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Delsate, Steur, Schneider, Thuy Lower to Middle Emsian flora and fauna from Consthum and Merkholtz 14 Superorder Isofilibranchia Iredale, 1939 vel Subclass Heteroconchia Hertwig, 1895 Superorder Palaeoheterodonta Newell, 1965 Order Modiomorphoida Newell, 1969 Family Modiomorphidae Miller, 1877 (= Modiolopsidae Fischer, 1887) Genus Goniophora Phillips, 1848 Goniophora sp Pl. 7, fig. 4 EIB119-120 (5-6 cm long axis) is the external surface mould of a Goniophora sp. Other Goniophora specimens (Plate 7, fig. 4) and another member of the Modiomorphidae, possibly Modiolopsis sp., are present on slab EIB131, with an interesting undetermined bivalve, byssally aached to plant material and needing further study. The pluricentimetric Goniophora are frequent in the upper level of the quarry. Modiomorphidae were abundant in Early Devonian high energy shallow water environments, but also in other facies. Subclass Heteroconchia Hertwig, 1895 Superorder Heterodonta Neumayr, 1883 Order Veneroida Adams and Adams, 1856 Superfamily Crassatelloidea Ferussac, 1822 Family Cardiniidae Ziel, 1881 Genus Cypricardinia Hall, 1859 Cypricardinia sp. The pericentimetric EIB121 and 2 specimens on slab EIB123 are probably Cypricardinia sp., but the hinge would be necessary for further determination. Note that Cox et al. 1969 include Cypricardinia in the Cardiniidae, but that Skelton and Benton 1993 exclude them from the family. This small bivalve was supposed to live in the sediment (= semi-infaunal). Subclass Pteriomorphia Beurlen, 1944 Superorder Eupteriomorphia Boss, 1982 Order Pterioida Newell, 1965 Superfamily Pterioidea Gray, 1847 Genus Leptodesma Hall, 1883 Leptodesma concentrica (Roemer) EIB122 (3 cm long axis) is most probably Leptodesma concentrica (Roemer) which is supposed to be a synonym of Leptodesma pseudolaevis (Oehlert). Superorder Prionodonta McNeill, 1937 Order Arcoida Stoliczka, 1871 Superfamily Arcoidea Lamarck, 1809 Family Parallelodontidae Dall, 1898 Genus Parallelodon Meek & Worthen, 1866 Parallelodon sp. The pericentimetric EIB124 and another specimen on slab EIB123 are probably Parallelodon sp. based on the relatively broad umbo placed in the anteriormost third of the straight hinge line. The pericentimetric Parallelodon and Cypricardinia (probably 2 genera oen identified as Grammysia) are frequent in many levels of the Consthum quarry. Arthropoda Rare Emsian Arthropods have been intensively studied by Stormer 1969, Heer 1972, Dunlop & Poschmann, 1997, Anderson et al.1998 and Dunlop et al. 2001, who described scorpios and arachnids, synziphosurine and chasmataspidida chelicerates, mostly from the Rheinisches Schiefergebirge. But sea scorpions (Eurypterids) remains are much more common in the Lower Devonian (see Geys 1991). Crustacea Class Ostracoda Latreille, 1802 Some badly preserved specimens show some Leperditellidae affinities. Phylum Chelicerata Heymons, 1901 Class Eurypterida Burmeister, 1843 Suborder Pterygotina Caster & Kjellesvig- Waering, 1964 Superfamily Pterygotoidea Clarke & Ruedemann, 1912 Ferrantia • 36 / 2003
Delsate, Steur, Schneider, Thuy Lower to Middle Emsian flora and fauna from Consthum and Merkholtz Ferrantia • 36 / 2003 Family Pterygotidae indet. Centimetric plates with tubercular ornamentation are aributable to pterygotids (sea scorpions). Problematica ? Mollusca incertae sedis Class Tentaculitoidea Ljashenko, 1957 = Criconarida Fisher, 1962 Order Tentaculitida Ljashenko, 1955 Tentaculites sp. Plurimillimetric elongated cones with annular ornamentation, Tentaculites are marine animals. Tentaculites was already reported by Asselberghs 1913 from the Schubourg Quartzophyllade. Ichnofossils Punctiform or fine oblique parallel sets of ridges tracks are aributed to trilobite’s walks, Diplichnites, while parallel irregular bars remain indeterminate. Vertebrata Osteichthyes Huxley, 1880 Class Sarcopterygii Romer, 1955 Infraclass Rhipidistia Cope, 1889 Order Holoptychiida Andrews, 1967 Family Porolepididae Berg, 1940 Porolepididae Berg, 1940 Porolepis sp. EIB125: Pl. 5, fig. 1 and 2 a,b Two scales have been recovered; they are perfectly preserved, and were prepared by Marcel Schneider. The beds yielding these scales are situated at the upper level, at the eastern part of the “Rinnen” quarry. The bigger one is 7 mm wide x 12 mm high; two adjacent borders are pectinated by ridges; some ridges bifurcate. The shiny surface (cosmine) is punctuated by pores, also present between the borders ridges. The smaller one, 7 mm x 10 mm, is incomplete, but the punctuations and the sometimes bifurcated ridges of two adjacent borders are present. Discussion Emsian vertebrates are limited to Chondrichthyans, the shield covered Agnathan and Placoderms, the tiny lozenge scaled Acanthodians, and early Actinopterygians. Centimetric scales of the Consthum type are much larger than Actinopterygian post-emsian Stegotrachelid Moythomasia (see for example Long & Trinajstic, 2000) and Palaeoniscoidea Orvikuina (see Bardenheuer, 1997) scales ; moreover these two genera usually have lozenge shaped elongated scales with well marked incisions of the posterior border rather than ridges. Actinopterygian Cheirolepis scales have a different morphology with irregular shaped ganoin surface. An Acanthodian aribution is discarded, as acanthodian scales have a typical structure with a constriction between the base and the outer part of the scale, and are usually much smaller than the Luxembourg specimens described here. The dimensions and the pores of the scale surface lead to a Sarcopterygian Porolepid aribution, thus the pectinated border is anterior (see Gross 1966). Lower Devonian porolepids have been described by Gross 1933a (Pl. 5:15) and Jessen 1965, from Germany (Rheinisches Schiefergebirge : Overath : Upper Siegenian; Ahrtal in Eifel (Ebert), Brexbachtal in Lower Coblencian (Heer)), and Spitsbergen. Lippert 1939 reports Porolepis scales from the Emsian (Clervaux Schists, E2) of Zweifelscheid and Irrhausen (D), near to the Luxembourg border. The first report of the genus dates from the Praguian (Siegenian) of Rhineland (Porolepis siegenensis Gross 1936), Spitsbergen (P. spitsbergensis Jarvik 1937, P. elongata Jarvik 1937, P. brevis Jarvik 1937); the last report is P. posnaniensis Kade 1858 from the Givetian of Estonia (Meseritz) (see Benton 1993). The genus is known from isolated scales and fragmentary bones (mandible: Gross 1933a: Plate 5: 13). Scales reach 30 m x 18 mm. 15
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B. Thuy Les échinides du Bajocien
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Authors notes according to the conc
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20 Les haies au Grand-Duché de Lux
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