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Plants of Casa Grande Ruins National Monument - NPS Inventory ...

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Pteridaceae<br />

Ferns<br />

36<br />

Cheilanthes wootonii<br />

beaded lipfern<br />

General: Slender, widely creeping rhizomes<br />

1–3 mm in diameter, densely scaly with loosely<br />

imbricated, oblong–ovate to lance–oblong,<br />

distantly denticulate, light reddish brown scales,<br />

2–3 mm long. Leaves: Several fronds, scattered<br />

7–35 cm long, noncircinate vernation, stipes<br />

slender 5–20 cm long, petiole dark brown,<br />

rounded adaxially; blade oblong–lanceolate,<br />

3–4 pinnate at base, 2–5 cm wide; pinnate not<br />

articulate, basal pair not conspicuously larger<br />

than adjacent pair; scales firmly attached,<br />

rounded to subcordate at base, ciliate. Sporangia: Few, false indusia<br />

marginal, weakly differentiated, 0.05–0.25 mm wide; sori more or less<br />

continuous around segment margins; sporangia containing 32 spores.<br />

Ecology: Found on rocky slopes and along ledges from 3,000–9,500 ft (914–<br />

2896 m); sporulating summer–fall. Notes: This species can be distinguished<br />

by the leaf blades appearing glabrous adaxially, with costal scales that are only<br />

ciliate in the proximal half, and the brown and loosely appressed stem scales.<br />

Ethnobotany: Used as a life medicine and as a lotion for gunshot wounds.<br />

Etymology: Cheilanthes is from Greek cheilos for lip and anthos for flower,<br />

while wootonii is named for Elmer Otis Wooton (1865–1945), an American<br />

botanist and former curator <strong>of</strong> the <strong>National</strong> Herbarium. Synonyms: None<br />

©2004 Patrick Alexander

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