Thelypteridaceae (PDF)

THELYPTERIDACEAE 36721. Pseudocyclosorus esquirolii (Christ) Ching, Acta Phytotax.Sin. 8: 324. 1963.西 南 假 毛 蕨 xi nan jia mao jueDryopteris esquirolii Christ, Bull. Acad. Int. Géogr. Bot.17: 144. 1907; Cyclosorus esquirolii (Christ) C. M. Kuo; D.eberhardtii Christ; Thelypteris esquirolii (Christ) Ching.Plants to 1.5 m tall. Rhizomes creeping. Fronds remote;stipes deep stramineous, glabrous above bases. Laminaebroadly oblong-lanceolate, ca. 130 × 30 cm, tapering to bases,pinnate-pinnatifid, pinnatifid-acuminate at apices; pinnae proximal9–11 pairs alternate, gradually reduced to triangular auricles,distal pairs alternate, sessile, spreading, lanceolate, 15–20 × 2–2.3 cm, bases rounded-truncate, symmetrical, pinnatifidnearly to costae, apices long caudate-acuminate; segments 30–35 pairs, spreading, lanceolate, 9–10 × 2.5–3 mm, entire, obtuseor acute at apices; proximal pair (particularly acroscopic one)clearly elongated. Veins visible, costae raised on both sides,8–12 pairs of veinlets per segment, proximal pair arising frombase of costules, acroscopic veinlet reaching bottom of sinus,basiscopic one reaching margin above sinus. Laminae darkgreen when dry, papery, glabrous on both surfaces of intercostalareas, abaxially acicular hairy along rachises and costae, adaxiallydensely appressed-setaceous along costal grooves, with 1or 2 setae on veins and margins. Sori orbicular, attached onmiddle of veinlets, 10–12 pairs per segment; indusia orbicularreniform,thickly membranous, brown, glabrous, persistent.On rocks by streams in valleys, in bamboo by streams; 400–2100m. Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan[N India, Myanmar, Nepal, Thailand].“Christella esquirolii” (H. Léveillé, Fl. Kouy-Tchéou, 474. 1915,nom. nud.) belongs here.Thelypteris ×erubesquirolica W. C. Shieh & J. L. Tsai (J. Sci. Engin.24: 7. 1987), described from Taiwan (Nantou), appears to be a presumedhybrid between Pseudocyclosorus esquirolii and Glaphyropteridopsiserubescens.22. Pseudocyclosorus linearis Ching & K. H. Shing ex Y. X.Lin, Fl. Reipubl. Popularis Sin. 4(1): 328. 1999.线 羽 假 毛 蕨 xian yu jia mao juePlants more than 1.6 m tall. Rhizomes not seen. Stipes ca.30 cm, bases dark brown, distally stramineous, glabrous; laminaelanceolate, ca. 140 × 40 cm, pinnate-pinnatifid, pinnatifidacuminateat apices; proximal 9 pairs of pinnae abruptlyreduced into auricles, middle normal pinnae more than 30 pairs,alternate, obliquely spreading, sessile, linear-lanceolate, ca. 20× 2.3 cm, not broadened at bases, pinnatifid, pinnatifid-acuminateat apices; segments ca. 40 pairs, ligulate, 9–11 × ca. 5mm, acute at apices. Veins not evident, ca. 12 pairs of veinletsper segment, proximal pair reaching bottom of sinus. Laminaedark green when dry, herbaceous; abaxially setaceous alongrachises, glabrous along costae, veins densely glandular, adaxiallydensely setaceous along costal grooves, with 1 or 2 setaeon veinlets, with few fine hairs along costules. Sori orbicular,attached on middle of veinlets; indusia orbicular-reniform,brown, thick, persistent, glabrous.● Mixed forests by streams; ca. 900 m. Sichuan (Emei Shan).Pseudocyclosorus linearis is similar to P. esquirolii but differs inthe pinnae linear-lanceolate and the proximal pair of veinlets on segmentsreaching bottom of sinus.23. Pseudocyclosorus submarginalis Ching ex Y. X. Lin, Fl.Reipubl. Popularis Sin. 4(1): 328. 1999.边 囊 假 毛 蕨 bian nang jia mao juePlants to 75 cm tall. Rhizomes creeping, woody, firm.Fronds remote; stipes ca. 23 cm, deep stramineous, bases withsparse deep brown lanceolate scales, distally glabrous; laminaebroadly lanceolate, ca. 50 × 25 cm, abruptly narrowed proximally,pinnate-pinnatifid, pinnatifid-acuminate at apices; proximal3 pairs of pinnae reduced and hastate; middle normal pinnaeca. 21 pairs, alternate, spreading, sessile, 12–14 × 1.5–1.7cm, bases nearly tapering, truncate, pinnatifid nearly to costae,apices pinnatifid and long acuminate; segments ca. 22 pairs,obliquely spreading, lanceolate, ca. 7 × 4 mm, entire, roundedobtuseat apices. Veins raised on both sides, 7–9 pairs of veinletsper segment, proximal pair arising above base of costules,acroscopic veinlet reaching bottom of sinus, basiscopic onereaching margin above sinus. Laminae greenish when dry, thinlypapery; abaxially sparsely long acicular hairy along rachises,glabrous along costae, sparsely finely hairy and glandularalong veins, adaxially with dense appressed setae along costalgrooves, sparsely setaceous along veinlets. Sori orbicular,attached above middle of veinlets or closer to margins; indusiaorbicular-reniform, brown, glabrous, persistent.● Sichuan.Pseudocyclosorus submarginalis is similar to P. esquirolii but differsin the plants smaller, laminae ca. 50 cm, proximal pair of veinletson segment arising above base of costules, and sori attached abovemiddle of veinlets and close to margins.24. Pseudocyclosorus qingchengensis Y. X. Lin, Fl. Reipubl.Popularis Sin. 4(1): 329. 1999.青 城 假 毛 蕨 qing cheng jia mao juePlants to 1.4 m tall. Rhizomes creeping, woody, firm, darkbrown, sparsely scaly; scales lanceolate, deep brown, fragileand easily deciduous. Fronds remote; stipes 32–37 cm, basesdark brown, distally stramineous, glabrous; laminae lanceolate,ca. 100 × 30 cm, abruptly narrowed at bases, pinnate-pinnatifid,pinnatifid-acuminate at apices; proximal 3–6 pairs of pinnaereduced and hastate; middle normal pinnae ca. 30 pairs, proximalones subopposite, distal ones alternate, sessile, narrowlylanceolate, 12–19 × ca. 2.5 cm, truncate at bases, pinnatifid,pinnatifid-acuminate at apices; segments ca. 37 pairs, 8–10 ×ca. 3 mm, entire, obtuse-pointed at apices. Veins evident, costulesraised on both sides, ca. 12 pairs of veinlets per segment,proximal pair arising above base of costules, acroscopic veinletreaching bottom of sinus, basiscopic one reaching margin abovesinus. Laminae dark green when dry, papery; abaxially withlong acicular hairs along rachises, sparsely shortly hairy alongcostae and veins, densely shortly hairy on intercostal areas,adaxially with dense appressed setae along costal grooves, with1 or more setae along veins and glabrous on intercostal areas.Sori orbicular, attached on middle of veinlets; indusia orbicularreniform,brown, thick, glabrous, persistent.