cryptic species within the dendrophidion vinitor complex in middle ...

SPECIES IN THE DENDROPHIDION VINITOR COMPLEX N Cadle 235
Cerro Delgadito, 2–4 mi W Santa Fe (Veraguas).
Approximately 08u309N, 81u079W (GE).
Cerro Malí (Darién). 08u079N, 77u149W (Fairchild
and Handley, 1966). According to Myers (1969:
25) Cerro Malí is about 1,410 m elevation and
southeast of Cerro Tacarcuna and the headwaters
of the Río Pucro (5 Río Pucuro; see Myers
and Lynch, 1997, figs. 1 and 2). It lies at the
southeastern end of the Serranía del Darién, a
ridge separating the Pacific-draining Río Tuira
system from the Atlantic lowlands of eastern
Panama and northern Colombia (the international
border follows the continental divide along this
ridge). Myers (1969: 24–25) described the
general topography and environment. Additional
perspectives and details are in Anthony (1916,
1923), Gentry (1983), and Myers and Lynch
(1997).
El Copé, continental divide north of, 600–700 m
(Coclé). The village of El Copé is at 08u379N,
80u359W (Fairchild and Handley, 1966). El Copé
(Omar Torrijos) National Park now encompasses
the continental divide.
Isla Popa, south end of, 1 km E of Sumwood
Channel (Bocas del Toro). Approximately
09u099N, 82u089W.
Laguna, 820 m (Darién). 08u049N, 77u199W (Fairchild
and Handley, 1966). According to Charles
W. Myers (personal communication), this locality
is on a ‘‘ridge south of the Río Tacarcuna (upper
tributary of Río Pucuro, Tuira drainage).’’ Also
referred to as ‘‘La Laguna,’’ it is not the village of
the same name near the coast in southwestern
Darién Province (e.g., as indexed in the NGA),
with which it is sometimes confused.
La Loma, W Panama (Bocas del Toro). 08u509N,
82u129W. Also known as Buenavista. 1,200 ft.
elevation (366 m) on the Atlantic slope along a
trail from Chiriquí Lagoon to David (Dunn, 1942:
478).
Peninsula Valiente, Bluefields, 70 m (Bocas del
Toro). 09u119N, 81u559W. Peninsula Valiente
delimits the eastern side of the Laguna de
Chiriquí.
Peninsula Valiente, Quebrada Hido (Bocas del
Toro). Not located.
Pequení–Esperanza ridge, near head of Río Pequení,
2,000 ft. [610 m] (Panamá). Approximately
09u299N, 79u249W. Now within Parque Nacional
Chagres. A ridge northeast of Lago Alajuela
(Madden Lake), running roughly northeast to
southwest and separating the Chagres–Esperanza
river system from the Pequení–Boquerón
system (Dunn and Bailey, 1939: 4, 15). Cadle
and Myers (2003: 15–17) described some of the
geography of the area as altered by damming
associated with construction of the Panama
Canal. Ibáñez et al. (‘‘1994’’ [1995], fig. 1)
provided an outline map of the major rivers of
the region.
Pequení–Esperanza ridge, junction main divide,
1,200 ft. [366 m] (Panamá). Approximately
09u209N, 79u209W. Location not precisely indicated
by Dunn and Bailey (1939) but presumably
at a lower elevation along the ridge closer to Lago
Alajuela than the above-listed locality near the
head of the Río Pequení (see above cited
references).
Summit site, border of Darién, 320 m (San Blas).
08u559N, 77u519W (coordinates from the collector
of FMNH 170138, Michael Duever). Also known
as Camp Summit, a canal survey camp in the
Serranía del Darién discussed by Myers (1969:
26–27, fig. 1), who gave the elevation as 358 m.
LITERATURE CITED
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Segunda edition. Tuxtla Gutierrez, Chiapas, Mexico:
Gobierno del Estado de Chiapas.
ALVAREZ DEL TORO, M. 1982. Los Reptiles de Chiapas.
Tercera edition. Tuxtla Gutierrez, Chiapas, Mexico:
Instituto de Historia Natural.
ANTHONY, H. E. 1916. Panama mammals collected in
1914–1915. Bulletin of the American Museum of
Natural History 35: 357–376 + foldout map.
ANTHONY, H. E. 1923. In the footsteps of Balboa.
Natural History 23: 312–324.
AUTH, D. L. 1994. Checklist and bibliography of the
amphibians and reptiles of Panama. Smithsonian
Herpetological Information Service 98: 1–59.
BARBOUR, T.,AND A. LOVERIDGE. 1946. First supplement
to typical reptiles and amphibians. Bulletin of the
Museum of Comparative Zoology 96: 59–214.
BELT, T. 1874. The Naturalist in Nicaragua, a Narrative
of a Residence at the Gold Mines of Chontales;
Journeys in the Savannahs and Forests; with
Observations on Animals and Plants in Reference
to the Theory of Evolution of Living Forms.
London: John Murray.
BOLAÑOS, F., J. M. SAVAGE, AND G. CHÁVES. 2010.
Anfibios y reptiles de Costa Rica (versión: 18 de
Agosto del 2010). Listas Zoológicas Actualizadas
UCR. San Pedro: Museo de Zoología, Universidad
de Costa Rica. Available from: http://museo.biologia.ucr.ac.cr/
Listas/LZAPublicaciones.htm. Accessed
1 January 2011.
BOULENGER, G. A. 1894. Catalogue of the snakes in the
British Museum (Natural History). Volume 2.
London: British Museum (Natural History).
CADLE, J. E. 2005. Systematics of snakes of the Dipsas
oreas complex (Colubridae: Dipsadinae) in western
Ecuador and Peru, with revalidation of D. elegans
(Boulenger) and D. ellipsifera (Boulenger). Bulletin
of the Museum of Comparative Zoology 158:
67–136.
CADLE, J. E. 2007. The snake genus Sibynomorphus
(Colubridae: Dipsadinae: Dipsadini) in Peru and
Ecuador, with comments on the systematics of
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