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Animal Diversity: Chordata

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The following alternate classification of amphibians is as given by Parker and Haswell:<br />

Table 2: Classification of amphibia<br />

Subclass 1:<br />

Apsidospondyli<br />

Super order 1:<br />

Labyrinthodontia<br />

Order 1:<br />

Ichthyostegalia<br />

Order 2:<br />

Rhachitomi<br />

Order 3:<br />

Stereospondyli<br />

Order 4:<br />

Embolomeri<br />

Order 5:<br />

Seymouriamorpha<br />

Super order 2:<br />

Salientia<br />

Order 1:<br />

Eoanura<br />

Order 2:<br />

Proanura<br />

Order 3:<br />

Anura<br />

Subclass 2:<br />

Lepospondyli<br />

Order 1:<br />

Aistopoda<br />

Order 2:<br />

Nectridia<br />

Order 3:<br />

Microsauria<br />

Order 4:<br />

Urodela<br />

Order 5:<br />

Apoda<br />

Labyrinthodonts The oldest amphibians were the swamp-dwelling labyrinthodonts.<br />

Ichthyostega was the earliest specimen appearing in the Devonian. Labyrinthodonts were a<br />

large, widely dispersed and diverse assemblage. On the basis of the morphology of their<br />

vertebrae, paleontologists have been of the opinion that fossil amphibians with<br />

stereospondylous and embolomerous vertebrae were not in the amniote line. Labyrinthodonts<br />

had many features seldom seen in modern amphibians. These included minute bony scales in<br />

the skin dermis; a fishlike tail supported by dermal fin rays; and skull similar to those of<br />

rhipidistian fishes. Labyrinthodonts, like their aquatic ancestors, had a sensory canal system<br />

of neuromast organs. One or another of the labyrinthodonts was ancestral to the first amniote.<br />

Temnospondyls was a group that was common in the Permian with its fossil record<br />

extending back to the Mississippian. Members of the temnospondyls have achieved skeletal<br />

similarities to modern frogs and salamanders, suggestive of their close relationship. A<br />

number of lissamphibian skeletal features and their relatively smaller size can be explained as<br />

the retention of juvenile ancestral temnospondyl features. The condition in caecilians does<br />

not fit easily into this scenario, possibly suggesting an independent origin from microsaurs.<br />

6

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