The genus Trichosanthes L. (Cucurbitaceae) in Thailand ...

THE GENUS TRICHOSANTHES L. (CUCURBITACEAE) IN THAILAND (B.E.E. DUYFJES & K. PRUESAPAN) 99
Leaves of adult plants shallowly or deeply 3-or 5-lobed, up to 4 ⁄5 deep. Male
bracts deeply and finely incised, up to ca. ⅓ deep. Male sepals with margin serrate or
margin with side-lobes.
Thailand.— NORTHERN: Mae Hong Son, Chiang Mai, Lampang; NORTH-
EASTERN: Sakon Nakhon, Nong Khai; CENTRAL: Lob Buri; EASTERN: Chaiyaphum;
SOUTH-EASTERN: Chanthaburi (Bo Rai, E of Makham).
Distribution.— S China (?), Laos, Cambodia, Vietnam (type), one collection
from Peninsular Malaysia.
Ecology.— Secondary roadside thickets, in mixed deciduous forest edges; on
limestone and granitic bedrock; at 100–1,200 m altitude.
Vernacular.— Khuea mak khi ka (เขือหมากขี้กา)
(Champasak, Laos), khi ka daeng.
(ขี้กาแดง).
Note.— Trichosanthes tricuspidata subsp. tricuspidata as here defined appears
quite homogeneous, except for the seeds. Mature seeds are known from Thailand from a
comparatively large number of collections, and show a rather problematic variation in
seed size and shape, comprising compressed forms with a square edge (e.g. Pooma et al.
2672), or more less-compressed forms with a rounded edge. Possible taxonomic
implications of this variation are as yet unclear.
subsp. javanica Duyfjes & Pruesapan, subsp. nov. a subspecies typica bracteis
masculis vadose grosse dentatis, sepalis masculis integris differt. Typus: Indonesia,
Java, W. J. de Wilde & Duyfjes 21777 (holotypus L; isotypus BO, K, L). Fig. 6B.
Trichosanthes tricuspidata auct. non Lour.: Blume, Bijdr. Fl. Ned. Ind.: 935, excl.
type. 1826; Cogn. in A. & C. DC., Monogr. Phan. 3: 374. 1881; Rugayah & W. J. de
Wilde, Blumea 42(2): 481, fig.1b, 2b & 2c. 1997; Rugayah, Trichosanthes
(Cucurbitaceae) in Malesia: 120, pl. 10. 1999.— T. bracteata auct. non. (Lam.) Voigt:
Backer in Backer & Bakh. f., Fl. Java 1: 304. 1963.— T. tricuspis Miq., Fl. Ind. Bat. 1
(1): 679. 1856.
Leaves of adult plants 3-lobed, to ⅓–½. Male bracts shallowly and coarsely
dentate. Male sepals (almost) entire.
Thailand.— SOUTH-WESTERN: Phetchaburi (Phonsena, De Wilde & Duyfjes
3982); SOUTH-EASTERN: Chanthaburi (Larsen et al. 32315; Seidenfaden 2878).
Distribution.— West Malesia (Java, type) to the Moluccas and Lesser Sunda
Islands (not in Philippines).
Vernacular.— Khi ka daeng (ขี้กาแดง).
Note.— Collections from Kaeng Krachan NP have conspicuously bright yellow
petal fringes, not white as in the specimens from the type locality (Java).
15. Trichosanthes truncata C. B. Clarke in Hook. f., Fl. Brit. Ind. 2: 608. 1879; Cogn.
in A. & C. DC., Monogr. Phan. 3: 364. 1881; Chakrav., Ind. J. Agric. Sc. 16(1): 22. 1946;
Monogr. Ind. Cucurbitaceae, 17(1): 39. 1959; S. K. Chen & C. H. Yueh, Acta Phytotax.