HUNTERIA - Gregory S. Paul
HUNTERIA - Gregory S. Paul
HUNTERIA - Gregory S. Paul
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plete (Table 1).Assuming a typical sauropod mass/femur length<br />
ratio it may have weighed over 50 tons. Lacking further data<br />
it is not possible to estimate the animal's total length. Another<br />
super-sized sauropod is the femur of the titanosaur Antarctosaurus<br />
giganteus (Huene, 1929; Van Valen, 1969;<br />
Table 1). It is in the same class as the biggest brachiosaur<br />
femora, but since the rest of the animal is poorly known, the<br />
best that can be said is that it was probably similar in mass<br />
too. The South African (Mclachlan and McMillan, 1976)and<br />
Laotian (Hoffet, 1942)femora Anderson et a1.(1985)cited are<br />
not from uniquely large individuals (Table 1).<br />
In the final analysis, B. (G.) brancai and B. (B.) altithorax<br />
were about as big as any other known sauropods. Just<br />
as importantly, B. (G.) brancai holds the record for the<br />
biggest species for which all of the skeleton is known. It was<br />
also the tallest, but not the longest. The largest known<br />
sauropods appear to cluster around 50 tons in lean condition,<br />
perhaps a third more in prime fat-bearing condition. This<br />
should not be taken as an ultimate limit. The sample of all<br />
known sauropods is a tiny fraction of the sampled populations<br />
available for many single species of living animals. Even<br />
larger sauropods certainly await discovery, and it is improbable<br />
that the largest were preserved in the fossil record. It is interesting<br />
that, in living tetrapods, extremely rare "world record"<br />
individuals are often twice as massive as average individuals<br />
(McWhirter and McWhirter, 1986).In this view sauropods of<br />
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<strong>HUNTERIA</strong> VOL. U, no. 3, pp. 12, February 19, 1988<br />
GREGORY S. PAUL<br />
CITED REFERENCES<br />
100tons are not unrealistic, especially if bearing large amounts<br />
of seasonal fat.<br />
In comparison, Balaenoptera musculus typically weighs<br />
80-100 tons, and may reach 200 in feeding season (Ellis, 1980;<br />
McWhirter and McWhirter, 1986). Unfortunately there has<br />
never been a rigorous study of the mass of baluchitheres and<br />
the largest fossil proboscideans, which may rival each other<br />
as the biggest of terrestrial mammals. The baluchitheres and<br />
bigger mammoths appear to be rather gracile, and may not<br />
have been as massive as sometimes suggested. At perhaps 20<br />
tons or less, they certainly do not match the bigger sauropods<br />
in size. As for the greatest living land animal,Loxodonta<br />
africana bulls average 5 tons, often reach 7.5, and rarely reach<br />
about 10+ (Laws et al., 1975; McWhirter and McWhirter,<br />
1986).<br />
ACKNOWLEDGEMENTS<br />
Special thanks go to J. McIntosh and his bottomless store of Information<br />
on giant Mesozoic monsters. R. Bakker's many discussions on dinosaur<br />
anatomy were critical to this paper, as was his request that I prepare it.<br />
F. Boothman has provided important information and enjoyable discussion<br />
on the problem of big dinosaurs. K. Carpenter, M. Brett-Surman,<br />
D. Berman, D. Russell, R. Molnar, Dong Z., P: Currie and D. Gasparini<br />
have provided much helpful information and discussion on sauropods. Many<br />
thanks to H. Jaeger for his hospitality during my visit to the HMN. P. Sites<br />
helped reproduce the figures.<br />
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.:.