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5.3 Class Magnoliopsida – flowering plants - Cambridge University ...

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epidermal glands but in different families the glands are adapted<br />

to produce either mucilage (in Polygonaceae), excrete salt as in the<br />

halophytes, or digestive enzymes in the insectivorous Droseraceae and<br />

Nepenthaceae. In the latter, the gland-type is shared but the subsidiary<br />

insect-trapping apparatus is quite diverse ranging from pitchers<br />

(Nepenthes), sticky traps (Drosera, Drosophyllum) to spring traps<br />

(Dionaea, Aldovandra).<br />

Several families in the order have a peculiar flower development<br />

that starts in a polymerous way but becomes organised into a pseudodiplostemonous<br />

way. Stamens appear to be produced in pairs. In<br />

the Caryophyllaceae and Plumbaginaceae stamens are antipetalous<br />

and arise with the petal as a unit.<br />

Some families in the Caryophyllalaes (Caryophyllaceae, Chenopodiaceae)<br />

generally lack mycorrhizae perhaps because they tend to<br />

occupy nutrient rich fresh soils. In contrast, the insectivores in the<br />

Nepenthaceae, Droseraceae and Drosophyllaceae can inhabit nutrientpoor<br />

soils.<br />

santalales (sandlewoods)<br />

All five families of the Santalales (Santalaceae ∼500 species, Olacaceae<br />

∼200 species, Opiliaceae 28 species, Misodendraceae 8 species, and<br />

the Loranthaceae ∼940 species) include tropical parasitic species. The<br />

Loranthaceae are the most specialised parasites. The shrubby, liane<br />

or twining habit is common throughout the order.<br />

saxifragales (saxifrages, currants and stonecrops)<br />

The Saxifragales is a very diverse order and includes: trees<br />

(Hamamelidaceae ∼100 species of witch hazel and sweet gum, Cercidiphyllaceae<br />

-- katsura, Altingiaceae), shrubs (Grossulariaceae ∼325<br />

species of currants and gooseberies) and showy ornamentals like the<br />

Paeoniaceae (∼34 species of peony), but the two largest families are<br />

herbs, mainly rosette-forming Saxifragaceae (∼475 species of Astilbe<br />

and saxifrage) or leafy succulents, the Crassulaceae (∼1280 species of<br />

Sempervivum, Echeveria, Sedum and Kalanchoe). Many of these rosette<br />

formers in the Crassulaceae, Saxifragaceae and some other small<br />

families, are linked by a similar look to their flowers and a number<br />

of characters such as a persistent scarious calyx and cellular<br />

endosperm.<br />

vitales (vines)<br />

This order is of supreme importance to us as the source of wine from<br />

the grape vine Vitis. Many other genera are twining vines with or<br />

without tendrils (Rhoiocissus, Cissus). Cyphostemma is a caudiciform and<br />

Leea a shrub and small tree.<br />

berberidopsidales<br />

This tiny order of two families, one with only one species Aextoxicon<br />

from Chile and the other with only two genera Berberidopsis and<br />

Streptothamnus from Chile and eastern Australia.<br />

<strong>5.3</strong> CLASS MAGNOLIOPSIDA <strong>–</strong> FLOWERING PLANTS 239<br />

Figure 5.87. Aizoaceae:<br />

Mesembryanthemum.<br />

Figure 5.88. Grossulariaceea:<br />

Ribes.<br />

Figure 5.89. Paeoniaceae:<br />

Paeonia.<br />

Figure 5.90. Berberidopsidales:<br />

Berberidopsis.

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