the genus Coccocarpia (Peltigerales: Coccocarpiaceae) - Instituto ...

Table 1. Species of Coccocarpia recorded for Costa Rica. Detailed collection data, including color images,
can be retrieved online from The Field Museum's Botany Collections Database at
http://emuweb.fieldmuseum.org/botany/search_crf.php.
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Coccocarpia adnata L. Arvidss.
Coccocarpia dissecta Swinsc. & Krog
Coccocarpia domingensis Vain.
Coccocarpia epiphylla (Fée) Kremp.
Coccocarpia erythroxyli (Spreng.) Swinsc. & Krog
Coccocarpia filiformis L. Arvidss.
Coccocarpia gallaicoi Lücking, Chaves & Umaña sp. nov.
Coccocarpia aff. gallaicoi Lücking, Chaves & Umaña
Coccocarpia glaucina Kremp.
Coccocarpia guimarana (Vain.) Swins. & Krog.
Coccocarpia aff. imbricascens Nyl.
Coccocarpia microphyllina Lücking & Aptroot sp. nov.
Coccocarpia neglecta Aptroot & Lücking sp. nov.
Coccocarpia palmicola (Spreng.) L. Arvidss. & D. Gall.
Coccocarpia pellita (Ach.) Müll. Arg.
Coccocarpia prostrata Lücking, Aptroot & Sipman sp. nov.
Coccocarpia stellata Tuck.
Coccocarpia tenuissima Müll. Arg.
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The morphology and position of isidia and lobules plays an important role in distinguishing
species of Coccocarpia. The distinction of isidia, phyllidia, lobules, and other similar
structures, is not always clearcut in the literature (see discussion in ARVIDSSON 1983: 12),
and we have here used a combination of morphological and functional definitions (Table
4). The term "phyllidia" was originally introduced for flattened isidia, but in our opinion
covers perfectly the marginal isidioid, strongly branched structures found, for example, in
Sticta species (MCDONALD et al. 2003). The structures have the same color as the thallus
(while true isidia in Sticta are usually darker) and do not seem to act as dispersal organs,
but rather increase the thallus surface for enhanced gas exchange. So defined, phyllidia
differ from both flattened isidia (in function) and accessory lobules (in the absence of
rhizines). The new species Coccocarpia microphyllina is the first in the genus in which
such phyllidia are found; all other species either produce isidia in the strict sense or
accessory lobules.
Table 2. Revision of names in Coccocarpia published by DODGE (1933) for Costa Rica and on labels of
specimens in FH, their current nomenclatural status, and taxonomic identification of the material (see also list
of cited specimens under each taxon entry below). *names not published by Dodge (1933) but used on
specimen labels, **superfluous new combinations published by DODGE (1933) of names already recombined
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