A NEW INFRAGENERIC CLASSIFICATION AND SyNOPSIS OF THE ...

366 LANKESTERIANA
19. Flowers very large; lip more than 8 cm long; stem conspicuously grooved and minutely papillose; leaves
oblong V. insignis
19. Flowers not very large; lip less than 6 cm long; stem terete, smooth; leaves elliptic V. helleri
20. Leaves elliptic; species from Brazil V. schwackeana
20. Leaves oblong-elliptic; species from Central America (but escaped in all tropical areas) 21
21. Leaves oblong, about four times as long as broad V. planifolia
21. Leaves lanceolate, about six times as long as broad V. tahitiensis
22. Stems more than 8 mm thick; leaves succulent; basal bracts of the inflorescence not conspicuously distichous;
lip with a thickened axial cushion near the lip apex, the veins of the distal part of the lip not thickened 23
22. Stems less than 8 mm thick, leaves coriaceous, not conspicuously succulent, basal bracts of the inflorescence
conspicuously distichous, lip without a thickened axial cushion near the lip apex, the veins of the distal part of
the lip thickened 28
23. Lip conspicuously longer than the petals, the midlobe of the lip strongly emarginated 24
23. Lip about as long as the tepals, the midlobe rounded to slightly notched 25
24. Tepals flared, ovary rather calyculate; species from Central America and NW South America V. calyculata
24. Tepals not flared, rather concave, species from SE South America V. chamissonis
25. Leaves very large, usually more than 22 × 11 cm, widely elliptic 26
25. Leaves medium-sized, less than 16 × 7 cm, oblong to oblong elliptic 27
26. Midlobe of the lip well-defined V. pompona
26. Midlobe of the lip obscure V. grandiflora
27. Lip strongly concave V. pseudopompona
27. Lip gently concave V. columbiana
28. Lip entire; sepals not conspicuously papillose, the papillae, if present, scattered 29
28. Lip obscurely 3-lobed; ovary, sepals and dorsal keel of the petals conspicuously granulose in the outer surface 30
29. Lip acute to subacute; leaves oblong, not acuminate V. gardneri
29. Lip apex widely rounded; leaves oblanceolate, acuminate-cuspidate V. dressleri
30. Inflorescence many-flowered, up to 60-flowered; leaves up to 26 × 8 cm; lip mid-lobe broadly ovate, with c. 10
thickened keels in the lip apex extending to the lateral lobes; species from Amazonia and Guyanas V. hostmanii
30. Inflorescence with less than 15 flowers; leaves less than 18 × 7 cm 31
31. Mid-lobe of the lip subquadrate-ovate, with 5-6 thickened keels on the apical, central part of the lip, not extending to
the lateral lobes, axial keel low; sepals rounded, slightly subcalyptrate; species from Central America V. cribbiana
31. Mid-lobe of the lip ovate, with c. 8 thickened keels, rather warty on the apical part of the lip, extending to
the lateral lobes, 1-3 elevated, axial keels; sepals acute and conspicuously calyptrate; species from Peru and
Bolivia V. weberbaueriana
Listed species, recognized as valid appear in bold
face. Those described in Vanilla and considered to be
synonyms are italicised.
Vanilla Plum. ex Mill., Gard. Dict. abridged de. 4, 3:
without page number (1754). Lectotype: Epidendrum
vanilla L. = Vanilla mexicana Mill. (Mansf. in
Kulturpfl. 2: 587, 1959)
Myrobroma Salisb., Parad. Lond. t. 82 (1807).
Vanillophorum Neck., Elem. Bot. 3: 184 (1790),
nom. inval.
LANKESTERIANA 9(3), January 2010. © Universidad de Costa Rica, 2010.
SyNOPSIS OF VANILLA
1. V. abundiflora J.J. Sm. in Bull. Jard. Bot.
Buitenzorg ser. 3, 2: 21 (1920). Types: Borneo, Koetei,
Soengei Djongkang, Tengaroeng, March 1918, W.C.
van Gelder 1; Soengei Landak, J.E. Teysmann s.n.;
Kapoeas, J.E. Teysmann 109898 (all syn. BO).
distribution: Borneo (Kalimantan Timur, Sarawak).
Similar to V. kinabaluensis because of its long
inflorescences with more than 100 successive flowers
and large leaves, but V. abundiflora has smaller flowers,
and the apical hairs of the lip are less conspicuous.