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Agglutinated Middle-Upper Eocene foraminifera in Jabal Hafit, Al ...

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20 H. S. ANAN<br />

Superfamily Lituolacea de BLAINVILLE, 1827<br />

Family Haplophragmoididae MAYNC, 1952<br />

Genus Haplophragmoides CUSHMAN, 1910<br />

Type species : Nonion<strong>in</strong>a canariensis dʼORBIGNY, 1839<br />

Haplophragmoides walteri (GRZYBOWSKI, 1898)<br />

Pl. I, fig. 4<br />

1898. Trochamm<strong>in</strong>a walteri GRZYBOWSKI, p. 290, p1. 11, fig.<br />

31.<br />

1981. Haplophragmoides walteri (GRZYBOWSKI).- MORGIEL &<br />

OLSZEWSKA, p. 10, p1. 3, fig. 15.<br />

1982. Haplophragmoides walteri (GRZYBOWSKI).- MILLER et<br />

al., p. 20, p1. 2, fig. 7.<br />

The species is characterized by its <strong>in</strong>volutes planispiral<br />

test, agglut<strong>in</strong>ated wall with f<strong>in</strong>e quartz gra<strong>in</strong>s with<br />

siliceous cement, small umbonal depression on both<br />

sides, elongate triangular chambers <strong>in</strong>creas<strong>in</strong>g slowly <strong>in</strong><br />

size, slightly depressed and curved sutures.<br />

In the Polish External Carpathians the species ranges<br />

from Senonian to <strong>Eocene</strong> (MORGIEL & OLSZEWSKA, 1981)<br />

and from Early to <strong>Middle</strong> <strong>Eocene</strong> <strong>in</strong> Labrador Sea, North<br />

Atlantic (MILLER et al., 1982).<br />

In this study, the species occurs <strong>in</strong> the top <strong>Middle</strong> <strong>Eocene</strong><br />

(P14) of <strong>Jabal</strong> <strong>Hafit</strong>.<br />

Genus Ammobaculites CUSHMAN, 1910<br />

Type species : Spirol<strong>in</strong>a agglut<strong>in</strong>ans dʼORBIGNY, 1846<br />

Ammobaculites cubensis CUSHMAN & BERMUDEZ, 1937<br />

Pl. I, fig. 5<br />

1937. Ammobaculites cubensis CUSHMAN & BERMUDEZ, p. 106,<br />

p1. 16, figs 4, 16-18.<br />

1955. Ammobaculites cubensis CUSHMAN & BERMUDEZ.-<br />

ANSARY, p. 17, p1. 1, fig. 1.<br />

1994. Ammobaculites cubensis CUSHMAN & BERMUDEZ.- BOLLI<br />

et al., p. 325, fig. 62.3.<br />

The species was orig<strong>in</strong>ally described from the <strong>Eocene</strong><br />

of Cuba by CUSHMAN & BERMUDEZ (1937) and from the<br />

<strong>Middle</strong> and <strong>Upper</strong> <strong>Eocene</strong> of Egypt (ANSARY, 1955) and<br />

Tr<strong>in</strong>idad (BOLLI et al., 1994).<br />

It is recorded here <strong>in</strong> the top <strong>Middle</strong> <strong>Eocene</strong> (P14) of<br />

<strong>Jabal</strong> <strong>Hafit</strong>.<br />

Superfamily Spiroplectamm<strong>in</strong>acea CUSHMAN, 1927<br />

Family Spiroplectamm<strong>in</strong>idae CUSHMAN, 1927<br />

Subfamily Spiroplectamm<strong>in</strong><strong>in</strong>ae CUSHMAN, 1927<br />

Genus Spiroplect<strong>in</strong>ella KISELʼMAN, 1972<br />

Type species : Spiroplecta wrightii SILVESTRI, 1903<br />

Spiroplect<strong>in</strong>ella nuttalli (LALICKER, 1935)<br />

Pl. I, fig. 6<br />

1935. Spiroplectamm<strong>in</strong>a nuttalli LALICKER, p. 43, p1. 6, figs<br />

3, 4.<br />

1951. Spiroplectamm<strong>in</strong>a nuttalli LALICKER.- CUSHMAN &<br />

STAINFORTH, p. 143, p1. 25, figs 19, 20.<br />

1955. Boliv<strong>in</strong>opsis nuttalli (LALICKER) - ANSARY, p. 19, p1. 1,<br />

fig. 7.<br />

1997. Boliv<strong>in</strong>opsis nuttalli (LALICKER).- HUSSEIN, p. 11, fig.<br />

3.10.<br />

2000. Boliv<strong>in</strong>opsis cf. nuttalli (LALICKER).- ABUL-NASR, p. 72,<br />

fig. 14.11.<br />

LOEBLICH & TAPPAN (1988, p. 111, 112) noted that<br />

the biserial stage of the genus Spiroplect<strong>in</strong>ella is<br />

characterized by its lozenge shaped <strong>in</strong> cross section and<br />

may have a marg<strong>in</strong>al keel, while it is long and nearly<br />

constant width throughout <strong>in</strong> Boliv<strong>in</strong>opsis and ovoid <strong>in</strong><br />

Spiroplectamm<strong>in</strong>a. Hence, the species is related here to<br />

the genus Spiroplect<strong>in</strong>ella.<br />

The species was orig<strong>in</strong>ally described from the <strong>Eocene</strong> of<br />

Venezuela by LALICKER (1935), then <strong>in</strong> the same horizon<br />

of Ecuador (CUSHMAN & STAINFORTH, 1951) and Egypt<br />

(ANSARY, 1955 ; HUSSEIN, 1997 ; ABUL-NASR, 2000).<br />

It is recorded here from the top <strong>Middle</strong> to early Late<br />

<strong>Eocene</strong> (P14 - P15) of <strong>Jabal</strong> <strong>Hafit</strong>.<br />

Subfamily Vulvul<strong>in</strong><strong>in</strong>ae SAIDOVA, 1981<br />

Genus Vulvul<strong>in</strong>a dʼORBIGNY, 1826<br />

Type species : Vulvul<strong>in</strong>a capreolus CUSHMAN, 1928<br />

Vulvul<strong>in</strong>a haer<strong>in</strong>gensis (GÜMBEL, 1868)<br />

1868. Venil<strong>in</strong>a haer<strong>in</strong>gensis GÜMBEL, p. 71, pl. 2, fig. 84.<br />

1975. Vulvul<strong>in</strong>a haer<strong>in</strong>gensis (GÜMBEL).- PROTO DECIMA & de<br />

BIASE, p. 91, pl. 1, figs 4, 23.<br />

1975. Vulvul<strong>in</strong>a haer<strong>in</strong>gensis (GÜMBEL).- BRAGA & GRÜNIG,<br />

p. 102, pl. 5, fig. 5.<br />

1978. Vulvul<strong>in</strong>a haer<strong>in</strong>gensis (GÜMBEL).- PROTO DECIMA &<br />

BOLLI, p. 797, pl. 1, fig. 5.<br />

1988. Vulvul<strong>in</strong>a haer<strong>in</strong>gensis (GÜMBEL).- PARISI & COCCIONI, p.<br />

103, pl. 1, figs 8, 9.<br />

1996. Vulvul<strong>in</strong>a haer<strong>in</strong>gensis (GÜMBEL).- ANAN, p. 149, fig.<br />

3.3.<br />

The species was recorded <strong>in</strong> Italy (PROTO DECIMA & de<br />

BIASE, 1975 ; BRAGA & GRÜNIG, 1975 ; PARISI & COCCIONI,<br />

1988) and also <strong>in</strong> South Atlantic, DSDP Leg 40, west<br />

coast of Africa (PROTO DECIMA & BOLLI, 1978).<br />

The species was recorded <strong>in</strong> the Early <strong>Eocene</strong> (P9) of<br />

<strong>Jabal</strong> <strong>Hafit</strong> (ANAN, 1996). It occurs also <strong>in</strong> the <strong>Middle</strong>-<br />

Late <strong>Eocene</strong> (P14-P15) of the study section.<br />

Vulvul<strong>in</strong>a jarvisi CUSHMAN, 1932<br />

Pl. I, fig. 7<br />

1932. Vulvul<strong>in</strong>a jarvisi CUSHMAN, p. 84, pl. 10, fig. 20.<br />

1976. Vulvul<strong>in</strong>a jarvisi CUSHMAN.- BERGGREN & AUBERT, p.<br />

314, pl. 1, fig. 1.

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