POACEAE 609composed of 3–5(–)8 racemes, subdigitate; racemes 3–8 cm,tinged purplish; rachis internodes and pedicels ciliate with longsilky hairs. Sessile spikelet 3–4.5 mm; lower glume narrowlyelliptic, cartilaginous, back concave, 5–7-veined, glossy, sparselyhirtellous to silky-pilose below middle, a circular pit abovehairs, 2-keeled, margins keeled and scabrid near apex; awn ofupper lemma 1–2 cm. Pedicelled spikelet male or barren, purplish,subequal to sessile spikelet, glabrous. Fl. and fr. Jul–Oct.2n = 40, 60.Grassy hills, disturbed ground; 1200–1500 m. Guangdong, Sichuan,Yunnan [India, Indonesia, Malaysia, Nepal, Pakistan, Thailand,Vietnam; introduced in Australia and United States].The type of Bothriochloa nana is a stunted specimen of B. pertusawith the spikelets infected by a smut fungus.204. SEHIMA Forsskål, Fl. Aegypt.-Arab. 178. 1775.沟 颖 草 属 gou ying cao shuSun Bixing ( 孙 必 兴 Sun Bi-sin); Sylvia M. PhillipsPerennial or annual. Culms tufted, simple or sparingly branched. Leaf blades narrowly linear; ligule a line of hairs. Inflorescencea single terminal raceme, spikelets paired, dissimilar; rachis internodes and pedicels subinflated, stoutly linear to subclavate,densely white-ciliate along margins. Sessile spikelet bisexual, narrow, compressed between internode and pedicel; callus rounded, insertedinto shallowly hollowed internode apex; lower glume leathery, back concave or longitudinally grooved, strongly veined oneither side of groove but midvein absent, 2-keeled, keels lateral or becoming dorsal toward base, barely winged, apex elongate, scarious,2-toothed; upper glume boat-shaped, finely awned; lower floret staminate, well developed with palea; upper lemma 2-lobed,awned from sinus; awn geniculate, column glabrous or ciliolate. Pedicelled spikelet large, conspicuous, usually staminate, lanceolate,strongly dorsally compressed, distinctly veined, midvein present, awnless. x = 10 and 20.Five species: E Africa through India to SE Asia and Australia; one species in China.1. Sehima nervosum (Rottler) Stapf in Prain, Fl. Trop. Africa9: 36. 1917.沟 颖 草 gou ying caoAndropogon nervosus Rottler, Ges. Naturf. Freunde BerlinNeue Schriften 4: 218. 1803 [“nervosum”]; Ischaemum laxumR. Brown; I. nervosum (Rottler) Thwaites.Perennial. Culms erect, wiry, 30–100 cm tall, nodesbearded or glabrous. Leaf sheaths glabrous or hispid withtubercle-based hairs; leaf blades linear, flat, tough, pale green,10–45 × 0.2–0.7 cm, scabrid, tapering to a filiform apex; ligule2–3 mm. Raceme 3–12(–18) cm, straight or gently curved;rachis internodes and pedicels stoutly linear, 3.5–5 mm. Sessilespikelet yellowish green, 7–9 mm; lower glume narrowlyoblong, deeply grooved between keels in lower part, with 6prominent laterally placed intercarinal veins, inner veinsanastomosing toward apex, apex scarious, 1/4–1/3 glumelength, shortly 2-toothed; upper glume with straight, 7–13 mmawn; awn of upper lemma stout, 1.7–3.5 cm, column brown,ciliolate along spiral, limb pallid. Pedicelled spikelet usuallyflushed purple, 7–10 mm, lower glume conspicuously 7-veined,margins ciliate. Fl. and fr. Jul–Oct.Dry grasslands; at low elevations. Hainan, Yunnan [India, Indonesia,Laos, Myanmar, New Guinea, Pakistan, Philippines, Sri Lanka,Thailand, Vietnam; E Africa, SW Asia, Australia].205. ISCHAEMUM Linnaeus, Sp. Pl. 2: 1049. 1753.鸭 嘴 草 属 ya zui cao shuSun Bixing ( 孙 必 兴 Sun Bi-sin); Sylvia M. PhillipsPerennial, or sometimes annual. Culms often decumbent and much branched. Leaf blades linear to lanceolate, narrowed toligule, sometimes pseudopetiolate; ligule membranous; sheath auricles often present. Inflorescence of paired racemes, or occasionallysubdigitate, terminal and axillary, exserted or sometimes supported by a spatheole; racemes 1-sided, when paired oftenlocked back to back and appearing as a single cylindrical raceme, spikelets of a pair similar or not; rachis internodes and pedicelsstoutly linear to thickly clavate, U-shaped or triquetrous in cross section. Sessile spikelet dorsally compressed; callus truncate or obtuse,inserted into hollowed internode apex; lower glume papery to leathery, shallowly convex or almost flat, 2-keeled or rounded onflanks, sometimes winged, often rugose; upper glume boat-shaped, awnless or with straight awn from apex; lower floret male, welldeveloped with palea; upper lemma usually deeply 2-lobed, awned from sinus, rarely awnless; awn geniculate, glabrous. Pedicelledspikelet as large as sessile or much smaller, upper lemma sometimes geniculately awned.About 70 species: throughout the tropics but mainly in Asia, especially India; 12 species (one endemic) in China.1a. Margins of lower glume of sessile spikelet broadly rounded in lower part, narrowly inflexed and keeled above, notcoarsely rugose or knobbly.2a. Spikelets of a pair without obvious awns, or only sessile spikelet awned.3a. Inflorescence base enclosed in uppermost sheath; plant strongly rhizomatous ................................................ 1. I. muticum3b. Inflorescence long exserted from uppermost sheath; plant loosely tufted.
610POACEAE4a. Racemes villous; sessile spikelet 8–10 mm ..................................................................................... 2. I. anthephoroides4b. Racemes glabrous; sessile spikelet 5.5–8 mm .......................................................................................... 3. I. aristatum2b. Spikelets of a pair both clearly awned.5a. Lower glume of sessile spikelet winged on keels, apex rounded, emarginate.6a. Racemes paired; rhizomes absent; culms less than 60 cm ............................................................................ 4. I. ciliare6b. Racemes (2–)3–6; rhizomes present; culms 60–100 cm .................................................................... 5. I. polystachyum5b. Lower glume of sessile spikelet not winged on keels, apex sharply bicuspidate.7a. Spikelets ± glabrous; upper glume of sessile spikelet not winged on keel; callus hairs less than1 mm .......................................................................................................................................................... 6. I. timorense7b. Spikelets hispid; upper glume of sessile spikelet winged on keel; callus hairs ca. 2 mm ................ 7 I. thomsonianum1b. Margins of lower glume of sessile spikelet narrowly and evenly inflexed and keeled along entire length, oftencoarsely rugose or knobbly.8a. Lower glume of sessile spikelet not transversely rugose or knobbly.9a. Lower glume of sessile spikelet wingless, keels pilose; awn 1.8–2.5 cm ........................................................... 8. I. aureum9b. Lower glume of sessile spikelet winged, glabrous; awn 1–1.2 cm .................................................................. 9. I. setaceum8b. Lower glume of sessile spikelet transversely rugose or knobbly on flanks.10a. Plant annual; lower glume of sessile spikelet coarsely rugose with 4–7 sharp transverse ridges acrossback ................................................................................................................................................................ 10. I. rugosum10b. Plant perennial; lower glume of sessile spikelet with rounded knobs on keels in lower part, sometimesalso with 2–4 weak ridges across back.11a. Culms up to 1 m tall; sessile spikelet 5–7 mm, awned, awn 1–1.5 mm ........................................... 11. I. barbatum11b. Culms 1–2 m tall; sessile spikelet 6.5–8 mm, awnless or awn less than 1 mm .................................. 12. I. magnum1. Ischaemum muticum Linnaeus, Sp. Pl. 2: 1049. 1753.无 芒 鸭 嘴 草 wu mang ya zui caoPerennial, strongly rhizomatous; rhizomes clothed in cataphylls.Culms often red, much branched, stoloniferous orscrambling, several meters long, flowering culms erect, up to60 cm, nodes glabrous. Leaf sheaths ciliate along outer margin,otherwise glabrous or sparingly appressed hairy; leaf bladeslanceolate, tinged reddish brown, 2–10(–18) × 0.3–1.7 cm, glabrousor abaxial surface sparingly pilose, margins smooth orscaberulous, base cordate, very shortly pseudopetiolate, apexacute; ligule 0.2–0.6 mm. Racemes usually paired, appressedback to back, 2–5 cm, base enclosed by subtending sheath;rachis internodes and pedicels oblong, triquetrous, outer anglenarrowly winged, inner angles glabrous or ciliolate. Sessilespikelet lanceolate, 4.8–7 × 2.5–2.8 mm; lower glume leatherywith expanded rounded flanks in lower 2/3, herbaceous, stronglyveined and sharply 2-keeled below apex, glabrous, wingedfrom near base, apex entire; upper glume winged on upper keel;upper lemma subentire, mucronate or with ca. 1 mm awnlet.Pedicelled spikelet laterally compressed, otherwise resemblingsessile or smaller, awnless.Sands near the sea; below 100 m. Taiwan [Cambodia, India, Indonesia,Japan (S Ryukyu Islands), Malaysia, Myanmar, New Guinea,Philippines, Sri Lanka, Thailand, Vietnam; Australia (Queensland)].This seashore plant forms extensive colonies that bind the sand atthe back of sandy beaches.2. Ischaemum anthephoroides (Steudel) Miquel, Ann. Mus.Bot. Lugduno-Batavi 3: 193. 1867 [“antephoroides”].毛 鸭 嘴 草 mao ya zui caoRottboellia anthephoroides Steudel, Flora 29: 22. 1846[“antephoroides”]; Andropogon anthephoroides (Steudel) Steudel;A. anthephoroides var. eriostachyus (Hackel) Honda; Ischaemumeriostachyum Hackel.Perennial. Culms loosely tufted, geniculately ascending toshortly decumbent and rooting at lowest nodes, 30–70 cm tall,branching in lower part, nodes bearded. Leaf sheaths loose, villous;leaf blades linear-lanceolate, 5–20 × 0.3–0.9 cm, villous,margins thickened, smooth, base constricted, apex acuminate;ligule 1–3 mm. Racemes terminal, paired, appressed back toback, 5–8 cm, long exserted; rachis internodes and pedicelsstoutly cuneate, triquetrous, villous. Sessile spikelet broadlyoblong or obovate, 8–10 × 2.5–3.5 mm; lower glume leatherytoward base, flanks rounded in lower 1/3, villous with stiff ca. 3mm hairs, papyraceous and puberulous above, keeled andwinged above middle, wings ca. 0.8 mm wide with ciliolatemargin, apex truncate-denticulate; upper glume keeled andwinged above middle, villous on midline below wing; awn ofupper lemma weakly geniculate, up to 1.5 cm. Pedicelledspikelet laterally compressed, otherwise resembling sessile,awnless. Fl. and fr. Jun–Sep.Sand dunes, sandy slopes, near the sea. Hebei, Shandong, Zhejiang(Dachen Dao) [Japan, Korea].This species occupies a similar seashore habitat to Ischaemummuticum, but has a much more limited, more northerly distribution. It isa much hairier species, with exserted racemes of larger spikelets, andlacks the widely spreading, scaly rhizomes of I. muticum.3. Ischaemum aristatum Linnaeus, Sp. Pl. 2: 1049. 1753.有 芒 鸭 嘴 草 you mang ya zui caoPerennial. Culms loosely tufted, erect or geniculately ascending,40–80 cm tall, simple or branching, nodes glabrous.Leaf sheaths glabrous or pilose; leaf blades linear-lanceolate, 5–25 × 0.4–1 cm, glabrous or thinly pilose, margins smooth becomingscabrid toward apex, base attenuate or contracted, apexacuminate; ligule 2–3 mm. Racemes terminal, paired, appressedback to back, 4–7 cm; rachis internodes clavate, triquetrous,scabrid or ciliate along outer angle, inner angles glabrous orshortly ciliate. Sessile spikelet oblanceolate to obovate, 5.5–8 ×2–2.3 mm; lower glume leathery with rounded flanks below
- Page 1 and 2: 570POACEAEsmooth or scaberulous. Sp
- Page 3 and 4: 572POACEAE10a. Raceme rachis tough,
- Page 5 and 6: 574POACEAE7a. Leaf blades villous;
- Page 7 and 8: 576POACEAEGansu, Guangdong, Guizhou
- Page 9 and 10: 578POACEAE2. Saccharum formosanum (
- Page 11 and 12: 580POACEAEwith long soft hairs on b
- Page 13 and 14: 582POACEAEveined, puberulous; upper
- Page 15 and 16: 584POACEAEImperata conferta (Presl)
- Page 17 and 18: 586POACEAE10b. Leaf blades green or
- Page 19 and 20: 588POACEAEAndropogon leschenaultian
- Page 21 and 22: 590POACEAEapex mucronate or awned;
- Page 23 and 24: 592POACEAE2. Pogonatherum paniceum
- Page 25 and 26: 594POACEAEThe name Microstegium gla
- Page 27 and 28: 596POACEAE6. Microstegium petiolare
- Page 29 and 30: 598POACEAENormally the lower floret
- Page 31 and 32: 600POACEAEous, ca. 1 mm. Raceme 1(-
- Page 33 and 34: 602POACEAEgreen to dark brown or bl
- Page 35 and 36: 604POACEAEMountain slopes; ca. 2500
- Page 37 and 38: 606POACEAE1b. Sessile spikelet 3-3.
- Page 39: 608POACEAEbranched peduncles; racem
- Page 43 and 44: 612POACEAEMoist places on grassy pl
- Page 45 and 46: 614POACEAE206. APLUDA Linnaeus, Sp.
- Page 47 and 48: 616POACEAE1a. Raceme rachis scabrid
- Page 49 and 50: 618POACEAEtinate-setose from stout
- Page 51 and 52: 620POACEAEtatherus Hackel; A. cilia
- Page 53 and 54: 622POACEAE1. Schizachyrium sanguine
- Page 55 and 56: 624POACEAE211. CYMBOPOGON Sprengel,
- Page 57 and 58: 626POACEAECulms 20-40 cm tall; old
- Page 59 and 60: 628POACEAEgrown on a large scale in
- Page 61 and 62: 630POACEAEdense; spatheoles pale re
- Page 63 and 64: 632POACEAE1b. Racemes reflexed at m
- Page 65 and 66: 634POACEAE7b. Sessile spikelet with
- Page 67 and 68: 636POACEAEdifferent levels, male or
- Page 69 and 70: 638POACEAEnius; H. polystictus (Ste
- Page 71 and 72: 640POACEAE(April 1889), as was ackn
- Page 73 and 74: 642POACEAEglabrous. Leaf sheaths lo
- Page 75 and 76: 644POACEAEThaumastochloa chenii Hsu
- Page 77 and 78: 646POACEAErooting and branching, fl
- Page 79 and 80: 648POACEAEper glume equal to lower
- Page 81 and 82: 650POACEAEslightly inflated, sparse