404 <strong>Nuytsia</strong> Vol. 8, No. 3 (1992) (PERTH); 17 km NW of Koonjarra, D.W. Goodall 2511 (PERTH); 49 km NNE of Balladonia, G.J. Keighery 7623 (PERTH). Habitat. In shallow open depressions (dongas) that have a layer of heavy soil over limestone. Flowering time. July to October. Rhodanthe nullarborensis resembles H. tietkensii with which it has been confused in herbaria. The latter species may be recognised by its larger leaves, its larger and more hairy capitula, and by the presence of receptacular bracts. Johnson & Baird (1970) in referring to this species as Helipterum tietkensii state that it is heavily scented and that it forms dense colonies in the dongas. Rhodanthe polycephala (A. Gray) Paul G. Wilson, comb. nov. Pteropogon polycephalus A. Gray, Hooker’s J. Bot. Kew Gard. Misc. 4:268(1852). - Helipterum polycephalum (A. Gray) Benth., Fl. Austral. 3:649(1867). - Argyrocome polycephala (A.Gray) Kuntze, Revis. Gen. Pl. 1:309(1891). Type citation:'Swan river, Drummond’, n.v. Cassinia cuprea F. Muell., Fragm. 3:139(1863). Lectotype (here chosen): Near the Murchison, A. Oldfield (MEL 110308). Rhodanthe psammophila Paul G. Wilson, sp. nov. (Figure 4) Herba erecta ad 40 cm alta. Folia caulina alternata, anguste oblonga vel linearia, ad 6 cm longa, sessilia, parce decurrentia, lanata et minute stipitato-glandulosa. Capitula heterogama, sessilia, glomerata. Involucrum anguste turbinatum, c. 4 mm altum; bracteae 4-6-seriatae, obovatae, 3-4 mm longae, apice albo, rotundato, margine eroso; unguis bractearum interiorum dense lanato- ciliatus. Flosculi semper quinque, uno fertili, quatuor masculinis. Corolla actinomorpha, sparse puberula; lobi intra papillosi. Achenium compresso-obovoideum, c. 2.5 mm longum, sparse et minute puberulum; carpophorum annulare; pericarpium tenuiter crustaceum; testa membranacea pericarpium adnata. Pappus achenii fertile tarde deciduus; setae c. 13, anguste lineares, acuminatae, c. 2.0 mm longae, sparse ciliatae pilis gracilibus acuminatis. Achenium bracteis interioribus consociatis deciduum. Typus: Western Australia, 20 km north of Carnarvon, red sand dune, 25 September 1987, Paul G. Wilson 12604 (holo: PERTH; iso: AD, CANB, K, MEL, NSW, S). Erect shortly woolly annual branching at and above base, to 40 cm high. Leaves cauline, alternate, narrow-oblong to linear, acuminate, to 6 cm long, sessile, slightly decurrent along the margins, woolly all over and with minute shortly stipitate reddish brown globular glands. Capitula heterogamous, sessile in dense clusters, these shortly stipitate forming subglobular heads on slender peduncles 1-3 cm long. Involucre narrow-turbinate, c. 4 mm high; bracts 6-seriate, obovate, 3-4 mm long, rounded at apex, hyaline in lower 2/3, white petaloid (with flat cells) and wrinkled above with erose margin; claw of inner bracts densely long woolly ciliate on margin, the wool enveloping the ovaries; stereome linear, hard, green, slightly shorter than claw, passing downwards into a very short hard stipe. Receptacle hemispherical, c. 3 mm high and wide, glabrous, smooth. Florets consistently 5 only one
Paul G. Wilson, Helipterum: Part 1 Figure 4. Rhodanthe psammophila. A - Habit. B - Cluster of capitula. C - Capitulum. D - Florets with and without corolla. E - Carpopodium. F - Pappus bristle. G - Corolla. H - Style apex. I - Achene with pappus. J - Pericarp. K - Duplex myxogenic achenial hairs. From Paul G. Wilson 12604. 405
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