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Fungi with septate hyphae and a dikaryophase

Fungi with septate hyphae and a dikaryophase

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5. F\ngt wtth septale haphae a nd a dtkargophase<br />

3. Pseudoparaphyses are <strong>hyphae</strong> found in some<br />

bitunicate Ascomycota that originate from above the<br />

asci <strong>and</strong> grow down between the developing asci <strong>and</strong><br />

become attached to the base of the ascal cavity.<br />

They often become free from the upper part.<br />

Pseudoparaphyses are often regularly <strong>septate</strong>, show<br />

regular anastomosis <strong>and</strong> are often broader than<br />

paraphyses, e.g. Pleospora spp.<br />

4. Paraphysoids resemble pseudoparaphyses but<br />

usually have fewer septa <strong>and</strong> are narrower;<br />

anastomosis common, e.g. Patellaria spp.<br />

5. Perlphysoids are short <strong>hyphae</strong> similar to<br />

pseudoparaphyses but which do not reach the base<br />

of the ascal cavity, e.g. Nectna spp.<br />

6. Periphyses are <strong>hyphae</strong> that form a -fringe on the<br />

inside of the ostiole or pore of ascomata <strong>and</strong> grow<br />

towards the opening. They are <strong>septate</strong>, unbranched<br />

<strong>and</strong> do not anastomose, e.g. Gibberella spp.<br />

7. Hamathecial tissue absent (not figured), e.g.<br />

Dothidea spp.<br />

63<br />

3. @@<br />

The traditional system of classi$ring the Ascomycota is based on the tytrle of<br />

ascoma, the structure of the ascus wall, the method of ascus dehiscence <strong>and</strong> the<br />

method of development of the centrum (the structures <strong>with</strong>in the ascoma such as<br />

the asci <strong>and</strong> inter-ascal tissue). This system, which is now recognised as being<br />

artificial because it does not indicate natural phylogenetic relationships, is<br />

outlined inTable 5.1.<br />

Table 5.1 Key to the classes of Ascomycota.<br />

A" Asci unitunicate or if bitunicate then in the exposed h5rmenium of an apothecium.<br />

B. Asci naked, formed as discrete free cells or in a hymenium of indefinite extent<br />

but not bounded bv a stroma or bv ascoma tissue: asci indehiscent.<br />

Class Hemiascomycetes<br />

B.* Asci formed in ascomata<br />

C. Asci scattered at various levels <strong>with</strong>in a cleistothecium or a beaked<br />

perittrecium; asci indehiscent.<br />

Class Plectomycetes<br />

C.* Asci formed at a common level in a hymenial layer or in a fascicle (a<br />

group) at the base of the ascoma.<br />

D. Ascoma usually a perithecium, less often a cleistothecium <strong>with</strong><br />

basal fasciculate asci or an ascostroma <strong>with</strong> unitunicate asci: asci<br />

inoperculate, <strong>with</strong> an apical pore or slit.<br />

Class P5zrenomycetes<br />

D.* Ascoma an apothecium; asci operculate, inoperculate or<br />

indehiscent.<br />

Class Discomycetes<br />

Ar' Asci bitunicate <strong>and</strong> formed in an ascostroma but not in an apothecium.<br />

Class Loculoascomycetes<br />

5.<br />

6.

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