; Ren and Yang 2004b: 306; Yang 

and Ren 2005a: 186; Cherney and Fedorenko 2006: 56, 144; 

[China: Xinjiang (Sinkiang)]. 

quadrinodosum Reitter, 1904. – Ren 2002b: 390; [China: 

Fujian (Fukien)]. 

recticolle Motschulsky, 1866. – Medvedev 1992: 638; Yang and 

Ren 2005a: 186. 

remedellii Gridelli, 1945. – Medvedev and Abdurakhmanov 


reticulatum Motschulsky, 1852. – Medvedev and Kaszab, 

1973: 94 (not Gonocephalum mongolicum: Medvedev 

1973: 94); Medvedev 1992: 637; Ren 2001: 90; Ren and Shen 

2002: 383; Yang and Ren 2005a: 186. 

royi royi Ardoin, 1961. – Medvedev and Abdurakhmanov 

1988: 89; Abdurakhmanov and Medvedev 1994: 58 (Gonocephalum 

royi); [Sudan]. 

rusticum (Olivier, 1811) – Ménétriés 1832: 203 (Opatrum 

fuscum); Schneider and Leder 1877: 225; Koenig 1899: 

379; Bogdanov-Katjkov 1914: 474; Bogush 1936: 36; 

Bogachev 1938: 140; Kryzhanovsky 1965: 176; Medvedev 

1965: 370; Davletshina 1967: 530; Kelejnikova 1968: 228; 

Skopin 1964: 280, 1968: 73; Pirnazarov 1970: 25; Tajibaev 

1972: 275; Medvedev and Kaszab 1973: 94; Protsenko 

and Kaltaev 1974 (cited on many pages); Nepesova 1980: 

29; Abdurakhmanov 1983: 83; Medvedev and Abdurakhmanov 

1988: 89; Medvedev and Nepesova 1990: 886; 

Abdurakhmanov and Medvedev 1994: 58, 158; Abdurakhmanov 

et al. 1995: 201; Ovchinnikov 1996: 155; 

Nabozhenko 1999: 42; Keskin 1999: 217; Jambazishvili 

2000: 187; Abdurakhmanov and Abdulmuslimova 

2002: 44; Tezcan et al. 2004: 168–169; Cherney 2005: 180, 

187; Yang and Ren 2005a: 186; Cherney and Fedorenko 

2006: 56. 

sauteri Kaszab, 1952. – Yang and Ren 2005a: 186. 

schneideri Reitter, 1892. – Skopin 1964: 280; Kryzhanovsky 

1965: 176; Pirnazarov 1970: 25; Nepesova 1980: 29; Abdurakhmanov 

1983: 83; Medvedev and Abdurakhmanov 1988: 

90; Abdurakhmanov and Medvedev 1994: 58, 158; Abdurakhmanov 

et al. 1995: 201; Abdurakhmanov and Abdulmuslimova 

2002: 44; Yang and Ren 2005a: 186; [Azerbaijan 

(Apsheron peninsula)]. 

semipatruele Kaszab, 1952. – Kulzer 1956:652; Medvedev 


Medvedev 1994: 58. 

setulosum setulosum (Faldermann, 1837). – Koenig 1899: 

379; Kühnelt 1957: 71; Skopin 1964: 280; Davletshina 1967: 

530; Pirnazarov 1970: 25; Medvedev et al. 1975: 90; 

Nepesova 1980: 29; Abdurakhmanov 1983: 83; Medvedev 


Medvedev 1994: 58, 157; Abdurakhmanov et al. 1995: 200; 

Yu and Ren 1994: 91; Ovchinnikov 1996: 155; Nabozhenko 

1999: 42; Jambazishvili, 2000: 187; Ren et al. 2000: 3; 

Abdurakhmanov and Abdulmuslimova 2002: 44; Tezcan et 

al. 2004: 169; [Azerbaijan, Georgia]. 

sexuale (Marseul, 1876). – Medvedev 1992: 638; Ren 1998b: 

91; Yang and Ren 2005a: 186. 

simplex simplex (Fabricius, 1801). – Medvedev and Abdurakhmanov 

1988: 90 (Gonocephalum simplex); Abdurakhmanov 

and Medvedev 1994: 58 (Gonocephalum simplex); 

Drinkwater 1999: 183 (Gonocephalum simplex). 

simulatrix (Fairmaire, 1891). – Medvedev and Abdurakhmanov 


58; Yang and Ren 2005a: 186. 

sinicum Reichardt, 1936. – Yang and Ren 2005a: 187. 

soricinum soricinum (Reiche et Saulcy, 1857). – Medvedev 

and Abdurakhmanov 1988: 89 (Gonocephalum soricinum); 


soricinum). 

strigosum (Reiche, 1850). – Type designation by Ferrer 

(1995a: 33): lectotype, Benin (Dahomey). 

subrugulosum Reitter, 1887. – Medvedev and Kaszab, 1973: 

94 (not Medvedev 1973: 94); Ren et al. 2000: 3; Yang and 

Ren 2005a: 186. 

subspinosum (Fairmaire, 1894). – Ren 2002a: 139; Ren and 

Yang 2004a: 311; Ren and Ming 2005: 382; Yang and Ren 

2005a: 187, 2005b: 227; Ren and Hua 2006: 267; Ren and 

Yang 2007: 255; [China: Gansu (Kansu), Guizhou (Kweichow), 

Shaanxi (Shensi)]. 

tellinii Gridelli, 1945. – Medvedev and Abdurakhmanov 1988: 


terminale Reichardt, 1936. – Medvedev 1992: 638. 

tibetanum Kaszab, 1952. – Yang and Ren 2005a: 187. 

tibiale Grimm, 2011: 254. Distribution: ORIENTAL REGION: 

Indonesia (Borneo). 

titschacki Kaszab, 1952. – Ren 2002a: 139; Yang and Ren 

2005a: 187. 

tomentosum (Walker, 1871). – Medvedev and Abdurakhmanov 


58. 

tschilianum Kaszab, 1952. – Yang and Ren 2005a: 187. 

tuberculatum (Hope, 1831). – Medvedev and Abdurakhmanov 


58; Ren 1998b: 91, 2002a: 138; Yang and Ren 2005a: 186; 

Ren and Hua 2006: 267. 

tuberipenne Grimm, 2011: 251. Distribution: ORIENTAL 

REGION: Indonesia (Borneo). 

vagum (Steven, 1829). – Kulzer 1954: 20; Yang and Ren 

2005a: 186. 

wui Ren, 1995. – Ren and Saihong 2001: 347; Ren 2001: 90; 

Yang and Ren 2005a: 187.

zoltani Iwan, Ferrer et Raś 2010. – Medvedev and Abdurakhmanov, 

1988: 90 (Gonocephalum paralellum, 

misprint, preoccupied name); Abdurakhmanov and Medvedev, 

1994: 58 (Gonocephalum paralellum, misprint, 

preoccupied name). 

Melanesthes medvedevi Kaszab, 1973, in: Medvedev and 

Kaszab (1973: 103) [replacement name for Melanesthes 

maxima Kaszab, 1965c: 341]. – Misinterpreted by Iwan et 

al. (2010: 304) as Gonocephalum medvedevi Kaszab, 

1965. 

Notes on the revision of European and African 

Gonocephalum species by Ferrer (1993, 1995, 

2000) 

A revision of the African and Europaean species of 

Gonocephalum was done by Ferrer and published in 

three separated papers (Ferrer 1993, 1995, 2000). 

The first one (Ferrer 1993) contains the key to all 

species, including propositions of new taxa and new 

names, but the descriptions of the new species were 

presented in both subsequent papers (Ferrer 1995, 

2000). According to the rule of International Code of 

Zoological Nomenclatures (ICZN 1999, Article 13), all 

the names used for the new species by Ferrer in 1993, 

should be treated as nomina nuda, and consequently 

are unavailable. The same names used in the later 

papers (Ferrer 1995, 2000) are available, and take 

authorship and date from these acts of establishment 

(ICZN 1999, Articles 21, 50). There are the list of the 

names: 

G. albertii Ferrer, 1995 (1993a: 77) 

G. alticola exile Ferrer, 2000 (1993a: 104) 

G. alticola kinschassae Ferrer, 2000 (1993a: 104) 

G. alticola nossibense Ferrer, 2000 (1993a: 104) 

G. alticola seychellense Ferrer, 2000 (1993a: 104) 

G. alticola zulu Ferrer, 2000 (1993a: 104) 

G. angelicae Ferrer, 1995 (1993a: 81) 

G. angolense escalerai Ferrer, 2000 (1993a: 99) 

G. bartolozzii Ferrer, 1995 (1993a: 99) 

G. batesi Ferrer, 2000 (1993a: 96) 

G. bremeri Ferrer, 1995 (1993a: 90) 

G. brincki Ferrer, 2000 (1993a: 105) 

G. compressum Ferrer, 1995 (1993a: 76) 

G. confusum Ferrer, 2000 (1993a: 106) 

G. demeteri Ferrer, 1995 (1993a:83) 

G. dermestoides latum Ferrer, 1995 (1993a: 91) 

G. deserticola Ferrer, 2000 (1993a: 105) 

G. devylderi Ferrer, 1995 (1993a: 93) 

G. folini Ferrer, 1995 (1993a: 81) 

G. frederici Ferrer, 1995 (1993a: 73) 

G. germari germari Ferrer, 2000 (1993a: 95) 

G. germari hemmingi Ferrer, 2000 (1993a: 95) 

G. gerstaeckeri Ferrer, 1995 (1993a: 79) 


G. girardi Ferrer, 1995 (1993a: 89) 

G. giraudini rex Ferrer, 1995 (1993a: 89) 

G. grimmi Ferrer, 1995 (1993a: 89) 

G. hiekei Ferrer, 1995 (1993a: 75) 

G. imitator Ferrer, 1995 (1993a: 90) 

G. inaequale danielssoni Ferrer, 1995 (1993a: 73) 

G. indictum indictum Ferrer, 1995 (1993a: 79) 

G. inquinatum palisoti Ferrer, 2000 (1993a: 98) 

G. inquinatum suberinitum Ferrer, 2000 (1993a: 98) 

G. juanense Ferrer, 2000 (1993a: 100) 

G. kaszabi Ferrer, 1995 (1993a: 84) 

G. kempsteri Ferrer, 2000 (1993a: 78) 

G. lesnei Ferrer, 1995 (1993a: 78) 

G. martini Ferrer, 1995 (1993a: 87) 

G. merensi estevezi Ferrer, 2000 (1993a: 97) 

G. merkli Ferrer, 2000 (1993a: 95) 

G. obscurum aethiops Ferrer, 1995 (1993a: 24) 

G. patrizii lagosicum Ferrer, 1995 (1993a: 91) 

G. politum Ferrer, 2000 (1993a: 95) 

G. royi clarkei Ferrer, 1995 (1993a: 86) 

G. simplex aldabraense Ferrer, 2000 (1993a: 103) 

G. simplex andreinii Ferrer, 2000 (1993a: 103) 

G. simplex sudanicum Ferrer, 2000 (1993a: 102) 

G. simplex ubanguiense Ferrer, 2000 (1993a: 102) 

G. simplex yemenicum Ferrer, 2000 (1993a: 103) 

G. soricinum faraonicum Ferrer, 2000 (1993a: 97) 

G. subsetosum erichsoni Ferrer, 2000 (1993a: 99) 

G. swazilandense Ferrer, 1995 (1993a: 80) 

G. triste Ferrer, 2000 (1993a: 105) 

G. tuberculiferum Ferrer, 1995 (1993a: 81, 85) 

G. validum Ferrer, 2000 (1993a: 98) 

G. zambesense Ferrer, 1995 (1993a: 94) 

G. pilosa Ferrer, 2000 (1993a: 107) 

Types deposited in the Hungarian Natural 

History Museum, Budapest, Hungary (HNHM) 

Unclear designation of types by Kaszab (1952), 

mentioned in the first part of the catalogue (Iwan et al. 

2010), leads to several questions. One of many examples 

is the species Gonocephalum abnormale Kaszab, 

1952. Kaszab (1952: 635) in the “Untersuchungsmaterial” 

listed the number of specimens and their place 

of deposition: “7 Exemplare (4 Exempl. British Mus., 2 

Exempl. Mus. Budapest; 1 Exempl. Mus. Frey)”, and 

then explicitly appointed one specimen as type deposited 

in HNHM: “Typus befindet sich in der Sammlung des 

Ungarischen Naturwissenschaftlichen Museum in 

Budapest“. Are the specimens deposited at the Natural 

History Museum (UK, London) and NHMB types? Do 

the specimens have labels with the type designations 

written by Kaszab? Currently, there are 3 specimens 

(labelled as HT and 2 PT) of G. abnormale Kaszab, 

1952 deposited in HNHM.


List of type specimens preserved in HNHM 

G. (Opatropis) aalbui Ferrer, 2000 – 11 PT 

G. abnormale Kaszab, 1952 – HT, 2 PT 

G. acuticolle Kaszab, 1952 – 3 PT 

G. adpressiforme Kaszab, 1951 – 8 PT 

G. andamanense Kaszab, 1952 – 1 PT 

G. andrewesi Kaszab, 1952 – 27 PT 

G. angolense escalerai Ferrer, 2000 – HT, 2 PT 

G. arisi Reitter, 1905 – HT 

G. balmeae Kaszab, 1952 – 1 PT 

G. baloghi Kaszab, 1969 – HT 

G. belli Kaszab, 1952 – 1 PT 

G. bengalense Kaszab, 1952 – 2 PT 

G. besnardi Kaszab, 1982 – HT, 6 PT 

G. bigranulatum Kaszab, 1952 – HT 

G. bimaculatum Ferrer, 1995 – HT, 4 PT 

G. birmanicum Kaszab, 1952 – HT 

G. biroi Kaszab, 1952 – HT, AT, 2 PT 

G. biseriatum Kaszab, 1975 – HT 

G. blairi Kaszab, 1952 – 4 PT 

G. borosi Kaszab, 1952 – 1 HT 

G. brachelytra Kaszab, 1952 – 5 PT 

G. brazzavillae Kaszab, 1969 – HT, 1 PT 

G. bremeri Ferrer, 1995 – HT, AT, 1 PT 

G. brevisetosum Kaszab, 1952 – MT = HT 

G. brittoni Kaszab, 1952 – 1 PT 

G. buitenzorgense Kaszab, 1952 – 1 ST 

G. celebense Kaszab, 1952 – 2 PT 

G. civicum Kaszab, 1952 – 14 PT 

G. clavigerum Kaszab, 1980 – HT 

G. coenosum Kaszab, 1952 – HT, 55 PT 

G. confusum Ferrer, 2000 – 29 PT 

G. cookae Kaszab, 1952 – 1 PT 

G. corallinum Kaszab, 1961 – 2 PT 

G. coriaceum takara Nakane, 1963 (= Gonocephalum 

takara) – 3 PT 

G. crassepunctatum Kaszab, 1952 – HT, 3 PT 

G. curiosum Kaszab, 1952 – 7 PT 

G. curvicolle Reitter, 1889 – 3 ST 

G. csikii Kaszab, 1952 – HT 

G. dasiforme Kaszab, 1952 – 4 PT 

G. deliensis Kaszab, 1952 – HT 

G. demeteri Ferrer, 1995 – HT 

G. dentipes Kaszab, 1952 – 11 PT 

G. dermestoides latum Ferrer, 1995 – HT, 5 PT (according 

to the description these types are in the 

coll. Bremer, but in fact in the HNHM!) 

G. deserticola Ferrer, 2000 – 9 PT 

G. elytrale Kaszab, 1952 – 1 PT 

G. endroedii Kaszab, 1952 – MT = HT 

G. ermischi Kaszab, 1960 – 6 PT 

G. espagnoli Kaszab, 1965 – HT 

G. feae Gebien, 1920 – 1 cotype 

G. foveicolle Kaszab, 1952 – HT, 2 PT 

G. freudei Kaszab, 1960 – 3 PT 

G. gebienianum Kaszab, 1952 – 5 PT 

G. germari Ferrer, 2000 – HT, 11 PT 

G. gestroi Gebien, 1920 – 1 cotype 

G. girardi Ferrer, 1995 – 1 PT 

G. granulatipenne Kaszab, 1978 – 4 PT 

G. granulipenne Gridelli, 1945 – 1 PT 

G. greensladei Kaszab, 1980 – HT, 11 PT 

G. gridellianum Kaszab, 1952 – 1 PT 

G. grimmi Ferrer, 1995 – HT, 1 PT 

G. griseovittatum Gridelli, 1940 – 1 PT 

G. guignoti Kaszab, 1960 – 5 PT 

G. hauschildi Kaszab, 1952 – HT, 3 PT 

G. helaeoides Kaszab, 1952 – 4 PT 

G. helferi Kaszab, 1952 – 4 PT 

G. himalayense Kaszab, 1952 – HT 

G. hingstoni Kaszab, 1952 – HT, 3 PT 

G. hintoni Kaszab, 1952 – 2 PT 

G. hispidulum Kaszab, 1952 – 4 PT 

G. horni Kaszab, 1952 – 26 PT 

G. impressiceps Kaszab, 1952 – HT 

G. inaequale Gridelli, 1945 – 1 PT 

G. indicum Kaszab, 1952 – HT 

G. indochinense Kaszab, 1972 – 1 PT 

G. javanicum Kaszab, 1952 – HT, 3 PT 

G. kandaharicum Kaszab, 1960 – 5 PT 

G. karakorumense Kaszab, 1961 – HT 

G. kaszabi Ferrer, 1995 – HT, AT 

G. kempsteri Ferrer, 2000 – 1 PT 

G. klapperichi Kaszab, 1952 – HT, 2 PT 

G. kochi Kaszab, 1952 – 6 PT 

G. kondoi Masumoto, 1983 – 2 PT 

G. koreanum Kaszab, 1952 – HT 

G. kuehnelti Kaszab, 1961 – HT, 41 PT 

G. kuluanum Kaszab, 1952 – 3 PT 

G. kulzeri Kaszab, 1952 – 3 PT 

G. kuntzeni Kaszab, 1952 – 3 PT 

G. laosense Kaszab, 1952 – HT, 1 PT 

G. longitarse Kaszab, 1952 – 2 PT 

G. longulum Gebien, 1911 – 1 cotype 

G. macrophthalmum Kaszab, 1952 – 1 PT 

G. madurense Kaszab, 1952 – HT, 4 PT 

G. malayanum Gebien, 1935 – 1 cotype 

G. marani Kaszab, 1952 – 1 PT 

G. martensi Kaszab, 1977 (=Gonocephalum simulatrix 

Fairmaire, 1891) – 5 PT 

G. merkli Ferrer, 2000 – HT 

G. microphthalmum Ardoin, 1965 – 2 PT 

G. mongolicum Reitter, 1889 (= Gonocephalum reticulatum 

Motschulsky, 1854) – 2 ST 

G. mysorense Kaszab, 1952 – 1 HT 

G. nepalicum Kaszab, 1973 – 28 PT 

G. niloticum Ardoin, 1965 – 2 PT 

G. novaecaledoniae Kaszab, 1982 – 3 PT 

G. nyassae Ferrer, 1995 – HT 

G. obenbergeri Kaszab, 1952 – 2 PT 

G. obscurum aethiops Ferrer, 1995 – HT, 3 PT

G. oceanicum Kaszab, 1985 – HT 

G. oculare Kaszab, 1952 – HT, 3 PT (= Gonocephalum 

(Opatropis) oculare) 

G. okinawanum Chûjô, 1963 – 2 PT 

G. papuanum Gebien, 1920 – 1 cotype 

G. parallelum Kaszab, 1952 – HT, 1 PT (= Gonocephalum 

zoltani Iwan, Ferrer et Ra?, 2010) 

G. parcesetosum Kaszab, 1952 – 1 PT 

G. patricium Kaszab, 1952 – 3 PT 

G. patrizii lagosicum Ferrer, 1995 – HT, 1 PT 

G. peguanum Kaszab, 1952 – 1 PT 

G. planicolle Kaszab, 1952 – HT 

G. platipenne Kaszab, 1960 – 1 PT 

G. politum Ferrer, 2000 – 1 PT 

G. potanini Reichardt, 1936 (= G. guerryi Chatanay, 

1917) – 2 ST 

G. pottsi Kulzer, 1957 – 1 PT 

G. pseudopubens Kaszab, 1952 – HT, 9 PT 

G. puberulum Kaszab, 1952 – 2 PT 

G. pusillum afghanicum Gridelli, 1954 (= Gonocephalum 

granulatum afghanicum Gridelli, 

1954) – 1 PT 

G. recticolle var. terminale Reichardt, 1936 (= Gonocephalum 

terminale Reichardt, 1936) – 5 ST 

G. remedellii Gridelli, 1945 – 1 ST 

G. rileyi Kaszab, 1952 – 1 PT 

G. rondoni Kaszab, 1972 – 3 PT 

G. roseni Kaszab, 1952 – HT, 9 PT 

G. rostratum Bremer et Ferrer, 2002 – 2PT 

G. rotundatum Gridelli, 1940 – 1 PT 

G. royi clarkei Ferrer, 1995 – HT, 3 PT 

G. sauteri Kaszab, 1952 – HT 

G. schneideri Reitter, 1898 – HT 

G. schusteri Kaszab, 1952 – HT, 2 PT 

G. semipatruele Kaszab, 1952 – 5 PT 

G. setulosum var. pubiferum Reitter, 1904 (= Gonocephalum 

pubiferum Reitter, 1904) – 9 ST 

G. shimoganum Kaszab, 1952 – 1 PT 

G. sikkimense Kaszab, 1952 – 3 PT 

G. simplex montisrolandi Ardoin, 1962 – 1 PT 

G. simplex ubanguiense Ferrer, 2000 – 1 PT 

G. sinicum Reichardt, 1936 – 1 ST 

G. somalicum Gridelli, 1945 (= Gonocephalum alternicostis 

somalicum) – 1 ST 

G. spangleri Kaszab, 1980 – 2 PT 

G. stoeckleini Kaszab, 1952 – HT, 40 PT 

G. subcontractum Gridelli, 1945 – 1 PT 

G. subrugulosum Reitter, 1887 – 1 ST 

G. subsetosum Reitter, 1904 (= Gonocephalum subpinosum 

(Fairmaire, 1894)) – 2 ST 

G. szekessyi Kaszab, 1952 – 1 PT 

G. tellinii Gridelli, 1945 – 5 PT 

G. tenasserimicum Kaszab, 1952 – HT, 9 PT 

G. tenuicorne Kaszab, 1952 – HT, 6 PT 

G. tenuipes Kaszab, 1952 – HT 

G. tibetanum Kaszab, 1952 – 1 PT 


G. titschacki Kaszab, 1952 – HT 

G. tonkinense Kaszab, 1952 – 3 PT 

G. topali Kaszab, 1975 – HT, 13 PT 

G. triste Ferrer, 2000 – 2 PT 

G. tschilianum Kaszab, 1952 – HT 

G. tuberculiferum Ferrer, 1995 – HT 

G. ulugurense Ardoin, 1976 – 2 PT 

G. uniseriatum Kaszab, 1952 – HT 

G. validum Ferrer, 2000 – 25 PT 

G. veigai Koch, 1956 – 8 PT 

G. vientianeum Kaszab, 1972 – 2 PT 

G. vincenti Ardoin, 1965 – 6 PT 

G. wau Kaszab, 1970 – HT, AT, 29 PT 

G. wittmeri Kaszab, 1975 – 22 PT 

G. woynarovichi Kaszab, 1973 – HT, 12 PT 

Opatrum scutellare Perty, 1831 (= Gonocephalum 

scutellare) – 1 ST 

Leichenum impictum Fairmaire, 1849 (= Gonocephalum 

impictum) – 1 ST 

Hasticollinum podagrarium Kaszab, 1939 (= Gonocephalum 

podagrarium) – 2 PT 

Hopatrum dorsogranosum Fairmaire, 1896 (= Gonocephalum 

dorsogranosum) – 1 ST 

Pseudoblaps simulatrix Fairmaire, 1891 (= Gonocephalum 

simulatrix) – 3 ST. 

ACKNOWLEDGEMENT 

We are grateful to Haoyu Liu (China, Hebei) 

and Maxim Nabozhenko (Russia, Rostov-on-Don) 

for the provide the copies of Chinese and Russian 

papers. 

REFERENCES 

Abdurakhmanov, G. M. 1983. On faunistic links of Coleoptera 

of arid regions of the Eastern part of the Caucasus Major 

and Middle Asia. Entomologicheskoe obozrenie, 62(3): 

481–497. (In Russian). 

Abdurakhmanov, G. M. and G. S. Medvedev. 1994. Catalogue of 

darkling beetles of Caucasus. Makhachkala, Daghestan 

State Pedagogical Institute, 212 pp. (In Russian). 

Abdurakhmanov, G. M., Ismailov, Sh. I. and A. L. Lobanov. 

1995. A new approach to the problem of objective zoogeographical 

districting. Part 5. Computer base of dates 

TENKAV about fauna of darkling beetles of Caucasus. 

Makhachkala, Dagestanian State Pedagogical Institute, 

325 pp. (In Russian). 

Abdurakhmanov, G. M. and K. M. Abdulmuslimova. 2002. Composition, 

morphoecological structure and zoogeographic 

peculiarities of tenebrionid populations in the Caucasus. 

Russian entomological journal, 11(1): 41–48. (In Russian). 

Abdurakhmanov, G. M., Tochieva, F. T. and T. Yu. Tochiev. 

2005. Darkling beetles of Dzheirakh and Targim hollows of


Respublic Ingushetiya. Makhachkala, Yupiter, 120 pp. (In 

Russian). 

Allard, E. 1883. Mélanges Entomologiques. Annales de la 

Société Entomologique de Belgique, 27: 5–49. 

Angelov, P. A. and G. S. Medvedev. 1981. Tenebrionid beetles 

(Coleoptera, Tenebrionidae) of Bulgaria. Entomologicheskoe 

Obozrenie, 60: 300–315. (In Russian). 

Ardoin, P. 1961. Le Parc national de Niokolo Koba, Coleoptera, 

Tenebrionidae. Mémoires de l’Institut Français d’Afrique 

Noire, 62: 140–149. 

Ardoin, P. 1962. Nouvelles formes africaines de Coléoptères 

Ténébrionidaes appartentant aux collections de l’Ifan. 

Bulletin de l’Institut Français d’Afrique Noire, Série A, 24: 

354–359. 

Ardoin, P. 1963. Essai de révision des Amarygmini africains 

(2 me , 3 me , 4 me , et 5 me parties). Bulletin de l’Institut 

Français d’Afrique Noire, Série A, 25: 9–116. 

Ardoin, P. 1965a. Contribution à la faune du Congo (Brazzaville) 

(Mission A. Villiers et A. Descarpentries). VIII. Coléoptères 

Ténébrionides. Bulletin de l’Institut Français 

d’Afrique Noire, Série A, 27: 963–1011. 

Ardoin, P. 1965b. Contribution à l’étude des Ténébrionides 

africains (Col.). Bulletin de la Société Entomologique de 

France, 70: 73–85, pl. II. 

Ardoin, P. 1965c. Note préliminaire sur les ténébrionides 

récoltés aux Monts Loma (Sierra-Leone). Bulletin 

de l’Institut Français d’Afrique Noire, Série A, 27: 

1315–1320. 

Ardoin, P. 1966. Contribution a la conaissance des Ténébrionides 

de l’Ile Europa (Col.) Mémoires du Muséum National 

d’Histoire Naturelle (nouvelle série), série A, Zoologie, 41: 

187–190, pl. II. 

Ardoin, P. 1967. Les Gonocephalum de la Région de l’Omo. 

Annales de la Société entomologique de France, N. S., 

3(1): 207–214. 

Ardoin, P. 1971. Le massif des Monts Loma (Sierra Leone). 

XII. Coleoptera Tenebrionidae. Mémoires de l’Institut 

Français de l’Afrique Noire, 86: 283–290. 

Ardoin, P. 1972. Liste des Ténébrionides récoltés au Soudan 

par les expéditions finlandeises (1962–1964), no 27, Commentationes 

Biologicae. Societas Scientiarum Fennica 

(Helsinki, Helsingfors), 49: 3–20. 

Ardoin, P. 1976. Mission Entomologique du Musée Royal de 

l’Afrique Centrale aux Monts Uluguru, Tanzanie. 20. 

Coleoptera Tenebrionidae. Revue Zoologique et Botanique 

Africaine, 90: 723–768, pls 7–10. 

Bach, M. 1856. Käferfauna für Nord- und Mitteldeutschland 

mit besonderer Rückshicht auf die preussischen. Coblenz, 

J. Bölscher, Rheinlande, Dritter Band, 292 pp. 

Baudi di Selve, F. 1875a. Coleotteri Tenebrioniti delle 

collezioni italiane. Bollettino della Società Entomologica 

Italiana, [1874] 6: 275–301. 

Baudi di Selve, F. 1875b. Catalogo dei Tenebrioniti della fauna 

europea e circummediterranea apparteneti alle 

collezioni del Museo Civico di Genova. Parte seconda. 

Annali del Museo Civico di Storia Naturale di Genova, 7: 

684–703. 

Baudi di Selve, F. 1876. Europaeae et circummediterraneae 

faunae Tenebrionidum specierum, quae Comes Dejean in 

suo Catalogo, editio 3 a consignavit, ex ejusdem collectione 

in R. Taurinensi Museo asservata, cum auctorum 

hodierne denominatione collatio. Pars altera. Deutsche 

Entomologische Zeitschrift, 20: 1–74. 

Bates, F. 1879. Characters of the new genera and species of 

Heteromera collected by Dr. Stoliczka during the Forsyth 

expedition to Kashgar in 1873–4. Cistula Entomologica, 2: 

467–484. 

Bates, F. 1890. Heteromera, pp. 55–79. In: Ball, V. (ed.): Scientific 

results of the second Yarkand Mission; based upon 

the collections and notes of the late Ferdinand Stoliczka, 

Ph. Dr. Calcutta: Office of superindendent of Goverment 

Printing, India, 79 pp, 2 pls. 

Besser, W. S. J. G. v. 1832. Additamenta et observatiunculae in 

Cel. Steven Tentyrias et Opatra collectionis Stevenianae 

nunc musei Universitatis Mosquensis. Nouvelle Mémoires 

de la Société Impériale des Naturalistes de Moscou, 2: 

5–21. 

Billberg, G. J. 1815. Insecta ex ordine Coleopteroum descripta. 

Nova Acta Upsalae, 72: 271–281, pl. 9. 

Blackburn, T. 1892. Coleoptera. Scientific results of the Elder 

Exploring Expedition. Transactions of the Royal Society 

of South Australia, 16: 16–61. 

Blackburn, T. 1894. Further notes on Australian Coleoptera, 

with descriptions of new genera and species. Part XVI. 

Transactions of the Royal Society of South Australia 18: 

200–240. 

Blackburn, T. 1896. Coleoptera (exclusive of the Carabidae), 

pp. 254–308. In: Spencer, B. (ed.), Report of the Horn Scientific 

Expedition to Central Australia. Part 2. Zoology. 

Dulau & Co., London. 

Blackburn, T. 1907. Further notes on Australian Coleoptera, 

with description of new genera and species, xxxvii. Transactions 

of the Royal Society of South Australia, 31: 

231–299. 

Blair, K. G. 1921. Types of Heteromera described by F. Walker 

now in the British Museum. Transactions of the Entomological 

Society of London, 1921: 268–283. 

Blair, K. G. 1923. New species of Heteromerous Coleoptera 

from Mesopotamia. The Entomologist’s Monthly Magazine, 

(3)9: 118–126. 

Blair, K. G. 1928. Heteromera, Bostrychoidea, Malacodermata 

and Buprestidae. Insects of Samoa, Part IV. Fasc. 2: 

67–109. 

Blanchard, E. 1853. Description des insectes. In: Hombros, J. 

& Jacquinot, H. Zoologie. Tome quatrième. In: Dumontd’Urville, 

J. Voyage au pôle sud et dans l’Océanie sur les 

corvettes l’Astrolabe et la Zélée; exécuté par ordre du Roi 

pendant les années 1837–1838–1839–1840, sous le commandement 

de M. J. Dumont d’Urville, capitaine du vaisseau, 

publié par ordre du gouvernement, sous la direction 

supérieure de M. Jacquinot, capitaine de vaisseau, commendant 

de la Zélée. Paris: Gide & J. Baudry, 422 pp., 

19 pls. 

Boddy, D. W. 1965. Family Tenebrionidae, pp. 130–189. In: 

Hatch, M. H. (ed.), The beetles of the Pacific northwest. 

Part IV: Macrodactyles, Palpicornes, and Heteromera. 

Seattle: University of Washington Press. 

Bogachev, A. V. 1938. List of species of beetles from famelies 

Tenebrionidae and Scarabaeidae (collected in Nakhichevan 

Autonomic SSR in 1933). Proceedings of Zoological 

Institute of Azerbaijanian branch of Academy of Sciences 

of USSR. 8: 135–154. (In Russian).

Bogdanov-Katjkov, N. N. 1914. To the fauna of darkling 

beetles of Kuban region (Coleoptera, Tenebrionidae). 

Russkoe entomologicheskoe obozrenie, 14: 473–474. (In 

Russian). 

Bogush, P. P. 1936. Darkling beetle Gonocephalum rusticum 

Ol., as pest of corn-shoots in Bairam-Ali region. 

Bulletin of Turkmenian zoological station, 1: 35–37. (In 

Russian). 

Boisduval, J. B. A. D. de. 1835. Voyage de découvertes de l’Astrolabe 

exécuté par l’ordre du Roi, pendant les années 

1826–1827–1828–1829, de M. Dumont d’Urville. Faune 

Entomologique de l’Océan pacifique, avec l’illustration des 

insectes nouveaux recueillis pandent le voyage. Deuxième 

partie. Coléoptères et autres ordres. Paris: J. Tastu, vii + 

716 pp. 

Bremer, H. J. and J. Ferrer. 1992. Zwei neue Gonocephalum- 

Arten aus der orientalischen Region (Col., Tenebrionidae, 

Opatrini). Entomologische Blaetter fuer Biologie und Systematik 

der Kaefer, 88: 83–88. 

Bremer, H. J. and J. Ferrer. 2002. Description of a new species 

of Gonocephalum Solier, 1834 from Burma (Coleoptera, 

Tenebrionidae, Opatrinae, Opatrini). Entomofauna, 23: 

221–226. 

Brisout de Barneville, C. 1863. [new taxa], pp. iv + 3–79 + 135 

pp. In: Grenier, A. (ed.), Catalogue des coléoptères de 

France et matériaux pour servir à la faune des coléoptères 

français, par MM. E. Allard, D r . Ch. Aubé, Ch. Brisout 

de Barneville, A. Chevrolat, L. Fairmiare, Al. Fauvel, Dr. A. 

Grenier, Dr. Kraatz, J. Linder, L. Reiche, et Félicien de 

Saulcy. Paris: L. Toinon. 

Brullé, A. 1832. Expedition scientifique de Morée. 3 e Série 

Zoologique. Partie des Insectes. Paris: Levrault, pp. 129, 

64–395, pls 22. 

Brullé, A. 1838. Insectes, pp. 54–95, pls 4, 5. In: Webb P. B. 

& Berthelot S. 1836–1844, Histoire naturelle des îles 

Canaries. Tome deuxième. Deuxième Partie. Contenant la 

Zoologie. Paris: Bèthune. 

Canzoneri, S. 1970. I Tenebrionidae delle Isole Egadi. Memorie 

del Museo Civico di Storia Naturale di Verona, 18: 

55–89. 

Canzoneri, S. 1977. Contributo alla conoscenza de Tenebrionidi 

appenninici. (XXXII Contributo allo studio dei Tenebrionidi). 

Bollettino del Museo Civico di Storia Naturale di 

Verona, 4: 227–285. 

Canzoneri, S. and P. Vienna. 1987. I Tenebrionidae della Padania 

(Coleoptera Heteromera). Bollettino del Museo Civico 

di Storia Naturale di Venezia, 36: 7–62. 

Carter, H. J. 1914. Notes on Tenebrionidae in the South Australian 

Museum, collected by Mr. A. M. Lea, 1911–12, with 

descriptions of new species. Transactions of the Royal 

Society of South Australia, 38: 219–238. 

Carter, H. J. 1915. The Australian Strongyliinae and other 

Tenebrionidae with descriptions of new genera and 

species. (Family Tenebrionidae). Proceedings of the Linnean 

Society of New South Wales, 40: 522–539. 

Carter, H. J. 1928. Some new Australian Heteromera in the 

Germ. Entomolog. Museum Dahlem (Col.). Entomologische 

Mitteilungen, 17: 132–138. 

Carter, H. J. 1930. New Guinea and Australia Coleoptera. 

Notes and new species. Proceedings of the Linnean Society 

of New South Wales, 55: 532–549. 


Casey, T. L. 1890. Coleopterological notices, II. Annals of the 

New York Academy of Sciences, 5: 307–504. 

Champion, G. C. 1894. On the Tenebrionidae collected in Australia 

and Tasmania by Mr. James J. Walker, R.N., F.L.S., 

during the voyage of H.M.S. “Penguin”, with descriptions 

of new genera and species. Transactions of the Entomological 

Society of London, 1894: 351–408, pl. 8. 

Champion, G. C. 1895. A list of Tenebrionidae supplementary 

to the “Munich” Catalogue. Mémoires de la Société Entomologique 

de Belgique, 1: 1–264 pp. 

Chatanay, C. 1914. Tenebrionidae de Mahatsinjo. Annales de 

la Société entomologique de France, 83: 458–554. 

Chatanay, J. 1917. Matériaux pour servir à l’étude de la faune 

entomologique de l’Indo-Chine française réunis par M. 

Vitalis de Salvaza. Bulletin du Muséum National d’Histoire 

Naturelle, 23: 229–255. 

Cherney, L. S. 2005. Darkling-beetles (Coleoptera, Tenebrionidae). 

Fauna of Ukraine. Vol. 19. Beetles. Issue 10. Kiev, 

Naukova Dumka. 431 pp. (In Russian). 

Cherney, L. S. and L. V. Fedorenko. 2006. Keys to darkling 

beetles (Coleoptera, Tenebrionidae) of Fauna of Ukraine 

(imago, larvae, pupae). Kiev: Kolobig, 247 pp. (In Russian). 

Chevrolat, L. A. A. 1873. Description de quelques espèces nouvelles 

de coléoptères de Syrie. Annales de la Société Entomologique 

de France, 3: 203–218. 

Chikatunov, V., Lillig, M., Pavlicek, T., Blaustein, L. and E. 

Nevo. 1997. Biodiversity of insects at a microsite, ‘Evolution 

Canyon’, Nahal Oren, Mt. Carmel, Israel. Coleoptera: 

Tenebrionidae. Journal of Arid Environments, 37: 

367–377. 

Chûjo, M. 1963. A new species of Gonocephalum Solier from 

Okinawa and Takara Is. (Coleoptera, Tenebrionidae). 

Kontyû (Tokyo), 31: 149–153. 

Chûjô, M. 1966. Taxonomic study on the Tenebrionidae 

(Coleoptera) of the Ryukyu Islands. Journal of the Faculty 

of Agriculture, Kyushu University, 14: 1–32, 2 pls. 

Chûjô, M. 1968. Coleoptera from Southeast Asia (VIII). Memoirs 

of the Faculty of Education Kagawa University, Part 

II, 171: 21–23. 

Chûjô, M. 1973. Five new species of the Tenebrionidae from 

Japan and her adjacent regions, with a new record of 

Gonocephalum coenosum Kaszab from the Senkaku 

island (Coleoptera). Esakia, 8: 13–23. 

Chûjô, M. T. 1975. Tenebrionidae of the Nansei Islands I 

(Coleoptera). Esakia, 9: 15–24, 2 pls. 

Chûjô, M. 1977. Tenebrionidae of the Nansei Islands. II 

(Coleoptera). Esakia, 10: 1–19. 

Chûjô, M. T. 1978a. Tenebrionidae of the Nansei Islands III 


Chûjô, M. T. 1978b. Tenebrionidae of the Nansei Islands IV 


Chûjô, M. T. 1995. Gonocephalum ficifolium, a new species of 

Tenebrionidae from Japan (Coleoptera). Esakia, 35: 

113–116. 

Davletshina, A. G. 1967. Species composition and distribution 

of darkling beetles (Tenebrionidae) on biotops in southern 

western Kyzylkum. Zoologicheskiy zhurnal, 46(4): 

524–536. (In Russian). 

Dobrovolskiy, B. V. 1951. Pest beetles. Rostov-on-Don, Rostizdat. 

455 pp. (In Russian).


Dobrovolskiy, B. V. 1959. Distribution of pest insects. Moscow, 

State press “Soviet Science”. 216 pp. (In Russian). 

Drinkwater, T. W. 1999. Regional distribution of species of 

Gonocephalum (Col. Tenebrionidae) in fields in South 

Africa. African Entomology, 7: 183–187. 

Duftschmid, C. E. 1812. Fauna Austriae, oder Beschreibung 

der österreichischen Insekten für angehende Freunde der 

Entomologie. Zweyter Theil. Leipzig: Akademie Buchhandlung, 

viii + 311 pp. 

Erichson, W. F. 1843. Beitrag zur Insecten-Fauna von Angola 

in besonderer Beziehung zur geographischen Verbreitung 

der Insecten in Afrika. Archiv für Naturgeschichte, 9: 

199–267. 

Español, F. 1943. Misión cientifica E. Morales Agacino, Ch. 

Rungs y B. Zolotarevsky a Ifni y Sáhara Español. Tenebrionidae 

(Col.). 1.a parte. Eos, 19: 119–148. 

Español, F. 1944. Nuevos datos para el conocimiento de los 

Tenebriónidos (Col.) del Sáhara Español. Eos, 20: 7–30. 

Español, F. 1946. Tenebriónidos (Col.) nuevos o interesantes 

recogidos por D. Eugenio Morales y D. Joaquín Mateu en 

el Sáhara Español. Eos, 22: 107–122, pls. 8–10. 

Español, F. 1947. Coleópteros de Lanzarote. Graellsia, 5: 

83–97. 

Español, F. 1952. Los Gonocephalum de España (Col. Tenebrionidae). 

Graellsia, 10: 5–14. 

Español, F. 1953. Misiones saharianas de l’Institut scientifique 

Chérifien (1950–1951) Col. Tenebrionidae. Bulletin 

de la Société des Sciences Naturelles du Maroc, [1951] 31: 

287–312. 

Español, F. 1959. Opatrinae del Rif, Marruecos (Col. Tenebrionidae). 

Eos, 35: 243–255. 

Español, F. 1959. Los Gonocephalum de Canarias. Anuario de 

Estudios Atlánticos, 5: 105–113. 

Español, F. 1962a. Tenebriónidos del Médano (Tenerife). 

Coleotera. Graellsia, 229–235. 

Español, F. 1962b. Los Opatrinae de las Islas Canarias (Col. 

Tenebrionidae). Eos, 38: 203–221. 

Español, F. 1963. Datos para el conocimiento de los tenebriónidos 

del Mediterráneo occidental (Coleoptera). Eos, 

39: 185–209. 

Español, F. 1969. Tenebrionidos de la región de Sax-Salinas, 

provincia de Alicante (Coleoptera). Graellsia, [1968], 24: 

79–86. 

Español, F. and Viñolas, A. 1983. Revisión de los Gonocephalum 

del grupo prolixum (Col., Opatrinae). Eos, 59: 

31–39. 

Fabricius, J. C. 1775. Systema entomologicae, systens insectorum 

classes, ordines, genera, species, adiectis synonymis, 

locis, descriptionibus, observationibus. Flensburgi et Lipsiae, 

Libraria Kortii, xxxii + 832 pp. 

Fabricius, J. C. 1801. Systema Eleutheratorum secundum 

ordines, genera, species adiectis synonyxmis, locis, observationibus, 

descriptionibus. Tomus I. Kiliae, Bibliopolii 

Academici, xxiv + 506 pp; Tomus II. Kiliae, Bibliopolii 

Academici, 687 pp. 

Fairmaire, L. 1849. Essai sur les Coléoptères de la Polynésie 

(suite). Revue et Magazin de Zoologie Pure et Appliquée, 

1: 445–460. 

Fairmaire, L. 1863. Coléoptères d’Algérie rapportés par M. 

Obier de Baulny. Annales de la Société Entomologique de 

France, 4: 637–648. 

Fairmaire, L. 1882. Coléoptères Hétéromères de Sumatra. 

Notes from the Leyden Museum, 4: 219–265. 

Fairmaire, L. 1883. Essai sur les Coléoptères de l’Archipel de 

la Nouvelle-Bretagne. Annales de la Société Entomologique 

de Belgique, 27: 1–58. 

Fairmaire, L. 1888. Notes sur les Coléoptères des environs de 

Pékin (2 e partie). Revue d’Entomologie (Caen), 7: 111–160. 

Fairmaire, L. 1891: Descriptions de Coléoptères des Montagnes 

de Kashmir. Comptes-Rendus des Séances de la 

Société Entomologique de Belgique, 1891: lxxxviii–ciii. 

Fairmaire, L. 1893. Coléoptères du Haute Tonkin. Annales de 

la Société Entomologique de Belgique, 37: 303–325. 

Fairmaire, L. 1893b. Coléoptères nouveaux des Indes Orientales, 

de la famille des Scarabaeidae, Rhipidoceridae, 

Tenebrionidae et Oedemeridae. Notes from the Leyden 

Museum, 15: 17–64. 

Fairmaire, L. 1894. Coléoptères de l’Afrique intertropicale et 

australe. Troisième note. Annales de la Société Entomologique 

de Belgique, 38: 651–679. 

Fairmaire, L. 1894b. Hétéromères du Bengale. Annales de la 

Société Entomologique de Belgique, 38: 16–43. 

Fairmaire, L. 1896. Hétéromères de l’Inde recueillis par M. 

Andrewes. Annales de la Société Entomologique de Belgique, 

40: 6–62. 

Faldermann, F. 1837. Fauna entomologica trans-Caucasica. 

Pars II. Nouveaux Mémoires de la Société Impériale des 

Naturalistes de Moscou, 5: 1–433, 16 pls. 

Fauvel, A. 1867. Catalogue des coléoptères de la Nouvelle 

Calédonie et dépendances avec descriptions, notes et synonymies 

nouvelles. Bulletin de la Société Linnéenne de 

Normandie 2: 172–214, pl 1, figs 13–17. 

Fauvel, A. 1904. Faune analytique des Coléoptères de la Nouvelle 

Calédonie. 2 e Partie. Revue d’Entomologie, 23: 

113–208. 

Ferrer, J. 1993a. Essai de revision des espèces africaines et 

européennes appartenant au genre Gonocephalum Solier 

(Coleoptera, Tenebrionidae). Atti del Museo Civico di Storia 

Naturale di Trieste, 45: 59–150. 

Ferrer, J. 1994. Beitrag zur Kenntnis der Tenebrioniden-Fauna 

Burundis (Insecta, Coleoptera). Veröffentlichungen des 

Naturkundemuseums Erfurt, 13: 200–208. 

Ferrer, J. 1995a. Essai de revision des espèces africaines 

et europèennes appartenant au genre Gonocephalum 

Solier (Coleoptera, Tenebrionidae). Deuxième partie. 

Atti del Museo Civico di Storia Naturale di Trieste, 46: 

1–75. 

Ferrer, J. 1995b. Contribution to the knowledge of the Tenebrionidae 

of Somalia (Coleoptera). Frustula Entomologica, 

18: 1–76. 

Ferrer, J. 1996. Contributions to the knowledge of the Tenebrionidae 

of Kenya (Insecta: Coleoptera). Frustula Entomologica, 

19: 51–109. 

Ferrer, J. 1998. Contribution a la connaissance des Tenebrionidae 

de Madagascar (Insecta, Coleoptera). Entomofauna, 

19: 353–404. 

Ferrer, J. 2000a. Nouvelles espèces de Gonocephalum Solier, 

1834 et de Pachypterus Lucas, 1849 du Kenya et de Madagascar 

(Coleoptera, Tenebrionidae, Opatrini). New 

species of Gonocephalum Solier, 1834 and Pachypterus 

Lucas, 1849 from Kenya and Madagascar (Coleoptera, 

Tenebrionidae, Opatrini). Zoosystema, 22: 825–833.

Ferrer, J. 2000b. Revision des espèces africaines et européennes 

appartennantes au genre Gonocephalum Solier (Coleoptera, 

Tenebrionidae, Opatriini). Troisième partie. Atti 

del Museo Civico di Storia Naturale di Trieste, 48: 69–153. 

Ferrer, J. 2001. Contribucion al conocimiento de los tenebriónidos 

de Tanzania y descripcion de una nueva especie 

de Praeugena Maklin (Insecta, Coleoptera). Rivista 

Piemontese di Storia Naturale, 22: 225–237. 

Ferrer, J. 2004. Additions et corrections a ma revision des 

Gonocephalum Solier (1834), d’Europe et d’Afrique 

(Coleoptera Tenebrionidae). Additions and corrections to 

my revision of the European and African Gonocephalum 

Solier (1834) (Coleoptera Tenebrionidae). Bulletin de la 

Société Entomologique de France, 109: 83–91. 

Ferrer, J. 2007. Gonocephalum yelamosi Español, & Viñolas, 

1983 = Gonocephalum calcaripes (Karsch, 1881), (nouvelle 

synonymie) (Coleoptera, Tenebrionidae, Opatrini). 

Nouvelle Revue d’Entomologie, (N.S.), 24: 92–93. 

Ferrer, J. 2010. Taxonomic notes on the genus Gonocephalum 

Solier, 1834, with description of new taxa 

(Coleoptera, Tenebrionidae). Annales Zoologici, 62: 

231–238. 

Ferrer, J. and L. Soldati. 1999. Contribution à l’étude des 

Tenebrionidae de Turquie (Insecta, Coleoptera). Entomofauna, 

20: 53–92. 

Fischer von Waldheim, G. 1830. Note sur quelques nouvelles 

espèces d’insectes; dans une lettre à Mr. de Steven. Bulletin 

de la Société Impériale des Naturalistes de Moscou, 

2: 183–188. 

Fischer von Waldheim, G. 1844. Spicilegium Entomographiae 

Rossicae. Bulletin de la Société Impériale des Naturalistes 

de Moscou, 17: 3–144. 

Gardini, G. 1974. Materiali per lo studio dei Tenebrionidi dell’Arcipelago 

Toscano (Col. Heteromera) (Studi sulla Riserva 

Naturale dell’Isola di Montecristo. 17). Lavori della 

Societa Italiana di Biogeografia, 4: 637–723. 

Gebien, H. 1906. Ueber die von Fabricius beschriebenen 

Typen von Tenebrioniden in den Museen von Kopenhagen 

und Kiel. Deutsche Entomologische Zeitschrift, 1906: 

209–237. 

Gebien, H. 1908. Tenebrionidae, pp. 325–348. In: Michaelsen, 

W. and Hartmeyer, R. (eds.), Die Fauna Südwest-Australiens. 

Ergebnisse der Hamberger südwest-australischen 

Forschungsreise 1905. Vol 1. Part 10. Jena: Gustav 

Fisher. 

Gebien, H. 1910a. 7. Coleoptera. 19. Tenebrionidae, pp. 

363–396. In: Sjöstedt, Y. (ed.): Wissenschaftliche Ergebnisse 

der Schwedischen Zoologischen Expedition nach 

dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen 

Deutsch-Ostafrikas 1905–1906 unter der 

Leitung von Prof. Dr. Yngve Sjöstedt. 1. Band. Stockholm: 

Königl. Schwedisches Akademie der Wissenschaften, 

442 pp. 

Gebien, H. 1910b. Tenebrionidae, pp. 1–354. In: Schenkling, S. 

(ed.), Coleopterorum Catalogus. Vol. 18. Berlin: W. Junk. 

Gebien, H. 1911. Coleoptera, Tenebrionidae, pp. 57–79. In: 

Schubotz, H. (ed.): Wissenschaftliche Ergebnisse der 

Deutschen Zentral-Afrika-Expedition 1907–1908 unter 

Führung Adolf Friedrichs, Herzogs zu Mecklenburg. 

Band IV. Zoologie II. Leipzig: Klinkhardt & Biermann, 

485 pp. 


Gebien, H. 1913. H. Sauter’s Formosa-Ausbeute. Tenebrionidae 

(Coleopt.). Archiv für Naturgeschichte, 79A 9: 1–60, 

1 pl. 

Gebien, H. 1913b. Die Tenebrioniden der Philippinen. The 

Philippine Journal of Science, 8 (Sec. D): 373–433. 

Gebien, H. 1914. Fauna Simularensis. Coleoptera, Fam. Tenebrionidae. 

Notes from the Leyden Museum, 36: 61–80. 

Gebien, H. 1920a. Coleoptera Tenebrionidae. Nova Guinea, 13: 

213–500. 

Gebien H. 1920b. Käfer aus der Familie Tenebrionidae gesammelt 

auf der “Hamburger deutsch-südwestafrikanischen 

Studienreise 1911”. Hamburgische Universität Abhandlungen 

aus der Auslandskunde Band 5. Reihe C Naturwissenschaften 

Band 2. Hamburg: L. Friederichsen & Co., 168 

pp, 2 pls. 

Gebien, H. 1921. Die Tenebrioniden Westafrikas. Archiv für 

Naturgeschichte, Abteilung A, [1920], 86: 1–256. 

Gebien, H. 1922. The Percy Sladen Trust Expedition to the 

Indian Ocean in 1905, under the leadership of Mr. J. Stanley 

Gardiner, M. A. Vol. II. No. V. – Coleoptera, Heteromera: 

Tenebrionidae. Transactions of the Linnean Society 

of London, Series 2, Zoology, 18: 261–324. 

Gebien, H. 1927. Fauna Sumatrensis (Beitrag Nr. 31) Tenebrionidae 

(Col.). Supplementa Entomologica, 15: 22–58. 

Gebien, H. 1935. Tenebrionidae. In: Van Stralen, V. É. (ed.): 

Résultats Scientifiques du Voyage aux Indes Orientales 

Néerlandaises de LL. AA. RR. Le Prince et la Princesse 

Léopold de Belgique. Invertébrés: Hexapodes. Mémoires 

du Musée Royal d’histoire Naturelle de Belgique. Hors 

Série, 4(11): 53–77. 

Gebien, H. 1939. Katalog der Tenebrioniden. Mitteilungen der 

Münchner Entomologischen Gesellschaft, 29: 443–474. 

Gebler, F. A. v. 1860. Coleopterum species novae. a Dr. 

Schrenk in deserto kirgiso-songorico anno 1843 detectae. 

Bulletin de la Société Impériale des Naturalistes de 

Moscou, 3: 1–39. 

Gemminger, M. 1870. In: Gemminger M. & Harold B.: Catalogus 

coleopterorum hucusque descriptorum synonymicus 

et systematicus. Monachii, E. H. Gummi. Tom VII, pp. 

1801–2180 + (9). 

Germar, E. F. 1824. Species insectorum novae aut minus cognitae, 

descriptionibus illustratae. Volumen Primum. 

Coleoptera. Halae: J. C. Hendelii et filii, xxiv + 624 pp, 

pl. 1. 

Gerstaecker, A. 1855. Diagnosen neuer Coleopteren und Lepidopteren. 

Berlin: Monatsberichte der Berliner Akademie, 

1855: 636–639. 

Gerstaecker, A. 1871. Beitrag zur Insektenfauna von Zanzibar. 

Archives für Naturgeschichte, 37: 42–86. 

Gerstaecker, A. 1873. Gliederthiere (Insecten, Arachniden, 

Myriapoden und Isopoden). Fam. Melasoma, Latr., pp. 

162–192, pl. 9–10. In: Baron Carl Claus von der Decken's 

Reise in Ost-Africa in den Jahren 1859–1865. Dritter Band: 

Wissenschaftliche ergebnisse. Zweite Abtheilung. Leipzig 

und Heidelberg: C.F. Winter'sche Verlagshandlung. 

Gilyarov, M. S. (ed.). 1964. Key to soil-inhabiting larvae of 

insects. Moscow: Nauka, 919 pp. (In Russian). 

Girard, C. 1975. Étude des peuplements de Coléoptères Tenebrionides 

de la savane de Lamto (Côte D’Ivoire). Annales 

de la Société Entomologique de France (N.S.) II (2): 

335–381.


Gravely, F. H. 1915. XLII. Coleoptera, IX: Tenebrionidae. In: 

Zoological results of the Abor Expedition, 1911–12. 

Records of the Indian Museum, [1912–1922], 8: 519–536, 

pls 43, 44. 

Gridelli, E. 1930. Risultati zoologici della Missione inviata dalla 

R. Società Geografica Italiana per l’esplorazione 

del’oasi di Giarabub (1962–1927). Coleotteri. Annali del 

Museo Civico di Storia Naturale di Genova, 54: 1–486 + 

[1], 1 map. 

Gridelli, E. 1934. Materiali zoologici raccolti della spedizione 

italiana al Karakoram (1929 – Anno VII). Coleoptera – 

Tenebrionidae. Atti del Museo Civico di Storia Naturale 

Trieste, 12: 37–68, pls. 9, 10. 

Gridelli, E. 1939. Coleotteri dell’Africa orientale italiana. 6 e 

Contributo. Bollettino della Società Entomologica Italiana, 

71: 75–79. 

Gridelli, E. 1940. Coleotteri dell’Africa orientale italiana 

11 o Contributo. Materiali per lo studio della fauna Eritrea 

raccolti nel 1901–03 dal Dott. Alfredo Andreini. Memorie 

della Società Entomologica Italiana, [1939], 18: 

219–258. 

Gridelli, E. 1945. Coleotteri dell’Africa orientale (XIV. contributo). 

Appunti per una monografia delle specie etiopiche 

del genere Gonocephalum Sol. (Coleopt. Tenebr.). Atti del 

Museo Civico di Storia Naturale Trieste, 16: 1–28. 

Gridelli, E. 1948. Coleotteri dell’Africa Tropicale XVII contributo. 

Ulteriori appunti per una monografia delle specie 

africaine del genere Gonocephalum Sol. (Coleopt. Tenebr.). 

Atti del Museo Civico di Storia Naturale Trieste, 17: 

1–56. 

Gridelli, E. 1953. Catalogo ragionato delle specie di Coleotteri 

Tenebrionidi dell’Arabia. Atti del Museo Civico di storia 

naturale di Trieste, 19: 1–70. 

Gridelli, E. 1954. Catalogo ragionato delle species della 

famiglia Tenebrionidae a me nota dell’Afghanistan (Coleoptera). 

Atti del Museo Civico di Storia Naturale 

Trieste, 19: 169–292. 

Grimm, R. 1981. Die Fauna der Ägäis-Insel Santorin. Teil 2. 

Tenebrionidae (Coleoptera). Stuttgarter Beiträge zur 

Naturkunde Ser. A, 348: 1–14. 

Grimm, R. 1985. Zur Kenntnis der Tenebrioniden aus Süditalien 

(Insecta: Coleoptera). Stuttgarter Beiträge zur 

Naturkunde Ser. A, 379: 1–32. 

Grimm, R. 1986a. Tenebrionidae aus Niger und Mali (Coleoptera, 

Tenebrionidae). Mitteilungen Münchener Entomologischen 

Gesellschaft, 76: 71–78. 

Grimm, R. 1986b. Tenebrionidae vom Maltesischen Archipel 

(Insecta: Coleoptera). Stuttgarter Beiträge zur Naturkunde, 

Serie A: 392. 

Grimm, R. 1991. Tenebrioniden von der Insel Zypern (Insecta: 

Coleoptera). Biocosme Mesogéen, 8: 15–49. 

Grimm, R. 1995. Beitrag zur Kenntnis der Schwarzkaferfauna 

der Kanarischen Inseln (Coleoptera, Tenebrionidae). Contribution 

to the study of Tenebrionidae from the Canary 

Islands (Coleoptera). Mitteilungen Münchener Entomologischen 


Grimm, R. 2002. Tenebrionidae von Gambia (Insecta, Coleoptera). 

Entomofauna, 23: 353–380. 

Grimm, R. 2011. New and little known species of Tenebrionidae 

(Coleoptera) from Borneo (2). Stuttgarter Beiträge 

zur Naturkunde, Serie A, Neue Serie, 4: 249–257. 

Guérin-Ménéville, F. E. 1834. Matériaux pour une classification 

des Mélasomes. (Extraits d’une monographie de cette 

famille). Magasin de Zoologie, 1: 1–39, 19 pls. 

Harold, E. von. 1876. Bericht über eine sendung Coleopteren 

aus Hiogo. Abhandlungen vom Naturwissenschaftlichen 

Vereine zu Bremen, 5: 115–135. 

Harold, E. von. 1878. Beiträge zur Käferfauna von Japan. 

(Viertes Stück). Deutsche Entomologische Zeitschrift, 22: 

65–88. 

Heller, K. M. 1916. Die Käfer von Neu-Caledonien und den 

benachbarten Inselgruppen, pp. 229–365. In: Roux, F. S. a. 

J. (ed.), Nova Caledonia, Zoologie II, Wiesbaden. 

Herbst, J. F. W. 1793. Natursystem aller bekannten in-und 

ausländischen Insekten, als eine Forsetzung der von Büffon 

schen Naturgeschichte. Der Käfer fünfter Theil. Berlin: 

Pauli, 392 pp., pls. 44–59 + J–N. 

Hesse, A. J. 1935. Scientific results of the Vernay-Lang Kalahari 

expedition, March to September 1930. Tenebrionidae 

(Coleoptera). Annals of the Transvaal Museum, 16: 

525–579. 

Hope, F. W. 1831. Synopsis of the new species of Nepal Insects 

in the Collection of Major General Hardwicke, pp. 21–31. 

In: Gray, J. E. The Zoological Miscellany. London, Treuttel, 

Wurtz and Co. 

Hua, Li. 2002. Tenebrionidae, pp. 133–148. In: List of Chinese 

Insects, vol. 2. Guangzhou: Zhongshan (Sun Yat-sen) University 

Press. 

Iwan, D., Ferrer, J. and M. Raś. 2010. Catalogue of the World 

Gonocephalum Solier, 1834 (Coleoptera, Tenebrionidae, 

Opatrini). Part 1. List of the species and subspecies. Annales 

Zoologici, 60: 245–304. 

Israelson, G., Machado, A., Oromí, P. and T. Palm. 1982. 

Novedades para la fauna de las Islas Canarias, Vieraea, 2: 

109–134. 

Jambazishvili, M. Ya. 2000. Catalogue of darkling beetles 

(Coleoptera: Tenebrionidae) of Georgia. Trudy Instituta 

zoologii Academy of Sciences of Georgia, 20: 185–193. (In 

Russian). 

Kaluzhnaya, N. S. 1982. A review of the tenebrionid beetles 

(Coleoptera, Tenebrionidae) of the north-western Caspian 

lowland. Entomologicheskoe Obozrenie, 61: 67–80. (In 

Russian). 

Karsch, F. 1881. Die Käfer der Rohlf’s Afrikanischen Expedition. 

Berliner Entomologische Zeitschift, 21(1): 41–50, pl. 

2. 

Kaszab, Z. 1939. Zoologische Ergebnisse der ersten (VI–X. 

1936.) und zweiten (V–VIII. 1937.) Forschungsreise N. 

Vasvári’s in Kleinasien. II. Schwarzkäfer (Tenebrionidae). 

Matematischer und Naturwissenschaftlicher 

Anzeiger der Ungarischen Akademie der Wissenschaften, 

58: 582–590. 

Kaszab, Z. 1940. Beiträge zur Kenntnis der Tenebrioniden- 

Fauna Kleinasiens. Folia Entomologica Hungarica, 5: 2–7. 

Kaszab, Z. 1941. Tenebrioniden aus Formosa (Col.). Stettiner 

Entomologische Zeitung, 102: 51–72. 

Kaszab, Z. 1942. Beiträge zur Kenntnis der orientalischen 

Opatrinen (Col. Teneb.). Mitteilungen der Münchner Entomologischen 


Kaszab, Z. 1946. Monographie der Leiochrinen. Budapest: 

Ungarisches Naturwissenschaftliches Museum, 221 pp, 

1 pl.

Kaszab, Z. 1951. A new species of Gonocephalum (Col., Tenebrionidae) 

from the Philippine Islands. Annals and Magazine 

of Natural History, (12)4: 182–185. 

Kaszab, Z. 1952. Die indomalaischen und ostasiatischen 

Arten der Gattung Gonocephalum Solier (Coleoptera, 

Tenebrionidae). Entomologische Arbeiten aus dem Museum 

G. Frey, 3: 416–688. 

Kaszab, Z. 1954a. A csendes-óceáni szigetek Tenebrionidáinak 

(Coleoptera) biogeographiája. Zur Biogeographie der 

Tenebrioniden der Pazifischen Inseln. Állattani Közlemények, 

44: 71–79. 

Kaszab, Z. 1954b. Über die von Herrn J. Klapperich in der chinesischen 

Provinz Fukien gesammelten Tenebrioniden 

(Coleoptera). Annales Historico-Naturales Musei Nationalis 

Hungarici, Series nova, 5: 248–264. 

Kaszab, Z. 1957. Neue Tenebrioniden aus Iran 1954 (Coleopt.) 

(Ergebnisse der Entomologischen Reisen Willi Richter, 

Sttugart, im Iran 1954 und 1956, Nr. 7). Jahreshefte des 

Veriens für Vaterländsiche Naturkunde in Württemberg, 

112: 60–65. 

Kaszab, Z. 1960. Die Tenebrioniden Afghanistans, auf Grund 

der Ergebnisse der Sammelreise des Hern J. Klapperich in 

den Jahren 1952/53 (Col.). Entomologische Arbeiten aus 

dem Museum G. Frey, 11: 1–171. 

Kaszab, Z. 1961a. Wissenschaftliche Ergebnisse der von Dr. F. 

Schmid in Pakistan und Iran gesammelte Tenebrioniden 

(Coleoptera). Miscelánea Zoológica, 1: 115–121. 

Kaszab, Z. 1961b. Ergebnisse der österreichisechen Karakorum-Expedition 

1958 Coleoptera- Tenebrionidae. Annales 

Historico-Naturales Musei Nationalis Hungarici, 53: 

341–355. 

Kaszab, Z. 1961c. Zwei neue Tenebrioniden (Coleoptera) aus 

Arabien und Kleinasien. Abhandlungen und Berichte aus 

dem Staatlichen Museum für Tierkunde in Dresden, 26: 

169–175. 

Kaszab, Z. 1961d. Ergebnisse der Deutschen Afghanistan- 

Expedition 1956 der Landessamlungen für Naturkunde 

Karlsruhe. Tenebrionidae (Coleoptera). Beiträge zur 

Naturkundlichen Forschung in Südwestdeutschland, 19: 

235–238. 

Kaszab, Z. 1961e. Neue Tenebrioniden (Coleoptera) aus der 

Zoologischen Staatssammlung in Münnchen. Mitteilungen 

der Münchener Entomologischen Gesellschaft, 51: 213–230. 

Kaszab, Z. 1963a. Angaben zur Kenntnis der Tenebrioniden 

des Tschadsee-Gebietes, nebst einer Revision der 

afrikanischen Mesomorphus-Arten (Coleoptera). Revue de 

Zoologie et de Botanique Africaines, 68: 341–385. 

Kaszab, Z. 1963b. Ergebnisse der Zoologischen Nubien-Expedition 

1962 Teil X Coleoptera-Tenebrionidae. Annalen des 

Naturhistorischen Museums in Wien, 66: 453–456. 

Kaszab, Z. 1964. Über die Tenebrioniden einiger japanischen 

Inseln (I). Entomological Review of Japan, 16: 39–49. 

Kaszab, Z. 1965a. Wissenschaftliche Eregbnisse der von 

Dr. F. Schmid in Indien gesammelten Tenebrioniden 

(Coleoptera). Separata de Miscelànea Zoológica, 2: 

107–130. 

Kaszab, Z. 1965b. Österreichische entomologische Expeditionen 

nach Persien und Afghanistan. Beträige zur Coleopterofauna. 

Teil I: Tenebrionidae, Meloidae und Pedilide. 

Annalen des Naturhistorischen Museums in Wien, 68: 

667–670. 


Kaszab, Z. 1965c. Angaben zur Kenntnis der Tenebrioniden- 

Fauna der Mongolischen Volksrepublik (Coleoptera). Acta 

Zoologica Academiae Scientiarum Hungaricae, 11: 

295–346. 

Kaszab, Z. 1965d. Zoologische Ergebnisse der Forschungen 

von Dr. T. Pócs in der Volksrepublik Vietnam Tenebrionidae 

(Coleoptera). Annales Historico-Naturales Musei 

Nationalis Hungarici, 57: 287–296. 

Kaszab, Z. 1967a. Die Tenebrioniden der Westmongolei 

(Coleoptera). Acta Zoologica Academiae Scientiarum 

Hungaricae, 13: 279–351. 

Kaszab, Z. 1967b. Ergebnisse der Albanien-Expedition 1961 

des Deutschen Entomologischen Institutes. 70. Betrag. 

Coleoptera: Tenebrionidae. Beiträge zur Entomologie, 

3–4: 547–571. 

Kaszab, Z. 1968a. Tenebrionidae und Meloidae (Coleoptera) 

aus Nord-Korea, gesammelt von M. Mroczkowski und A. 

Riedel im Jahre 1965. Annales Zoologici, 26: 7–14. 

Kaszab, Z. 1968b. Beiträge zur Kenntnis der Fauna Afghanistans 

(Sammelergebniss von O. Jakeš 1963–64, D. Povolný 

1965, D. Povolný & Fr. Tenora 1966, J. Šimek 1965–66, 

D. Povolný, J. Geiser, Z. Šebek & Fr. Tenora 1967) Tenebrionidae, 

Col. Časopis Moravského Musea, Vědy přírodní 

53, Supplementum, 7–124 pp, 4 pls. 

Kaszab, Z. 1969. The Scientific Results of the Hungarian soil 

Zoological Expedition to the Brazzaville-Congo. 37. 

Coleoptera: Tenebrionidae. Annales Historico-Naturales 

Musei Nationalis Hungarici, 61: 225–265. 

Kaszab, Z. 1970a. Beiträge zur Kenntnis der Tenebrioniden- 

Fauna von Nepal (Coleoptera). Khumbu Himal, 3: 

422–434. 

Kaszab, Z. 1970b. The Scientific Results of the Hungarian Soil 

Zoological Expeditions to New Guinea. Tenebrionidae 

(Coleoptera). Annales Historico-Naturales Musei Nationalis 

Hungarici, 62: 257–280. 

Kaszab, Z. 1970c. Beiträge zur Kenntnis der Fauna Afghanistans 

(Sammelergebniss von O. Jakeš 1963–64, D. Povolný 

1965, D. Povolný & Fr. Tenora 1966, J. Šimek 1965–66, 

D, Povolný, J. Geiser, Z. Šebek & Fr. Tenora 1967). Tenebrionidae, 

Col. (Fortsetzung). Časopis Moravského Musea, 

Vědy přírodní 54, Supplementum: 5–182 pp, 23 pls. 

Kaszab, Z. 1971. The Scientific Results of the Hungarian Zoological 

Expeditions to Tanganyika. 15. Coleoptera: Tenebrionidae. 

Annales Historico-Naturales Musei Nationalis 

Hungarici, 63: 225–238. 

Kaszab, Z. 1972. Neue Tenebrioniden-Arten aus Laos (Coleoptera). 

Entomologische Arbeiten aus dem Museum G. 

Frey, 23: 287–297. 

Kaszab, Z. 1973. Neue Beiträge zur Kenntnis der Tenebrioniden-Fauna 

van Afghanistan (Coleoptera). Entomologia 

Scandinavica, 5: 35–58. 

Kaszab, Z. 1974. Neue Angaben zur Kenntniss der Tenebrioniden 

(Coleoptera) von Afghanistan. Annales Historico- 

Naturales Musei Nationalis Hungarici, 66: 187–203. 

Kaszab Z. 1975. Tenebrioniden (Coleoptera) aus Nepal, II. 

Annales Historico-Naturales Musei Nationalis Hungarici, 

67: 119–126. 

Kaszab, Z. 1976. Tenebrionidae der Nepal-Expeditionen von 

Dr. J. Martens (1969–1974) (Insecta: Coleoptera). Senckenbergiana 

Biologia, 57: 241–283. 

Kaszab, Z. 1977. Tenebrionidae det Nepal-Expeditionen von


Dr. J. Martens (1969–1974) (Insecta: Coleoptera). Senckenbergiana 

Biologica, 57: 241–283 

Kaszab, Z. 1978a. In: Kaszab Z., Schawaller, W. & N. G. 

Skopin. Systematik und Ökologie einiger Tenebrionidae 

aus Kashmir und Ladakh. Senckenbergiana Biologica, 59: 

215–234. 

Kaszab, Z. 1978b. Vier neue Tenebrioniden aus Nordbengal 

(Coleoptera). Folia Entomologica Hungarica, 31: 173–176. 

Kaszab, Z. 1979a. Tenebrioniden aus Südindien (Coleoptera). 

Acta Zoologica Academiae Scientiarum Hungaricae, 25: 

259–310. 

Kaszab, Z. 1979b. Faunistik der Tenebrioniden von Sri Lanka 

(Coleoptera). Folia Entomologica Hungarica, 32: 43–128. 

Kaszab, Z. 1979c. Insects of Saudi Arabia. Coleoptera: Fam. 

Tenebrionidae. Fauna of Saudi Arabia, 1: 257–288. 

Kaszab, Z. 1980a. Tenebrionidae (Coleoptera) aus Rennell 

und anderen Solomon Inseln. The Natural History of Rennell 

Island, Britisch Solomon Islands, 8: 33–48. 

Kaszab, Z. 1980b. Angaben zur Kenntnis der Tenebrioniden 

Nordvietnams (Coleoptera). Annales Historico-Naturales 


Kaszab, Z. 1982a. Die Tenebrionide 1978: 230; n Neukaledoniens 

und der Loyaute-Inseln (Coleoptera). Folia Entomologica 

Hungarica, 43: 1–294. 

Kaszab, Z. 1982b. Insects of Saudi Arabia. Coleoptera: Fam. 

Tenebrionidae (part 2). Fauna of Saudi Arabia, 4: 

124–245. 

Kaszab, Z. 1985. Beiträge zur Kenntnis der pazifischen Tenebrioniden 

(Coleoptera). Folia Entomologica Hungarica, 

46: 23–63. 

Kaszab, Z. 1986a. Tenebrioniden (Coleoptera) aus Neukaledonien. 

Annales Historico-Naturales Musei Nationalis 

Hungarici, 78: 151–175. 

Kaszab, Z. 1986b. Beiträge zur Kenntnis der Tenebrioniden 

(Coleoptera) aus Arabien. Annales Historico-Naturales 


Kaszab, Z. and M. Chûjô. 1964. 25. Family Tenebrionidae (2). 

Nature and Life in S. E. Asia, 3: 252–237 + 2 pls. 

Kaszab, Z. and M. Chûjô. 1966. Coleoptera from Southeast 

Asia (V). 14, Familie Tenebrionidae (1). Memoires of the 

Faculty of Education Kagawa University, Part II, No. 140: 

51–56 + 1 pls. 

Kaluzhnaya, N. S. 1982. A review of the tenebrionid beetles 

(Coleoptera, Tenebrionidae) of the north-western Caspian 

lowland. Entomologicheskoe Obozrenie, 61: 67–80. 

Kelejnikova, S. I. 1968. Soil larvae of darkling beetles (Coleoptera, 

Tenebrionidae) of the fauna of Soviet Union. I. 

Investigations on fauna of Soviet Union (Insects). Proceedings 

of Zoological museum of Moscow State University, 11: 

205–233. (In Russian). 

Keskin, B. 1999. Balçova baraji (Izmir; Türkiye) civar1 Tenebrionidae 

(Coleoptera) faunas1. Türkiye entomoloji dergisi, 

23: 211–224. 

Koch, C. 1935. Wissenschaftliche Ergebnisse der entomologischen 

Expedition seiner Durchlaucht des Fuersten A. Dalla 

Torre e Tasso nach Aegypten und auf der Halbinsel 

Sinai. VII. Tenebrionidae (Coleoptera). Bulletin de la 

Société Royale Entomologique d’Egypte, 19: 2–111. 

Koch, C. 1939. Die Käfer der libyschen Ausbeute des Herrn 

Georg Frey. Mitteilungen der Münchner Entomologischen 


Koch, C. 1945. Über die Tenebrioniden der Gipfelfauna des 

Hohen und Mittleren Atlas Marokkos. Ergebnisse der 

Forschungsreise R. Paulian und A. Villiers nach dem 

Hohen Atlas Marokkos 1938. XIX. Beitrag. Eos, [1944], 20: 

387–433. 

Koch, C. 1953. The Tenebrionidae of southern Africa. XXVI. 

New Port. East African species collected by Dr. A. J. Barbosa. 

Revista da faculdade de ciencias, Universidade de 

Lisboa. Serie C. Ciencias Naturais, 3: 239–310. 

Koch, C. 1955. The Tenebrionidae of Southern Africa. XXX. On 

Some Remarkable New Species of Gonocephalum. Annals 

of the Transvaal Museum, 22: 523–530. 

Koenig, E. 1899. Coleoptera Caucasica. P. 339–403. In: G. 

Radde. Die Sammlungen des Kaukasischen Museums. 

Part 1. Zoology. Tiflis, 474 pp. 

Kolbe, H. 1883. Neue Coleoptera von Westafrika. Berliner 

Entomologische Zeitschrift, 27: 15–36. 

Kolbe, H. 1897. Coleopteren. Die Käfer Deutsch-Ost-Afrikas. 

Berlin: Dietrich Reimer, 368 pp. 

Kolbe, H. 1902. Koleopteren der Aldabra-Inseln. Abhandlengen 

Senckenbergischen Naturforschenden Gesellschaft, 

26: 567–586. 

Krynicki, I. 1832. Enumeratio Coleopterorum Rossiae meridionalis 

et praecipue in Universitatis Caesareae Charkoviensis 

circulo obvenientium, quae annorum 1827–1831 

spatio observavit. Bulletin de la Société Impériale des Naturalistes 

de Moscou, 5: 65–179. 

Kryzhanovsky, O. L. 1965. Composition and origin of herpetobiontic 

fauna of Middle Asia (mainly on material on beetles). 

Leningrad: Nauka, 420 pp. (In Russian). 

Kulikovsky, E. A. 1897. Materials to the fauna of Coleoptera of 

Southern Russia. Mémoires de la société des naturalistes 

de la Nouvelle-Russie, 21: 1–274. (In Russian). 

Kulzer, H. 1954. Achter Beitrag zur Kenntnis der Tenebrioniden 

(Col.). Entomologische Arbeiten aus dem Museum 

G. Frey, 5: 20–73, pls 5–7. 

Kulzer, H. 1956. Bemerkenswerte Tenebrioniden aus der 

Thar-Wüste. 14. Beitrag zur Kenntnis der Tenebrioniden 

(Col.). Entomologische Arbeiten aus dem Museum G. Frey 

7: 635–653. 

Kulzer, H. 1957. Coleoptera: Tenebrionidae. Insects of Micronesia, 

17: 185–256. 

Kühnelt, W. 1957. Ergebnisse der österreichischen Iran-Expedition 

1949/50. Die Tenebrioniden Irans. Sitzungsberichte 

der österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche 

Klasse, 166: 65–102. 

Kühnelt, W. 1965. Tenebrionidae. 60 pp. In: Kanellis, A. Catalogus 

Faune Graeciae. Pars I. To Wono, Thessaloniki. 

Küster, H. C. 1845. Die Käfer Europa’s Nach der Natur Beschkrieben. 

Nürnberg, Bauer & Raspe, 3: 2 + 100 pp. 

Küster, H. C. 1849a. Die Käfer Europa’s. Nach der Natur 

Beschrieben. Nürnberg: Bauer & Raspe, 16: 2 + 100 pp, 

3 pls. 

Küster, H. C. 1849b. Die Käfer Europa’s. Nach der Natur 

Beschrieben. Nürnberg: Bauer & Raspe, 19: 2 + 100 pp, 

2 pls. 

Kwieton, E. 1981. Coléoptères Tenebrionidae constatés par 

l’expédition de Tis au Pakistan. Annotationes Zoologicae 

et Botanicae (Bratislava), 143: 1–20. 

Kwon, Y. J. and Y. S. Choi. 1986. Check list of family Tenebrionidae 

from Korea. Insecta Koreana, 6: 105–113.

Labrique, H. and G. Chavanon. 2008. Coléoptères nouveaux du 

Maroc oriental (Tenebrionidae, Scarabaeidae, Aphodiidae, 

Carabidae et Curculionidae). Bulletin de la Société 

entomologique de Mulhouse, 64: 19–36. 

Labrique, H. and A. Janati-Idrissi. 2009. Espèces nouvelles et 

localisations nuvelles pour le Moyen-Atlas central et oriental 

(Coleoptera, Tenebrionidae). Bulletin de la Société 

entomologique de Mulhouse, 65: 19–28. 

Labrique, H. and G. Chavanon. 2001. Résultats d’une campagne 

de prospection entomologique dans le Maroc oriental 

(Coleoptera Tenebrionidae). Cahiers Scientifiques. 

Centre de Conservation et d’Étude des Collections. 

Muséum d’Histoire Naturelle de Lyon, 2: 75–87. 

Labrique, H. and G. Chavanon. 2002. Résultats de la deuxième 

campagne de prospection entomologique dans le Maroc 

oriental: Tenebrionidae (Coleoptera) (2ème contribution 

à la connaissance des Tenebrionidae du Maroc). 

Cahiers Scientifiques. Centre de Conservation et d’Étude 

des Collections. Muséum d’Histoire Naturelle de Lyon, 2: 

65–81. 

Lacordaire, T. 1859. Histoire naturelle des Insectes. Genera 

des Coléoptères ou exposé méthodique et critique de tous 

les genres proposés jusqu’ici dans ce ordre d’Insectes. 

Tome Cinquième. Paris: Librairie Encyclopédique de 

Roret. Première partie, 1–400 pp, seconde partie, pp. 

401–750. 

LeConte, J. L. 1874. Descriptions of new Coleoptera, chiefly 

from the Pacific Slope of North America. Transactions of 

the American Entomological Society, 5: 43–72. 

Leo, P. and S. Fattorini. 2002. New records of tenebrionid beetles 

from the Aegean Islands (Coleoptera: Tenebrionidae). 

Biocosme Mesogéen, 18: 157–165. 

Lewis, G. 1894. On the Tenebrionidae of Japan. The Annals 

and Magazine of Natural History, (6)13: 377–400, 465–485, 

pl. 13. 

Lillig, M. and H. J. Bremer. 2002. Tenebrionidae der nördlichen 

Provinzen der Republik Sudan. (Coleoptera: Tenebrionidae). 

Coleoptera, 6: 35–101. 

Lillig, M. and T. Pavlíček. 2003. The Darkling Beetles of the 

Sinai Peninsula. Coleoptera: Tenebrionidae (excl. Lagriinae 

et Alleculinae). Zoology in the Middle East: Supplementum. 

Heidelberg: Kasparek Verlag, 87 pp. 

Lindberg, H. 1950. Beitrag zur Kenntnis der Käferfauna der 

Kanarischen Inseln. Commentationes Biologicae, 10: 1–20. 

Lindberg, H. 1962. Coleoptera Insularum Canariensium. III. 

Tenebrionidae. Commentationes Biologicae, 25: 1–85, 

11 pls. 

Lo Cascio, P. and A. Scupola. 2004. New data and some considerations 

on the tenebrionid beetles of Astypalea and 

Nisyros Islands (Dodecanese, Greece) (Coleoptera: Tenebrionidae). 

Biocosme Mesogéen, 20: 113–126. 

Lucas, P. H. 1846. Histoire naturelle des animaux articulés. 

Deuxième partie. Insectes. In: Exploration scientifique de 

l’Algérie pendant les années 1840, 1841, 1842 publiée par 

ordre du Gouvernement et avec le concours d’une Commission 

Académique. Sciences Physiques. Zoologie 2. 

Paris: A. Bertrand, 590 pp. 

Lucas, P. H. 1849. Histoire naturelle des animaux articulés. 

Deuxième partie. Insectes. In: Exploration scientifique de 

l’Algérie pendant les années 1840, 1841, 1842 publiée 

par ordre du Gouvernement et avec le concours d’une 


Commission Académique. Sciences Physiques. Zoologie 2. 

Paris: A. Bertrand, [1846], 590 pp. 

Marcuzzi, G. 1962. Studi ecologici e faunistici sui Tenebrionidi 

(Col. Heteromera) della Puglia. Memoria de Biogeografia 

Adriatica, 6: 1–79. 

Marcuzzi, G. 1970. Contributo a la conoscenza dei Tenebrionidi 

delle Egadi. Bolletino della Societá Entomologica Italiana, 

100: 87–91. 

Marseul, S.-A. 1876. Coléoptères du Japon recueillis par M. 

Georges Lewis. Enumération des Hétéromères avec la 

description des espèces nouvelles. Annales de la Société 

Entomologique de France, 6: 93–142, 315–340. 

Masumoto, K. 1983. Notes and descriptions of Japanese Tenebrionidae 

(III). Entomological Review of Japan, 38: 

173–176. 

Masumoto, K. 1985. The Japanese species of the genus 

Gonocephalum (Coleoptera, Tenebrionidae). Elytra, 12: 

27–37. 

Masumoto, K. 2006. Tenebrionid Beetles (Coleoptera) from 

the Palau Islands Collected by Keiichi Kakahashi. Entomological 

Review of Japan, 61: 143–161. 

Masumoto, K. and K. Ando. 2003. New or little–known tenebrionid 

species from Japan (Part 4) (Coleoptera). Entomological 

Review of Japan, 58: 173–182. 

Masumoto, K., Ando, K. and K. Akita. 2008. A New Synonym of 

Gonocephalum (Coleoptera: Tenebrionidae) from Japan. 

Entomological Review of Japan, 63: 121–123. 

Medvedev, G. S. 1965. Fam. Tenebrionidae – Darkling beetles, 

pp. 356–381. In: Key to Insects of European part of USSR. 

Vol. 2. Coleoptera. Moscow-Leningrad: Nauka. 668 pp. (In 

Russian). 

Medvedev, G. S. 1968. Coleoptera, Tenebrionidae, Platynotini, 

Dendarini, Pedinini, Dissonomini, Pachypterini, Opatrini, 

Heterotarsini. Fauna SSSR, 19, 2. Leningrad: Nauka, 285 

pp. (In Russian). 

Medvedev, G. S., Protsenko, A. I. and T. Kaltaev. 1975. To the 

fauna of darkling beetles of Kirgizia. Entomologicheskie 

issledovaniya v Kirgizii, 10: 82–95. (In Russian). 

Medvedev, G. S. and G. M. Abdurakhmanov. 1988. Check-list of 

beetles of the family Tenebrionidae of South West of 

Palaearctic, pp. 71–99. In: Abdurakhmanov, G. M. Eastern 

Caucasus by view of entomologist. Makhachkala: Dagestanian 

books press, 136 pp. (In Russian). 

Medvedev, G. S. 1990. Keys to the darkling beetles of Mongolia. 

Trudy Zoologicheskogo Instituta, 220: 1–254. (In Russian). 

Medvedev, G. S. 1992. 97 th Fam. Tenebrionidae – Darkling beetles, 

pp. . 621–659. In: Ler, P. A. Key to Insects of Far East 

of USSR. Vol. 3. Beetles. Part 2. St.–Petersburg: Nauka, 


Medvedev, G. S. 2004. To the fauna of darkling beetles (Coleoptera, 

Tenebrionidae) of Lower Volga, pp. 166–167. In: 

Entomological investigation in Volga region. Saratov: 

Saratov State University, Issue 3, 184 pp. (In Russian). 

Medvedev, G. S. and Kaszab, Z. 1973. Ergebnisse Mongolisch- 

Sowetischen biologischen Expeditionen in der Mongolischen 

Volksrepublik (Tenebrionidae, excl. Tentyriini 

(Coleoptera). Folia Entomologica Hungarica, 26: 

79–111. 

Medvedev, G. S. and M. G. Nepesova. 1990. Composition and 

geographical distribution of fauna of darkling-beetles


(Coleoptera, Tenebrionidae) of Kopet-Dag. Entomologicheskoe 

obozrenie, 9: 879–889. (In Russian). 

Medvedev, G. S. and A. L. Lobanov. 1990. A faunistic list of 

tenebrionids (Coleoptera, Tenebrionidae) of the Mongolian 

People’s Republic with coordinates of the localities. 

Nasekomye Mongolii, 11: 139–204. (In Russian). 

Medvedev, G. S. and M. G. Nepesova. 1985. Opredelitel zhukovchernotelok 

Tukmenistana. Ashkhabad: Ylym, 177 pp. (In 

Russian). 

Medvedev, S. I. and D. S. Shapiro. 1957. To the knowledge of 

fauna of beetles (Coleoptera) of Moldavskoy SSR and 

applied regions of USSR. Trudy NII biologii i biologicheskogo 

faculteta Kharkovskogo universiteta, 30: 173–206. 

(In Russian). 

Ménétriés, E. 1832. Catalogue raisonnée des objets de Zoologie 

recueillis dans un voyage au Caucase et jusqu’aux 

frontières actuelles de la Perse, entrepris par ordre de S. 

M. L’Empereur. St.-Pétersbourg: Académie des Sciences, 

XXXIII + 272 pp. 

Ménétriés, E. 1849. Catalogue des Insectes recueillis par feu 

M. Lehmann, avec les descriptions des nouvelles espèces. 

(Seconde et dernière partie.) Coléoptères hétéromères. 

Mémoires de l’Académie Impériale des Sciences de 

St.–Pètersbourg, Sixième série, 6: 217–328, pls 3–5. 

Miedel, J. 1880. Observations sur les Opatrum. Deutsche Entomologische 

Zeitschrift, 24: 136–140. 

Miwa, Y. 1931. A Systematic Catalogue of Formosan Coleoptera. 

Report of the Department of Agriculture, Governmental 

Research Institute of Formosa, No. 55. Taihoku 

[= Taipei]: Taiwan Research Institute of Formosa, xi + 

359 pp. 

Montrouzier, X. 1860. Essai sur la faune entomologique de la 

Nouvelle-Calédonie (Balade). Annales de la Société Entomologique 

de France, (3)8: 229–308. 

Mordkovich, V. G. 2002. Species richness, population structure 

and functional significance of black-beetles (Coleoptera: 

Tenebrionidae) in steppe of Northern Asia. Russian 

Entomological Journal, 11: 57–68. (In Russian). 

Motschulsky, V. de. 1845. Remarques sur la collection de 

Coléoptères Russes. Bulletin de la Société Impériale des 

Naturalistes de Moscou, 18: 1–127. 

Motschulsky, V. de. 1849. Coléoptères reçus d’un voyage de M. 

Handschuh dans le midi de l’Espagne, énumerés et suivis 

de notes. Bulletin de la Société Impériale des Naturalistes 

de Moscou, 22: 52–163. 

Motschulsky, V. de. 1854. Diagnoses de Coléoptères nouveaux, 

trouvés par MM. Tatarinoff et Gaschkéwitsch aux 

environs de Pékin. Études Entomologiques, [1853] 2: 

44–51. 

Motschulsky, V. de. 1857. Insectes du Japon. Études Entomologiques, 

6: 25–41. 

Motschulsky, V. de. 1860a. Coléoptères de la Sibérie orientale 

et en particulier des rives de l’Amur par MM. Schrenck, 

Maack, Ditmar, Voznessenski etc. In: Dr. L. v. Schrenk’s 

Reisen und Forschungen im Amur–Lande. Band II. Zweite 

Lieferung. Coleopteren. St.-Pétersbourg: Kaiserlicher 

Akademie der Wissenschaften, 79–257 + (1) pp, pls 4–11, 

1 map. 

Motschulsky, V. de. 1860b. Insectes du Japon. Études Entomologiques, 

9: 4–39. 

Motschulsky, V. de. 1866. Catalogue des insectes reçus du 

Japon. Bulletin de la Société Impériale des Naturalistes de 

Moscou, 39: 163–200. 

Mulsant, E. 1854. Histoire naturelle des coléoptères de 

France. Latigènes. L. Maison, Paris. x + 396 + 2 pp. 

Mulsant, E. and B. Godart. 1859a. Description de trois espèces 

nouvelles de Coléoptères. Opuscules Entomologiques, 9: 

177–183. 

Mulsant, E. and B. Godart. 1859b. Description de trois espèces 

nouvelles de Coléoptères. Annales de la Société Linnéenne 

de Lyon, 30: 241–252. 

Mulsant, E. and C. Rey. 1859a. Essai d’une division des 

derniers Mélasomes. Annales de la Société d’Agriculture 

de Lyon, 3: 129–201. 

Mulsant, E. and C. Rey. 1859b. Descriptions de quelques 

coléoptères nouveaux. Annales de la Société d’Agriculture 

de Lyon, 3: 222–240. 

Nabozhenko, M. V. 1999. Landscape–ecological characteristics 

and common regularities of the darkling-beetles 

(Coleoptera, Tenebrionidae) fauna distribution in Southern 

Russia. Izvestia kharkovskogo entomologicheskogo 

obshestva, 7: 40–54. (In Russian). 

Nakane, T. 1963. New of little known Coleoptera from Japan 

and its adjacent regions, XIX. Fragmenta Coleopterologica, 

6–7: 26–30. 

Nakane, T. 1966. Coleoptera from Afghanistan. Results of the 

Kyoto University Scientific Expedition to the Karakorum 

and Hindukush, 8: 231–248. 

Nepesova, M. G. 1980. Darkling beetles of Turkmenia. Ashhabad: 

Ylym, 211 pp. (In Russian). 

Normand, H. 1936. Contribution au Catalogue des Coléoptères 

de la Tunisie, 10e fascicule. Bulletin de la Société d’Histoire 

Naturelle de l’Afrique du Nord, 27: 355–383. 

Ogloblin, D. A. and A. N. Kolobova. 1927. Darkling beetles 

(Tenebrionidae) and their larvae, harmful to agriculture. 

Trudy Poltavskoy selskochozyaistvennoy opytnoy stanzii, 

61: 1–60. (In Russian). 

Olivier, A. G. 1811. Encyclopédie méthodique. Histoire 

Naturelle. Insectes. Tome huitième. Paris: H. Agasse, 

722 pp. 

Olliff, A. S, 1887. IV. Insecta. In: Millington, J. A. M. 1888. 

Report on a small zoological collection from Norfolk 

Island. Proceedings of the Linnean Society of New South 

Wales, (2)2: 1001–1014. 

Olliff, S. A. 1889. The insects fauna of Lord Howe Island. Memoirs 

of the Australian Museum. No. 2, 2: 75–98. 

Oromí, P. 1982. Los Tenebriónidos de las Islas Canarias, pp. 

265–292. In: Instituto de Estudios Canarios, 50 Aniversario 

1932–1982, I Ciencias. La Laguna: Instituto de Estudios 

Canarios & Aula de Cultura del Cabildo, 352 pp. 

Ovchinnikov, S. V. 1996. Fam. Tenebrionidae – Darkling beetles, 

pp. 149–157. In: Genetical fund cadastre of Kyrgyzstan. 

Vol. 3. Superclassis Hexapoda (Entognatha and 

Insecta). Bishkek: Institute for Biology and Pedology, 

Kyrg. Acad. Sci., 400 pp. 

Palisot de Beauvois, A. M. F. J. 1821. Insectes recuillis en 

Afrique et en Amérique. Paris: Levrault, fol. 16, 270 pp., 

90 pls. 

Perty, J. A. M. 1831. Observationes nonnullae in Coleoptera 

Indiae Orientalis. Dissertatio philosophico-entomologica, 

quam unacum praemissis thesibus auctoritate et 

consensu illustris philosophorum ordinis in Academia

Ludovico-Maximilianea facultatem legendi rite adepturus. 

Monachii: M. Lindauer, xxxxiv pp., 1 pl. 

Pirnazarov, B. 1970. Darkling beetles of Karakalpakiya. Vestnik 

Karakalpakskogo otdelenia Academii Nauk Uzbekskoi 

SSR, 4: 23–28. (In Russian). 

Portevin, G. 1934. Histoire naturelle des Coléoptères de 

France, vol. 3. Heteromera, Phytophaga. Encyclopédie 

entomologique, Série A, 17. Paris: P. Lechevalier, 374 pp., 

pl. 11–15. 

Protsenko, A. I. and T. Kaltaev. 1974. Materials on fauna 

of darkling beetles (Coleoptera, Tenebrionidae) of Kirgizia. 

Entomologicheskie issledovaniya v Kirgizii (Entomological 

investigations in Kirgizia), 9: 21–45. (In Russian). 

Redtenbacher, L. 1849. Fauna Austriaca. Die Käfer. Nach der 

analytischen Methode bearbeitet. Wien: C. Gerold, xxvii + 

883 pp., 2 pls. 

Redtenbacher, L. 1858. Fauna Austriaca. Die Käfer. Nach der 

analytischen Methode bearbeitet. Zweite, gänzlich 

umgearbeitete, mit mehreren Hunderten von Arten und 

mit der Charakteristik sämmtlicher europäischen Käfergattungen 

vermehrte Auflage. Wien: Carl Gerold’s Sohn, 

cxxxvi + 1017 pp., 2 pls. 

Redtenbacher, L. 1874. Fauna Austriaca. Die Käfer , nach der 

analytischen Methode bearbeitet. Wien: Carl Gerold’s 

Sohn, 564 + 571 + viii + cliii pp. 

Reichardt, A. N. 1936. Darkling beetles of the tribe Opatrini 

(Coleoptera Tenebrionidae) of the Palearctic Region. Keys 

to the fauna of the USSR, published by the Zoological 

Institute of the Academy of Sciences 19. Moscow- 

Leningrad: Izdatelstvo Akademii Nauk SSSR, 224 pp. (In 

Russian). 

Reiche, L. J. 1850. Entomologie, pp. 257–532. In: Ferret. A. 

and J. G. Galinier (eds.), Voyage en Abyssinie dans les 

provinces du Tigre, du Samen et de l’Amhara. Tome 

Troisième. Paris: Paulin, 536 pp. 

Reiche, L. and F. Saulcy. 1857. Espèces nouvelles ou peu connues 

de Coléoptères, requeillis par M. F. de Saulcy, membre 

de l’Institut, dans son voyage en Orient. Annales de la 

Société Entomologique de France, (3)5: 169–276. 

Reitter, E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia 

centrali novissime lecta. IX. Tenebrionidae. Horae Societatis 

Entomologicae Rossicae, 21: 355–389. 

Reitter, E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia 

novissime lecta. XIII.. Tenebrionidae. Horae Societatis 

Entomologicae Rossicae, 23: 678–710. 

Reitter, E. 1904. Bestimmungs-Tabelle der Tenebrioniden- 

Unterfamilien: Lachnogyini, Akidini, Pedinini, Opatrini 

und Trachyscelini aus Europa und den angrenzenden 

Ländern. Verhandlungen des Naturforschenden Vereines 

in Brünn, [1903], 42: 25–189. 

Reitter, E. 1905. Acht neue Coleopteren aus der palaearktischen 

Fauna. Wiener Entomologische Zeitung, 24: 90–94. 

Reitter, E. 1911. Fauna Germanica. Die Käfer des Deutschen 

Reiches. Nach der analytischen Methode bearbeitet. III. 

Band. Stuttgart: K. G. Lutz, 436 pp, pls 81–128. 

Ren, G.-D. 1995b. Coleoptera: Tenebrionidae, pp. 233–236. 

In: Wu, H. (ed.), Insects of Baizhanzu Mountain Eastern 

China, China Forestry Publishing House, xiii + 586 pp. (In 

Chinese with English summary). 

Ren, G. D. 1998. Coleoptera: Tenebrionidae, pp. 108–114. In: 


Insects of Longwangshan Nature Reserve. China Forestry 

Publishing House, Beijing. 

Ren, G. 2001. Tenebrionidae, pp. 89–90. In: Fang, Z. and Wu, 

H. (eds.), A checklist of Insects from Zhejiang. 

Ren, G. 2002a. Tenebrionidae, pp. 137–170. Fauna of Insects 

in Fujian Province of China, vol 6. 

Ren, G. 2002b. Coleoptera: Tenebrionidae, pp. 388–392. In: 

Huang, F.S. (ed.), Forest Insects of Hainan. Beijing: Science 

Press, 1064 pp. 

Ren, G., Dai, J. and Y. Yu. 2000. The Form of Seven Species of 

Larvae of Gonocephalum (Coleoptera: Tenebrionidae). 

Journal of Ningxia Agricultural College, Vol. 21 No. 1: 1–6. 

Ren, G. and H. Hua. 2006. Tenebrionidae, pp. 265–274. Fanjingshan 

Jingguan Kunchou. 

Ren, G. and B. Ming. 2005. Coleoptera, Tenebrionidae. Insect 

Fauna of Middle–West Qinling Range and South Mountains 

of Gansu Province, 3: 379–389. 

Ren, G. and D. Saihong. 2001. Coleoptera, Tenebrionidae, pp. 

347–349. Insects of Tianmusshan National Nature 

Reserve. 

Ren, G. and X. Shen. 2002. Coleoptera: Tenebrionidae, pp. 

383–388. In: Shen, X. C and Y. Q. Zhao (eds.), Insects of 

the mountains Taihang and Tongbai Regions. Beijing: China 

Agricultural Science and Technology Press, 453 pp. 

Ren, G. and X. Yang. 2004a. Coleoptera, Tenebrionidae, 

pp. 311–319. Insects from Mt. Shwandashan Area of 

Guangxi. 

Ren, G. and X. Yang. 2004b. A New Species and Twelve New 

Records of The Tribe Opatrini in China (Coleoptera, Tenebrionidae). 

Acta Zootaxonomica Sinica, 29(2): 305–309. 

Rey, C. 1892. Remarques en passant. L’ Exhange, Revue Linnéenne, 

8: 42. 

Rosenhauer, W. G. 1847. Beiträge zur Insekten-Fauna 

Europas. Erstes Bändchen; enthält die Beschreibung von 

sechzig neuen Käfern aus Bayern, Tyrol. Ungarn. etx., so 

wie die Käfer Tyrols, nach dem Ergebnisse von vier 

Reisen. Erlangen: Theodor Blaesing, x + 160 pp., 1 pl. 

Sahlberg, C. R. 1823. Periculi Entomographici, species Insectorum 

nondum descriptas proposituri, Fasciculus. Aboae: 

Typis Frenckelliorum, 82 + (1), 4 pls. 

Schawaller, W. 1996. Tenebrionidae (Coleoptera) aus Nord- 

Griechenland: Habitate, Artengesellschaften und Verbreitung. 

Entomologische Blätter, 92: 3–18. 

Schawaller, W. 1997. The genus Gonocephalum Chevrolat 

(Coleoptera: Tenebrionidae) in the Nepal Himalayas. 

Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). 

559: 1–18. 

Schawaller, W. 2004. New species and records of Tenebrionidae 

(Coleoptera) from the Socotra Archipelago. Fauna of 

Arabia, 20: 439–458. 

Schawaller, W. 2006. New species, new records and new synonyms 

of Tenebrionidae (Insecta: Coleoptera) from Nepal. 

Veröffentlichungen des Naturkundemuseums Erfurt, 25: 

475–482, pls 9, 10. 

Schawaller, W. 2010. Order Coleoptera, family Tenebrionidae 

(with the exeption of the subfamily Alleculinae). Arthropod 

fauna of the UAE, 3: 253–278. 

Schneider, O. and Leder, H. 1878. Beiträge zur Kenntniss der 

kaukasischen Käferfauna. Verhandlungen der Naturforschenden 

Vereines in Brünn, 16: 3–258. 

Schuster, A. 1928. Tenebrionidae. Pp. 914–1024, pp. 817–1698


pp. In: Winkler, A. (ed.), Catalogus Coleopterorum regionis 

palaearcticae. Wien: Albert Winkler. 

Schuster, A. 1934. Tenebrioniden vom Toten Meer. Entomologische 

Blätter, 30: 180–182. 

Schuster, A. 1936. Tenebrionidae. Entomologische Sammelergebnisse 

der Deutschen Hindukusch-Expedition 1935 

der Deutschen Forschungsgemeinschaft. Arbeiten über 

Morphologische und Taxonomische Entomologie aus 

Berlin-Dahlem, 3: 192–201. 

Scupola, A. 1983. Tenebrionidi delle Regione Veronese, Coleoptera. 

Bolletino dell Museo di Storia Naturale de Verona, 

[1982], 2: 91–120, 1 tab. 

Seidlitz, G. von. 1891a. Fauna Baltica. Die Kaefer (Coleoptera) 

der Deutschen Ostseeprovinzen Russlands. Zweite neu 

bearbeitete Auflage. Königsberg: Hartungsche Verlagsdruckerei, 

LVI +192 + 818 pp, 1 pl. 

Seidlitz, G. K. M. von. 1891b. Fauna Transsylvanica. Die Kaefer 

(Coleoptera) Siebenbürgens. Königsberg: Hartungsche 

Verlagsdruckerei, lvi + 914 pp. 

Seidlitz, G. von. 1894. Tenebrionidae, pp. 401–608, May 1894. 

In: Kiesenwetter H. von. & Seidlitz G. von.: Naturgeschichte 

der Insecten Deutschlands. Erste Abteilung 

Coleoptera. Fünfter Band. Erste Hälfte. Berlin: Nicolaische 

Verlags-Buchhandlung, xxviii + 877 pp. 

Shalaby, F., El-Haidari, H. and A. I. Derwesh. 1966. Contribution 

to the Insect Fauna of Iraq. Bulletin de la Société 

Royale d’Entomologique d’Egypte, 77: 77–89. 

Skopin, N. G. 1961. Materialy po faune i ekologii chernotelok 

(Coleoptera, Tenebrionidae) yugo-vostochnogo Kazakhstana. 

Trudy Nauchno-Isledovatelskogo Instituta Zashchtity 

Rasteniy Kazakhstanskoy Akademii Selskokhozyastvennykh 

Nauk, 6: 172–208. 

Skopin, N. G. 1964. Materialy po faune i ekologii chernotelok 

(Coleoptera, Tenebrionidae) Mangyshlaka i severo-zapadnogo 

Ustyurta. Trudy Nauchno-Issledovatelskogo Institute 

Zashchity Rastenii Kazakhstanskoy Akademii Selskokhozyastvennykh 

Nauk, 8: 277–297. 

Skopin, N. G. 1966. Neue Tenebrioniden aus Zentralasien IV., 

nebst einigen systematischen sowie synonymischen 

Bemerkungen. Annales Historico-Naturales Musei Nationalis 

Hungarici, 58: 325–342. 

Soldati, F. 2010. Troisième campagne de prospection entomologique 

en Corse (Coleoptera, Tenebrionidae). R.A. 

R.E., 19(1): 1–10. 

Steven, C. 1829. Tentyriae et Opatra collectionis Stevenianae 

nunc Musei Universitatis Mosquensis. Nouvelles Mémoires 

de la Société Impérial des Naturalistes de Moscou, 1: 

83–100. 

Sturm, J. 1807. Deutschlands Fauna in Abbildungen nach der 

Natur mit Beschreibungen. V. Abtheilung. Die Insecten. 

Zweites Bändchen. Käfer. Nürnberg: Verfasser, 279 pp, 

52 pls. 

Sturm, J. 1843. Catalog meiner Insecten-Sammlung. Nürnberg: 

Verfasser, xii + 386 pp., 6 pls. 

Tajibaev, M. 1972. Darkling-beetles of (Coleoptera, Tenebrionidae) 

of low mountains of Southern Tajikistan. Entomologicheskoye 

obozrenie. 51: 274–281. (In Russian). 

Tezcan, S., Ferrer, J. and B. Keskin. 2000. Contribution to 

the study of tenebrionid beetles (Coleoptera: Tenebrionidae) 

in ecological cherry orchards in Izmir et Manisa 

provinces of Turkey. Türkiye entomoloji dergisi, 24: 

243–248. 

Tezcan, S., Karsavuran, Yu., Pehlivan, E., Keskin, B. and J. 

Ferrer. 2004. Contributions to the knowledge of the Tenebrionidae 

(Coleoptera) from Turkey. Part II. Opatrinae, 

Tenebrioninae, Adeliinae. Türkiye entomoloji dergisi, 28: 

163–180. 

Thomson, J. 1858. Voyage au Gabon. Histoire naturelle des 

Insectes et des Arachnides recueillis pendant un voyage 

fait au Gabon en 1856 et 1857 par M. Henry C. Deyrolle. 

Deuxième Partie. Insectes. I. Ordre Coléoptères. Archives 

Entomologiques, 2: 29–376. 

Uyttenboogaart, D. L. 1930. Contribution to the knowledge of 

the fauna of the Canary-Island. Synopsis of the results of 

the collecting-excursions 1925–1927. Tijdschrift voor 

Entomologie, 73: 211–235. 

Walker, F. 1858. Characters of some apparently undescribed 

Ceylon Insects. Annals and Magazine of Natural History, 

(3)2: 280–286. 

Walker, F. 1871. List of Coleoptera collected by J. K. Lord, 

Esq. in Egypt, Arabia and near the African shore of the 

Red Sea. With characters of the undescribed species. London: 

E. W. Janson, 19 pp. 

Wollaston, T. V. 1854. Insecta Maderensia; being an account of 

the insects of the islands of the Madeiran group. London: 

John van Voorst, xliii + 634 pp, 13 pls. 

Wollaston, T. V. 1861. On certain Coleoptera from the Island of 

St. Vincent. Annals and Magazine of Natural History, (3)7: 

197–206. 

Wollaston, T. V. 1864. Catalogue of the coleopterous insects of 

the Canaries in the collection of the British Museum. London: 

British Museum, xiii + 648 pp. 

Wollaston, T. V. 1865. Coleoptera Atlantidum, being an enumeration 

of the Coleopterous Insects of the Madeiras, Salvages, 

and Canaries. London: J. v. Voorst, 526 + 140 

(Appendix) pp, 1 map. 

Wollaston, T. V. 1867. Coleoptera Hesperidum, being an enumeration 

of the coleopterous insects of the Cape Verde 

Archipelago. London: J. van Voorst, xxxix + 285 pp. 

Yakobson, G. G. 1913. Beetles of Russia and Western Europe, 

pp. 721–864, tab. 76–83. Instructions to keys of beetles. 

Vol. 10. St.Petersburg: A.F. Devrien (in Russian). 

Yakobson, G. G. 1927. Key to beetles. Moskow-Leningrad: 

Gosudarstvennoe izdatelstvo, 522 pp. (in Russian). 

Yakobson, G. G. 1931. Key to beetles. Practical entomology. 

Iss. 7. Moskow-Leningrad: Gosudarstvennoe izdatelstvo, 


Yang, X. and G. Ren. 2005a. A list of the tribe Opatrini in China 

(Coleoptera: Tenebrionidae), pp. 183–189. In: Ren, G., 

Zhang, R. and F. Shi (eds), Classification and Diversity of 

Insects in China. 

Yang, X. and G. Ren 2005b. Coleoptera, Tenebrionidae pp. 

226–232. Guizhou Dashahe Kunchong. 

Yang, X. and G. Ren. 2007. Tenebrionidae, pp. 225–259. 

Leigongshan Jingguan Kunchong. 

Yu, Y. and G. Ren. 1994. Tenebrionid larvae of certain species 

of the opatrini – tribe from N. China (Coleoptera: Tenebrionidae), 

pp. 91–100. In: Lian, Z.M. (ed.), Insect Research, 

Vol. 1. Xi’an: Shaanxi Normal University Press. 

Received: May 10, 2011 

Accepted: June 20, 2011

CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834 ...

Create successful ePaper yourself

Delete template?

Save as template?