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Anna Saghatelyan - Botanical Research Institute of Texas

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410 Journal <strong>of</strong> the <strong>Botanical</strong> <strong>Research</strong> <strong>Institute</strong> <strong>of</strong> <strong>Texas</strong> 3(1)<br />

canadensis). Western American (Cordilleran) genera have six species (Muhlenbergia rigida, Epilobiium<br />

ciliatum), while 5 species <strong>of</strong> Amaranthus and Heliotropium are mostly Caribbean.<br />

4a. american trop/Subtr: widely distributed in tropical and subtropical (warm temperate) parts <strong>of</strong><br />

the Americas. This sub-element has 30 <strong>of</strong> the 101 sp. <strong>of</strong> the American element with the grasses<br />

(Cenchrus myosuroides) being especially numerous. Other examples are Tillandsia recurvata, Solanum<br />

elaeagnifolium, Phyla nodiflora, and Ipomoea cardiophylla.<br />

5. North american: more or less wide ranges in temperate regions <strong>of</strong> North America; 98 mesophytic species.<br />

They either belong to widely north temperate genera (Maianthemum racemosum, Carex hystericina, and<br />

Vicia americana), north American genera (Solidago gigantea, Lobelia cardinalis, Monarda fistulosa), or cosmopolitan<br />

genera with large sections in temperate latitudes (Euphorbia cyatophora). Herbaceous habit,<br />

especially in the grasses (18 sp.), sedges (8 sp.), Asteraceae, and Euphorbiaceae (6 sp. each) predominate.<br />

A few older montane woody species have interesting ranges: either absent in the south-eastern and<br />

south-central region (Prunus virginiana), poorly represented in the western and absent in the central<br />

(Cephalanthus occidentalis) region, or those that tend to the Rocky Mountain (Cheilanthes feei) Region.<br />

6. East North american: wide ranges in the Atlantic North American Region <strong>of</strong> Cronquist (1982). Representatives<br />

<strong>of</strong> ancient Laurasian genera are essential among the 37 sp. <strong>of</strong> this geoelement (Ostrya virginiana,<br />

Carya ilinoensis, and Clematis pitcheri). A few species however have tropical connections (Cocculus<br />

carolinus, Celastrus scandex, and Nothoscordum bivalve).<br />

6a. Gulf Coast-(Caribbean): with ranges in the namesake provinces (Thelipteris ovata, Melothria pendula).<br />

6b. appalachian: with ranges in the namesake province (Ostrya virginiana, Melica nitens).<br />

7. Comanchian/ SC US: Edwards Plateau westward to the Trans-Pecos extending northward into central<br />

Oklahoma or further to Ozark Plateau, southward into northeastern Mexico and western Lousiana,<br />

mainly in the limits <strong>of</strong> the Comanchian Subprovince <strong>of</strong> McLaughlin (2007). There are 46 species in<br />

this geoelement like Juniperus ashei, Lupinus texensis, and Scutellaria wrightii, with some having disjunct<br />

relictual ranges (Leptopus phyllanthoides).<br />

8. Prairie: wide ranges in the North American Prairies Province <strong>of</strong> Cronquist (1982); 85 mostly herbaceous<br />

species. Biggest group <strong>of</strong> 24 sp. is in Asteraceae (Liatris punctata, Hymenopappus scabiosaceous), especially<br />

in the Heliantheae s.l. (Panero & Funk 2002). Poaceae (Bouteloua dactyloides, Schedonnardus paniculatus),<br />

and papilionaceous Fabaceae (Astragalus lotiflorus, Dalea aurea) have 7 sp. each. Remaining species<br />

are in large temperate genera (Oenothera triloba, Salvia azurea) with big centers <strong>of</strong> diversity in western<br />

America and Mexico.<br />

8a. S Prairie/SC North american: from Central and South <strong>Texas</strong> extending northward to adjacent<br />

states and southward to northern Mexican Plateau (Eryngium leavenworthii, Argythamnia humilis,<br />

Rhus lanceolata, and Gaillardia suavis).<br />

9. W North american: wide ranges in the entire western North America or its parts mostly north <strong>of</strong> Mexico.<br />

Among the 133 sp. <strong>of</strong> this geoelement 28 sp. <strong>of</strong> Asteraceae (Brickelia californica, Erigeron divergens)<br />

prevail, followed by 18 sp. <strong>of</strong> Poaceae (Agrostis exarata) and 9 sp. <strong>of</strong> Brassicaceae (Stanleya pinnata).<br />

Western American genera have numerous species (Glossopetalon spinescens, Cryptantha cinerea) including<br />

the dominants in different types <strong>of</strong> communities. Examples range from those <strong>of</strong> montane forests (Pinus<br />

ponderosa, Quercus gambelii, and Cercocarpus montanus), Rocky Mountain and Madrean woodlands and<br />

shrublands (Juniperus scopulorum, Rhus trilobata, Holodiscus dumosus), to widely distributed xerophytic<br />

(Opuntia polyacantha) as well as riparian (Salix exigua) species.<br />

9a. WC US/ WC North american: includes western mountainous (not Pacific) region and western<br />

half <strong>of</strong> the Prairies Province in the United States/or southward to central Mexico; this element<br />

supports S. McLaughlin’s (2007) Western Region. Of the 133 sp. <strong>of</strong> W North American element,<br />

29 sp. are in this sub-element. They belong to large temperate genera (Astragalus, Cirsium, and<br />

Lithospermum), North American genera (Monarda pectinata, Oenothera albicaulis), or a few tropical<br />

(Heliotropium convolvulaceum) genera.

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