Thursley Common [Shrike Hill] (SU9041) & Sheets Heath (SU9457 ...

Thursley Common [Shrike Hill] 

(SU9041) & Sheets Heath 

(SU9457), 

Surrey [VC17] 

Bare ground invertebrate 

assemblage monitoring 

Written by Scotty Dodd M.Sc. 

January 2013 

Surrey Wildlife Trust, School Lane, Pirbright, Woking, Surrey GU24 0JN

SUMMARY..............................................................................................................................................................................2 

1. INTRODUCTION ...............................................................................................................................................................3 

2. METHODS...........................................................................................................................................................................3 

Table 1. Sampling techniques for invertebrates. A fuller description of standardised methods for each sampling 

protocol is provided in Drake et al. (2007)......................................................................................................................4 

3. RESULTS .............................................................................................................................................................................5 

Table 2: Invertebrate species with UKBAP/SPI, RDB or Nationally Scarce statuses. Nationally Scarce species fall 

into two categories, a & b (Na & Nb). An asterix (*) in the appropriate site column denotes presence at a site............5 

Table 3a: Sheets Heath ISIS (2010) analysis of species samples – Broad Assemblage Types (BATs).............................7 

Table 3b: Sheets Heath ISIS (2010) analysis of species samples – Specific Assemblage Types (SATs). .........................7 

Table 4a: Thursley Common ISIS (2010) analysis of species samples – Broad Assemblage Types (BATs). ...................7 

Table 4b: Thursley Common ISIS (2010) analysis of species samples – Specific Assemblage Types (SATs)..................7 

4. ENTOMOLOGICAL ASSESSMENT AND RECOMMENDATIONS..........................................................................9 

5. REFERENCES ....................................................................................................................................................................9 

6. ACKNOWLEDGEMENTS ..............................................................................................................................................10 

7. APPENDICES....................................................................................................................................................................11 

7.1. APPENDIX 1: STATUS CATEGORIES FOR UNCOMMON SPECIES...........................................................................................11 

7.2. APPENDIX 2: ANNOTATED SPECIES LIST FOR SHEETS HEATH: ..........................................................................................12 

7.3. APPENDIX 3: ANNOTATED SPECIES LIST FOR THURSLEY COMMON: ..................................................................................23 

7.4. APPENDIX 4: INVERTEBRATE SPECIES-HABITAT INFORMATION SYSTEM (ISIS, 2010) ASSEMBLAGE ASSESSMENT.................31 

7.5. APPENDIX 5: MAP OF SAMPLING LOCATIONS AT SHEETS HEATH.......................................................................................32 

7.6. APPENDIX 6: MAP OF SAMPLING LOCATIONS AT THURSLEY COMMON. .............................................................................32 

January 2013 1 C1811 Invertebrate Monitoring Report

Summary 

Each selected study site was visited on four occasions throughout the year (the first visit being a 

brief scoping visit). Surveys were conducted on the following dates: 

Sheets Heath: (1.v.), 25.vi., 5.vii. & 22.viii.2012. 

Thursley Common: (26.iii.), 26.vi., 6.vii. & 23.viii.2012 

The primary objective of this research was to use standardised survey techniques to monitor 

invertebrate colonisation of areas specifically managed for bare ground and early successional 

heathland habitat between 2009 and 2011 as a part of the SITA Trust funded Heath Tiger Beetle 

Project. 

Invertebrates were identified to species level and later assigned to an assemblage type using the 

ISIS programme. Assemblage types generated were then analysed in terms of quality (based upon 

presence of uncommon species) and relevance to the target habitat of bare ground on dry heathland 

(based upon presence of high-fidelity species). 

The total number of invertebrate species recorded from each survey area are as follows; figures in 

parentheses represent species, within the total, afforded published conservation statuses (i.e. UK 

Biodiversity Action Plan, Red Data Book or Nationally Scarce categories A or B) + those species 

regarded as nationally Local: 

Sheets Heath: 128 (17 + 30) 

Thursley Common: 108 (21 + 31) 

Suboptimal weather conditions throughout 2012 affected the results for both sites, with several 

expected species noted in previous years not found. However, both sites were found to be in 

Favourable Condition for the recorded invertebrate assemblages associated with bare sand 

(F111). This strongly suggests that the habitat management undertaken has enhanced opportunities 

for invertebrates associated with dry, bare ground at both study sites. The assemblage representing 

scrub-heath (F003) was also found to be in Favourable Condition. This result was expected 

given that both sites represent well-established heathland. 

Thursley Common, the larger site and a National Nature Reserve, was found to be the superior site 

with an early successional mosaic / bare sand assemblage rarity scoring of 217 (based upon 

presence of uncommon and high-fidelity species). Further analysis of species fidelity to the target 

habitat shows that 56% of the 108 species recorded have a medium (22%) or high (34%) fidelity 

association with bare sand on dry heathland. In comparison Sheets Heath, a much smaller site 

geographically adjacent to, but physically fragmented from, the much larger Pirbright Ranges 

heathland produced an assemblage rarity scoring of 188, with 40% of the 128 species recorded 

having a medium (13%) or high (27%) fidelity association. 


1. Introduction 

The SITA Trust funded Heath Tiger Beetle Project arranged this monitoring work via Surrey Wildlife 

Trust Consultancy. The primary objective of this research was to use standardised survey techniques to 

monitor invertebrate colonisation of areas specifically managed for bare ground and early successional 

heathland habitat between 2009 and 2011 and to provide a baseline list of species with basic ecological 

information. 

This report briefly discusses the invertebrates recorded at the site. Invertebrates were identified to 

species level and later assigned to an assemblage type using the ISIS programme. Assemblage types 

generated were then analysed in terms of quality and relevance to the target habitat, based upon 

presence of uncommon and high-fidelity species. 

2. Methods 

Two sites were selected, Sheets Heath (SU9457) and Thursley Common (SU9041) – see map 

Appendices 5 & 6. Originally it had been intended to survey Hankley Common and Ash Ranges. 

However, due to a combination of access restrictions due to military training and (for Ash Ranges) the 

perceived lack of colonization of scrapes (winter snow during the winter close-down period led to all 

scrapes on Ash Ranges being postponed until the final year of the project, i.e. the winter before 

intended survey) it was decided to survey sites with no access restrictions and a three year succession 

of bare earth scrapes, banks and bunds. 

Sheets Heath sampling areas: 

SU 9457 5731, SU 9458 5734, SU 9464 5736, SU 9461 5765 

Thursley Common sampling areas: 

SU 9078 4106, SU 9068 4108, SU 9064 4103, SU 9072 4100 

Each selected study site was visited on four occasions throughout the year (the first visit being a brief 

scoping visit to ensure habitat suitability). Surveys were conducted on the following dates: 

Sheets Heath: (1.v.), 25.vi., 5.vii. & 22.viii.2012. 

Thursley Common: (26.iii.), 26.vi., 6.vii. & 23.viii.2012 

Standardised and timed entomological sampling techniques were employed (see Table 1), based upon 

those in Drake et al. (2007). The time period protocol for sweep-netting was doubled from that given in 

Drake et al. (2007) in accordance with the latest Natural England ISIS sampling protocol guidelines 

(Heaver, pers. comm. 2012). Samples were taken in representative habitat (i.e. bare sand and sparsely 

vegetated heath) during each visit, focusing upon the turf stripped areas, bare earth scrapes and sandy 

bunds created during 2009 – 2011. Sandy tracks and early successional mosaics adjacent to these were 

incorporated into the sampling areas to ensure adequate spatial coverage. Field observations (e.g. tiger 


eetles and butterflies basking on sandy tracks) were made whilst in transit during sampling events and 

these are therefore also recorded. 

Table 1. Sampling techniques for invertebrates. A fuller description of standardised methods for each 

sampling protocol is provided in Drake et al. (2007). 

Technique Target Groups Target Habitats 

Timed sweep-netting 

(‘sweeping’) with a W&D 

calico-bag sweep-net. 4 x 

20min samples (of 4 x 5min 

sub-samples per sample). 

Timed spot-sweeping 

(‘netting’) with a fine mesh 

net. 4 x 30min samples. 

Each sample covering a 

min of 50 meters of suitable 

habitat. 

Timed hand searching 

(‘grubbing’). 4 x 30min 

samples (of 6 x 5min subsamples 

per sample). 

E.g. flies (Diptera), bugs 

(Hemiptera), beetles (Coleoptera) 

and spiders (Arachnida). 

E.g. bees and wasps etc 

(Hymenoptera) and flies (Diptera). 

A range of ground-dwelling 

invertebrates inc. beetles 

(Coleoptera), bugs (Hemiptera), 

ants (Hymenoptera) and spiders 

(Arachnida). 

All field layer vegetation 

communities, with particular focus 

upon potential food plants. 

Pollen and nectar sources; 

identified food plants; nesting 

sites. 

Ground layer habitats; e.g. under 

host plants; stones, turves etc 

Once secured in a clear tube the individuals were either identified in the field and subsequently released 

or dispatched, in an ethyl acetate killing jar / 70% Iso-Propyl alcohol, for later microscopic 

examination in the lab. Voucher specimens of notable species have been retained where critical 

examination is required to confirm identification. 

Rarity statuses for Red Data Book (RDB) and Nationally Scarce categories A & B species follow those 

given in the relevant published national species status reviews (e.g. Hyman & Parsons, 1992). Status 

categories for uncommon species are discussed in Appendix 1. The ISIS (20 10) system provides 

information on Local and widespread species. In addition, UK Biodiversity Action Plan (UK BAP) 

Priority Species and Species of Principle Importance (SPI), covered under The Natural Environment 

and Rural Communities Act 2006 Section 41 (NERC S41), are listed. The S41 list is intended to 

provide guidance to decision-makers, e.g. those representing public bodies and local/regional 

authorities, in implementing their duty under Section 40 of the NERC Act to ‘…have regard to the 

conservation of biodiversity in England when carrying out their normal functions’. 


3. Results 

Suboptimal weather conditions throughout 2012 affected the results for both sites, with expected 

species noted in previous years not found. The total number of invertebrate species recorded from each 

survey area are as follows; figures in parentheses represent species, within the total, afforded published 

conservation statuses (i.e. UK BAP/SPI, RDB or Nationally Scarce categories A or B) + those species 

regarded as nationally Local: 

Sheets Heath: 128 (17 + 30) 

Thursley Common: 108 (21 + 31) 

Species with published conservation statuses are summarised in Table 2. A full species list for each site 

(identifying Local and Widespread species etc) is given in Appendices 2 & 3. 

Table 2: Invertebrate species with UKBAP/SPI, RDB or Nationally Scarce statuses. Nationally Scarce 

species fall into two categories, a & b (Na & Nb). An asterix (*) in the appropriate site column denotes 

presence at a site. 

Order Family Taxon Vernacular National Status Surrey Status Sheets Heath Thursley 

Arachnida: 

Araneae 

Dictynidae Dictyna pusilla a spider UKBAP Rare (no modern 

records) 

Lepidoptera Satyridae Hipparchia 

semele 

Coleoptera Carabidae Cicindela 

sylvatica 

Arachnida: 

Araneae 

Gnaphosidae Haplodrassus 

dalmatensis 

Grayling UKBAP Local 

Heath Tiger 

Beetle 

Heath 

Grasper 

Spider 

Lepidoptera Lycaenidae Plebejus argus Silverstudded 

Arachnida: 

Araneae 

Oxyopidae Oxyopes 

heterophthalmus 

Hymenoptera Crabronidae Philanthus 

triangulum 

Arachnida: 

Araneae 

Uloboridae Uloborus 

walckenaerius 

Hemiptera: Lygaeidae Nysius 

Heteroptera 

helveticus 

Hymenoptera Chrysididae Hedychrum 

niemelai 

Hymenoptera Crabronidae Diodontus 

insidiosus 

Hymenoptera 

(Apidae) 

Arachnida: 

Araneae 

Blue 

Lynx 

Spider 

UKBAP / Na Local 

UKBAP / Nb Very Local 

UKBAP / Nb Locally Common 

RDB2 Locally Common 

Bee Wolf RDB2 Very common 

a spider RDB3 Very Local 

a ground 

bug 

a cuckoo 

wasp 

a digger 

wasp 

RDB3 Rare 

RDB3 Common 

RDB3 Local 

Halictinae Halictus a mining RDB3 Very Local 

confusus bee 

Gnaphosidae Zelotes petrensis a spider Na Rare 

January 2013 5 C1811 Invertebrate Monitoring Report 

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Order Family Taxon Vernacular National Status Surrey Status Sheets Heath Thursley 

Hymenoptera Crabronidae Crabro 

scutellatus 

a digger 

wasp 

Na Local 

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Hymenoptera 

(Apidae) 

Hymenoptera 

(Apidae) 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Andreninae Andrena 

argentata 

a mining 

bee 

Anthophorinae Nomada baccata a cuckoo 

bee 

Lycosidae Xerolycosa a wolf 

nemoralis spider 

Philodromidae Philodromus 

collinus 

Philodromidae Philodromus 

praedatus 

Salticidae Aelurillus vinsignitus 

Theridiidae Anelosimus 

aulicus 

Theridiidae Episinus 

truncatus 

Theridiidae Steatoda 

albomaculata 

a crab 

spider 

a crab 

spider 

a jumping 

spider 

Arachnida: Thomisidae Thomisus a crab 

Araneae 

onustus spider 

Coleoptera Carabidae Poecilus lepidus a ground 

beetle 

Coleoptera Curculionidae Smicronyx 

jungermanniae 

Dictyoptera Blattellidae Ectobius 

pallidus 

Hemiptera: 

Auchenorrhyncha 

Hemiptera: 

Heteroptera 

Hemiptera: 

Heteroptera 

Cicadellidae Zygina 

rubrovittata 

Lygaeidae Megalonotus 

dilatatus 

Lygaeidae Rhyparochromus 

pini 

Hymenoptera Formicidae Formica 

sanguinea 

Na Locally common 

Na Locally common 

Nb Local 

Nb Local 

Nb Locally Common 

Nb Local 

a spider Nb Local 

a spider Nb Rare? 

a spider Nb Rare 

Nb Local 


a weevil Nb Very Local 

Tawny 

cockroach 

Small 

Heather 

Leafhopper 

a ground 

bug 

a ground 

bug 

Slavemaker 

Ant 

Hymenoptera Pompilidae Evagetes dubius a spiderhunter 

wasp 

Hymenoptera Tiphiidae Methocha 

articulata 

a solitary 

wasp 


Nb Rare? 

Nb Local? 

Nb Local 


Nb Local 

Nb Local 

January 2013 6 C1811 Invertebrate Monitoring Report 

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Table 3a: Sheets Heath ISIS (2010) analysis of species samples – Broad Assemblage Types (BATs). 

BAT 

code 

BAT name 

Representation 

(1-100) 

Rarity 

score 

Condition 

BAT 

species 

richness 

F1 unshaded early successional mosaic 40 188 fav 51 

F2 grassland & scrub matrix 36 150 46 

A1 arboreal canopy 5 7 

A2 wood decay 3 4 

F3 shaded field & ground layer 2 3 

W3 permanent wet mire 2 3 

Table 3b: Sheets Heath ISIS (2010) analysis of species samples – Specific Assemblage Types (SATs). 

SAT 

code 

SAT name No. spp. Condition 

Percentage 

of national 

species pool 

F003 scrub-heath & moorland 20 fav 6 

Related 

BAT 

rarity 

score 

F111 bare sand & chalk 20 fav 5 188 

F001 scrub edge 8 4 

F002 rich flower resource 8 3 

A211 heartwood decay 1 1 

F112 open short sward 1 1 188 

A212 bark & sapwood decay 2 0 

Table 4a: Thursley Common ISIS (2010) analysis of species samples – Broad Assemblage Types (BATs). 

BAT code BAT name 

Representation 

(1-100) 

Rarity 

score 

Condition 

BAT 

species 

richness 

F1 unshaded early successional mosaic 49 217 fav 52 

F2 grassland & scrub matrix 36 203 fav 38 

A1 arboreal canopy 2 2 

W3 permanent wet mire 1 1 

Table 4b: Thursley Common ISIS (2010) analysis of species samples – Specific Assemblage Types 

(SATs). 

SAT code SAT name No. spp. Condition 

Percentage 

of national 

species pool 

F003 scrub-heath & moorland 26 fav 8 

Related 

BAT 

rarity 

score 

F111 bare sand & chalk 26 fav 6 217 

F002 rich flower resource 13 5 

F001 scrub edge 2 1 

F112 open short sward 2 1 217 


The results of the ISIS (2010) analysis found both study sites to be in Favourable Condition for the 

target habitat of bare ground on dry heathland, the target Specific Assemblage Type (SAT) being 

associated with bare sand - F111 ( see Tables 3b & 4b). This assemblage is a sub-category of the 

favourable Broad Assemblage Type (BAT) unshaded early successional mosaic – F1 (see Tables 3a 

& 3b). 

The assemblage representing scrub-heath (F003) was also found to be in Favourable Condition at 

both study sites. This latter result was expected given that both sites represent well-established 

heathland supporting a high proportion of heathland-specific invertebrate species, such as obligate 

Heather feeders. For Thursley Common this is further supported by the Favourable Condition of the 

BAT grassland & scrub matrix – F2. This BAT comprises many generalist grassland and scrub 

associated species, whilst also incorporating dwarf shrub specialists – i.e. those species directly 

associated with heathers on heathland. The BAT and SAT columns presented within the species lists 

for each study site in Appendices 2 & 3 denote the assemblage categories applied to each species by the 

ISIS programme. 

The columns in Appendicies 2 & 3 entitled ‘Hth’ (Heathland) and ‘BGrd’ (Bare Ground) list numeric 

values indicating each individual species fidelity to the featured habitats (where Low Fidelity = 1, 

Medium Fidelity = 2 & High Fidelity = 3). Totals (Ttl) for each species are given in the final column. 

These data detail those species with the highest fidelity to the target habitat – i.e. dry, bare ground on 

lowland heathland. It follows that a species scoring a maximum of 6 has a very high fidelity to the 

target habitat (stenotopic) and that a species scoring the minimum of 2 has no great affinity to either 

heathland or bare ground. 

Although this monitoring is in no way a direct comparison of the two sites, Thursley Common, the 

larger site and a National Nature Reserve, was found to be the superior site with 56% of the 108 species 

recorded have a Medium (22%) or High (34%) Fidelity association with bare sand on dry heathland. 

However, it should be noted that Thursley yielded slightly less species than Sheets Heath. In 

comparison analysis of the results for Sheets Heath, a much smaller site geographically adjacent to, but 

physically fragmented from, the much larger Pirbright Ranges heathland show that 40% of the 128 

species recorded have a Medium (13%) or High (27%) Fidelity association with the target habitat. 

These fidelity index results support the Favourable ISIS (2010) results , with an early successional 

mosaic (F1) / bare sand (F111) assemblage rarity scoring of 217 for Thursley and 188 for Sheets Heath 

(see Tables 3a, b & 4a, b), based upon the presence of uncommon and/or high-fidelity species 

associated with the target habitat. 


4. Entomological assessment and recommendations 

The analysis of the sample results clearly show that the SITA Trust funded creation and enhancement 

of bare ground habitat at Thursley Common and Sheets Heath has been successful for a range of 

invertebrates associated with the target habitat, including numerous UKBAP/SPI and nationally 

rare/scarce species. 

Subject to further project funding all sites within the Heath Tiger Beetle Project should be revisited 

with a view to creating further bare ground features and encouraging sustainable metapopulations of 

key invertebrate species at a site and landscape scale. 

During the 2012 monitoring several UKBAP/SPI species were conspicuous by their absence at the 

Thursley Common study area; Mottled Bee Fly Thyridanthrax fenestratus, Early Sunshiner Ground 

Beetle Amara famelica, Hornet Robberfly Asilus crabroniformis, Silver Studded Blue Butterfly Plebjus 

argus and Grayling Butterfly Hipparchia semele. However, all of these species were noted within the 

study area during 2011 and as previously mentioned in this report much invertebrate activity was 

suppressed (throughout Britain) due to sub -optimal weather conditions during spring and early 

summer. The activity period of Amara famelica ended before monitoring began in June. Observations 

for Thyridanthrax and Asilus were low across Surrey, as for many other Diptera species. Silver Studded 

Blue and Grayling Butterflies were least likely to be recorded by the survey methods used and where 

only recorded as field observations at Sheets Heath, whilst in transit between sample sites. 

The Heath Tiger Beetle Cicindela sylvatica was recorded at Thursley Common but not at Sheets Heath. 

Thursley supports a well established population, whilst Sheets Heath has never knowingly supported 

the species. However, Sheets Heath was selected by the project due to it’s proximity to populations at 

nearby Bisley ranges. It appears that C.sylvatica has not colonized this ‘stepping stone’ site as hoped. 

This may be due to insurmountable distance or habitat (e.g. woodland) between this site and the nearest 

known population, suggesting that manual translocation to suitable sites should be a consideration. 

5. References 

Drake, C.M., Lott, D.A., Alexander, K.N.A. & Webb, J. 2007a. Surveying terrestrial and freshwater 

invertebrates for conservation evaluation. Natural England Research Report NERR005. 

Sheffield: Natural England. 

Drake, C.M., Lott, D.A. & Lee, P. 2007b. Ducan’s Marsh survey report. Unpublished survey report. 

Hyman, P.S. (Revised and updated, Parsons. M .S.) 1992. A review of the scarce and threatened 

Coleoptera of Great Britain. Part 3. Peterborough: Joint Nature Conservancy Council. 

Shirt, D.B. (ed.) 1987. British Red Data Books: 2. Insects. Peterborough: Nature Conservancy 

Council. 

Lott, D.A. 2008. Synopsis of ISIS 2009 and its use in Common Standards Monitoring. ISIS 

development report. Leicestershire: Stenus Research. 

Webb, J.R. & Lott, D.A. 2006. The development of ISIS: a habitat-based invertebrate assemblage 

classification system for assessing conservation interest in England. Journal of Insect 

Conservation. 10: 179 - 188. 


6. Acknowledgements 

Thanks to the SITA Trust for funding both habitat creation and subsequent monitoring between 2009 

and 2012. Thanks also to James Giles of Natural England and Marcus Turley of The Surrey Heathland 

Project for permissions to conduct this research at the sites under their care, Thursley Common NNR 

and Sheets Heath respectively. 


7. Appendices. 

7.1. Appendix 1: Status categories for uncommon species. 

Criteria for allocation of Red Data Book (nationally rare) and Nationally Scarce (notable) statuses are varied and 

complex. I have followed the British Red Data Book for insects (Shirt, 1987) with reference to the subsequent 

JNCC reviews, which also include notable species. The Red Data Book and its subsequent reviews are already in 

need of updating and I have made reference to this in the text for individual species where it is applicable as 

many species have had range expansions or contractions in recent years. 

Red Data Book category 1. RDB1 - Endangered. Occurring only as a single population or otherwise in 

danger of extinction. 

Red Data Book category 2. RDB2 - Vulnerable. Likely to move into the RDB1 Endangered category if 

causal factors continue, species declining in their range, species in vulnerable habitats. 

Red Data Book category 3. RDB3 - Rare. Species estimated to occur in 15 or fewer of the 10km squares in 

the national Ordnance Survey grid since 1970. Localised within a restricted geographical area or thinly 

scattered over a more extensive range. 

Red Data Book category I. RDBI - Indeterminate. Taxa considered Endangered, Vulnerable or Rare, but 

where there is not enough information to assign to category (RDB1 – 3). 

Red Data Book category K. RDBK – Insufficiently known. Taxa suspected, but not definitely known due 

to lack of information, to be Endangered, Vulnerable or Rare. 

Provisional Red Data Book (X). pRDB(X). The prefix ‘p’ before any RDB category implies a provisional 

grading. 

Nationally Scarce (Notable) category A - NA. Very local species, thought to occur in 16 to 30 of the 10km 

squares of the national Ordnance Survey grid since 1970, or within 7 or fewer Vice-counties for less well 

recorded groups. 

Nationally Scarce (Notable) category B - NB. Very local species, thought to occur in 31 to 100 of the 

10km squares of the national Ordnance Survey grid since 1970, or between 8 and 20 Vice-counties for less 

well recorded groups. 

Nationally Scarce (Notable). Very local species, thought to occur in 16 to 100 of the 10km squares of the 

national Ordnance Survey grid since 1970, or between 1 and 20 Vice-counties for less well recorded groups. 

Sub-division into categories NA and NB has not been attempted due to lack of data. 

Local. Species that are restricted in distribution either geographically or by habitat. Also used for species 

that are widespread but infrequently encountered, e.g. encountered in no more than 300 10km squares of the 

national Ordnance Survey grid since 1970. 

Unknown. Usually indicates a lack of available data for difficult taxa but may also imply recent taxonomic 

confusion. 


7.2. Appendix 2: Annotated species list for Sheets Heath: 

Notes: There are two status columns, the first - National Status - indicates either the published status (e.g. UKBAP, RDB, Nationally Scarce) 

or the status given in ISIS (i.e. Local or Widespread). Widespread is perhaps an arbitrary status for those species considered to be well 

distributed and fairly common within their British geographic range. Local species tend to be those restricted either geographically or by 

habitat requirements. Naturalised and Migrant species are also indicated. Surrey Statuses are derived from published and provisional Surrey 

atlases and checklists (where available. Published atlases/checklists include: Butterflies, Larger Moths, Smaller Moths, Ants, Bees, Wasps, 

Hoverflies, Orthoptera, Shieldbugs & Beetles [checklist only]. Provisional works were consulted to inform upon Larger Brachycera Flies 

and Spiders. Other groups are based on the authors knowledge). 

The BAT and SAT columns denote the assemblage categories applied to each species by the ISIS programme and can be cross referenced 

for relevant BATs & SATs within the ISIS results tables in Section 3 of this report. A ‘0’ indicates that there is no given category. A detailed 

explanation of ISIS is provided in Appendix 4. 

The Comment column provides a brief ecological account of each species and is by no means comprehensive. Species listed as ‘Eurytopic’ 

are those species found in a wide range of habitats and situations. 

The columns titled ‘Hth’ (Heathland) and ‘BGrd’ (Bare Ground) list the numeric values of a fidelity index; Low Fidelity = 1, Medium 

Fidelity = 2 & High Fidelity = 3 – Totals (Ttl) for each species are given in the final column. These data show which species recorded during 

the survey have the highest fidelity to the target habitat - dry, bare ground (i.e. exposed dry sand) on lowland heathland. It follows that a 

species scoring a maximum of 6 has a very high fidelity to the target habitat (stenotopic) and that a species scoring the minimum of 2 has no 

great affinity to either heathland nor bare ground. 

Order Family Taxon Vernacular National 

Status 

Surrey Status BAT SAT Comment Hth BGrd Ttl 

Arachnida: Araneae Agelenidae Agelena Labyrinth Widespread Common F2 0 Eurytopic 1 1 2 

labyrinthica Spider 

Arachnida: Araneae Araneidae Agalenatea redii a spider Local Local F2 0 Heathland & other 

sand, chalk and clay 

habitats supporting 

rough grassland. 

2 1 3 

Arachnida: Araneae Araneidae Hypsosinga a spider Local Rare F2 0 Heathland & other sand 2 1 3 

albovittata 

and chalk habitats 

supporting rough 

January 2013 12 

C1811 Invertebrate Monitoring Report


Status 

Arachnida: Araneae Araneidae Mangora 

acalypha 

Arachnida: 

Araneae 


grassland. 

a spider Widespread Common F2 0 Heathland & other 



Dictynidae Dictyna pusilla a spider UKBAP Rare (no 

modern 

records) 


F2 0 Heathland, open 

forest etc. First 

Surrey record since 

1970 (Chobham 

Common). 


C1811 Invertebrate Monitoring Report 

2 1 3 

3 1 4 

Arachnida: 

Araneae 

Gnaphosidae Zelotes petrensis a spider Na Rare F1 F111 Heathland. 3 3 6 

Arachnida: Araneae Linyphiidae Bathyphantes a money Widespread Common 0 0 Eurytopic 1 1 2 

gracilis 

spider 

Arachnida: Araneae Linyphiidae Erigone 

a money Widespread Ubiquitous 0 0 Eurytopic 1 1 2 

dentipalpis spider 

Arachnida: Araneae Linyphiidae Lepthyphantes a money Widespread Common F3 0 Woodland 1 1 2 

flavipes 

spider 

Arachnida: Araneae Linyphiidae Lepthyphantes a money Widespread Ubiquitous 0 0 Eurytopic 1 1 2 

tenuis 

spider 

Arachnida: Araneae Linyphiidae Linyphia a money Widespread Common F3 0 Woodland 1 1 2 

hortensis spider 

Arachnida: Araneae Linyphiidae Linyphia a money Widespread Common 0 0 Eurytopic 1 1 2 

triangularis spider 

Arachnida: Araneae Linyphiidae Meioneta a money Widespread Common F1 0 Eurytopic 1 1 2 

rurestris spider 

Arachnida: Araneae Liocranidae Phrurolithus a spider Widespread Locally 

F2 0 Eurytopic. Associated 2 2 4 

festivus 

Common 

with ants, which the 

spider mimics. 

Arachnida: Araneae Lycosidae Arctosa perita a wolf Local Local F1 F111 Heathland & other 2 3 5 

spider 

sandy habitats 

supporting bare 

ground.


Status 


Arachnida: Araneae Lycosidae Pardosa a wolf Local Local F1 F111 Heathland & other 2 3 5 

hortensis spider 



bare ground. 

Arachnida: Lycosidae Xerolycosa a wolf Nb Local F1 0 Heathland & other 2 2 4 

Araneae 




bare ground. 

Arachnida: Araneae Philodromidae Philodromus 

aureolus 

a crab spider Widespread Common A1 0 Woodland, gardens etc 1 1 2 

Arachnida: Philodromidae Philodromus a crab Nb Locally A1 0 Associated with open 1 1 2 

Araneae 

praedatus spider 


grown Oak (swept 

from sand bank area 

adjacent to Oak). 

Arachnida: Salticidae Aelurillus v- a jumping Nb Local F1 F111 Heathland. Disturbed 3 3 6 

Araneae 

insignitus spider 

areas with bare 

ground. 

Arachnida: Araneae Salticidae Heliophanus a jumping Widespread Local F2 0 Heathland & other 2 2 4 

cupreus 

spider 


habitats. 

Arachnida: Araneae Salticidae Heliophanus a jumping Widespread Common F2 0 Heathland, grassland 2 1 3 

flavipes 

spider 

etc. 

Arachnida: Theridiidae Anelosimus a spider Nb Local F2 0 Heathlands. On Gorse 3 1 4 

Araneae 

aulicus 

etc. 

Arachnida: Araneae Theridiidae Anelosimus 

vittatus 

Arachnida: Araneae Theridiidae Enoplognatha 

latimana 

Arachnida: Araneae Theridiidae Enoplognatha 

ovata 

Arachnida: 

Araneae 

Theridiidae Episinus 

truncatus 

Arachnida: Araneae Theridiidae Paidiscura 

pallens 

a spider Widespread Common A1 0 Eurytopic. Arboreal - 

esp. on Gorse 

1 1 2 

a spider Local Local F2 0 Eurytopic (dry 

habitats) 

1 1 2 

a spider Widespread Common F2 0 Eurytopic. 1 1 2 

a spider Nb Rare? F2 0 Heathland. Amongst 

Heathers. 

a spider Widespread Common A1 0 Trees & shrubs. Swept 

from sand bank area 

adjacent to Oak. 



3 1 4 

1 1 2 

Arachnida: Araneae Theridiidae Simitidion simile a spider Local Local F2 F222 Heathland. Amongst 3 1 4


Status 

Arachnida: Araneae Theridiidae Theridion 

impressum 

Arachnida: Thomisidae Thomisus 

Araneae 

onustus 


heather. 

a spider Local Local F2 0 Heathland, grassland 

etc. Esp. on Gorse. 

a crab Nb Local F2 F222 Heathland. On 

spider 

Coleoptera Byrrhidae Byrrhus fasciatus Banded Pill 

Beetle 

Coleoptera Carabidae Carabus 

problematicus 

Coleoptera Carabidae Cicindela 

campestris 

a ground 

beetle 

Green Tiger 

Beetle 

Heathers. 



2 1 3 

3 2 5 

Widespread Rare F2 0 Heathland & other 

sandy habitats. 

2 2 4 

Widespread Local F2 0 Eurytopic (dry 

habitats) 

2 1 3 

Local Local F1 F111 Heathland & other 


habitats. 

2 3 5 

Widespread Common F1 0 Eurytopic 1 1 2 

Coleoptera Carabidae Harpalus affinis a ground 

beetle 

Coleoptera Carabidae Harpalus latus a ground 

beetle 

Coleoptera Carabidae Harpalus a ground Local Locally 

F1 F111 Heathland & other 

rufipalpis beetle 



Coleoptera Carabidae Notiophilus a ground Local Local F2 0 Heathland & other 

aquaticus beetle 


Coleoptera Carabidae Notiophilus a ground Local Local F3 0 Eurytopic (shaded 

rufipes 

beetle 

habitat) 

Coleoptera Carabidae Notiophilus a ground Local Common F1 0 Eurytopic (dry 

substriatus beetle 

habitats) 

Coleoptera Carabidae Syntomus a ground Widespread Locally 

F1 0 Heathland & other 

foveatus beetle 



Coleoptera Chrysomelidae Cryptocephalus a leaf beetle Local Local F1 F112 Heathland (on Sheep's 

fulvus 

Sorrel) & chalk etc (on 

Bird's-foot trefoil). 

Coleoptera Chrysomelidae Luperus 

longicornis 

Coleoptera Coccinellidae Coccinella 

septempunctata 

Birch Leaf 

Beetle 

7-spot 

Ladybird 

Widespread Common F2 0 Eurytopic 1 1 2 

Widespread Locally 


F2 0 Heathland etc. On 

Birch 

2 3 5 

2 2 4 

1 1 2 

1 1 2 

2 2 4 

2 3 5 

2 1 3 

Widespread Ubiquitous 0 0 Eurytopic 1 1 2


Status 


Coleoptera Coccinellidae Propylea 14-spot Widespread Ubiquitous 0 0 Eurytopic 1 1 2 

quattuordecimpunctata 

Ladybird 

Coleoptera Curculionidae Micrelus ericae Small 

Heather 

Weevil 

Coleoptera Curculionidae Neliocarus sus Heather 

Widespread Local F2 0 Heathland. Heatherspecialist 



3 1 4 

Local Local F2 0 Heathland. Heather- 3 1 4 

Weevil 

specialist 

Coleoptera Curculionidae Orchestes pilosus a weevil Local Common A1 0 On Oak (swept from 

sandy area adjacent to 

an Oak) 

1 1 2 

Coleoptera Curculionidae Rhinoncus castor Sorrel Local Local F1 0 Heathland & acid 2 2 4 

Weevil 

grassland. On Sheep's 

Sorrell. 

Coleoptera Curculionidae Sitona striatellus a weevil Widespread Locally 

F2 0 Heathland etc. On 2 1 3 


Gorse & Broom etc. 

Coleoptera Curculionidae Strophosoma Nut Leaf Widespread Common A1 0 Eurytopic 1 1 2 

melanogrammum Weevil 

Coleoptera Malachiidae Malachius Malachite Widespread Ubiquitous A2 A212 Eurytopic. Larvae 1 1 2 

bipustulatus Beetle 

saproxylic. 

Coleoptera Scarabaeidae Anomala dubia Dune Chafer Local Very Local F1 F111 Heathland & other 


2 3 5 

Coleoptera Staphylinidae Tachyporus 

hypnorum 

a rove beetle Widespread Common F2 0 Eurytopic 1 1 2 

Coleoptera Tenebrionidae Nalassus False Widespread Common 0 0 Heathland, woodland 1 1 2 

laevioctostriatus Ground 

etc. Under Heathers 

Beetle 

and under tree bark. 

Grazes algae. 

Diptera Asilidae Machimus 

atricapillus 

a robberfly Widespread Common F1 0 Eurytopic. Predatory. 1 1 2 

Diptera Asilidae Machimus a robberfly Local Common F1 F111 Heathland, grassland 2 2 4 

cingulatus 

etc. Predatory. 

Diptera Syrphidae Helophilus 

pendulus 

a hoverfly Widespread Common W3 0 Eurytopic 1 1 2


Status 


Diptera Syrphidae Myathropa florea a hoverfly Widespread Common A2 A211 Woodland. Larvae in 

rot holes of trees. 

1 1 2 

Diptera Syrphidae Sphaerophoria 

scripta 

a hoverfly Widespread Common F1 0 Eurytopic 1 1 2 

Diptera Syrphidae Xylota segnis a hoverfly Widespread Common A2 0 Eurytopic. Larvae in 

decaying timber. 

1 1 2 

Diptera Syrphidae Xylota sylvarum a hoverfly Local Common A2 A212 Woodland. Larvae in 

decaying roots of trees. 

1 1 2 

Diptera Tabanidae Chrysops 

viduatus 

a deer fly Local Local W3 0 Heathland etc. 2 1 3 

Hemiptera: Cercopidae Aphrophora alni a leafhopper Widespread Common 0 0 Eurytopic. Arboreal. 1 1 2 


bug 

Hemiptera: Cicadellidae Aphrodes a leafhopper Widespread Common F2 0 Eurytopic 1 1 2 


makarovi bug 

Hemiptera: Cicadellidae Thamnotettix Oak Widespread Common F2 F212 Oak trees. Swept from 1 1 2 


dilutior 

Leafhopper 

sand bank under Oak. 

ISIS allocations are in 

error. 

Hemiptera: Cicadellidae Ulopa reticulata Heather Widespread Locally 

F2 0 Heathland. Heather- 3 1 4 


Leafhopper 


specialist 

Hemiptera: Delphacidae Ditropis pteridis Bracken Widespread Common F2 0 Associated with 1 1 2 


Leafhopper 

Bracken in a range of 

habitats. 

Hemiptera: Lygaeidae Kleidocerys a ground Widespread Locally 


Heteroptera 

ericae 

bug 


specialist 

Hemiptera: Lygaeidae Kleidocerys a ground Widespread Common A1 0 Trees and shrubs, esp. 1 1 2 

Heteroptera 

resedae 

bug 

Birch. 

Hemiptera: Lygaeidae Macrodema a ground Widespread Locally 

F2 0 Heathland & other 2 2 4 

Heteroptera 

micropterum bug 



Hemiptera: Lygaeidae Megalonotus a ground Nb Local? F1 0 Heathland, grassland 2 2 4 

Heteroptera 

dilatatus bug 

and sandy habitats. 

Hemiptera: Lygaeidae Nysius helveticus a ground RDB3 Rare F1 F111 Heathland. 

3 3 6 

Heteroptera 

bug 

Associated with Bell 

Heather (Erica 

cinerea) 




Status 


Hemiptera: Lygaeidae Rhyparochromus a ground Nb Local F2 F222 Heathland. Under 3 1 4 

Heteroptera 

pini 

bug 

heathers. 

Hemiptera: Lygaeidae Scolopostethus a ground Widespread Locally 


Heteroptera 

decoratus bug 


specialist 

Hemiptera: Lygaeidae Trapezonotus a ground Widespread Locally 

F1 0 Heathland & chalk 2 2 4 

Heteroptera 

desertus bug 


grassland, esp. with 

bare ground. 

Hemiptera: Miridae Deraeocoris a plant bug Naturalised Common 0 0 Eurytopic. Trees & 1 1 2 

Heteroptera 

flavilinea 

shrubs. 

Hemiptera: Miridae Leptopterna a plant bug Widespread Common F2 0 Grassland. Associated 1 1 2 

Heteroptera 

dolabrata 

with various grasses. 

Hemiptera: Miridae Orthotylus a plant bug Widespread Locally 


Heteroptera 

ericetorum 


specialist 

Hemiptera: Miridae Pithanus a plant bug Widespread Common F2 0 Grassland. 1 1 2 

Heteroptera 

maerkelii 

Hemiptera: Miridae Stenodema a plant bug Widespread Common F2 0 Grassland. Associated 1 1 2 

Heteroptera 

laevigata 

with various grasses. 

Hemiptera: Nabidae Himacerus a nabid bug Widespread Common F2 0 Eurytopic (dry 

1 1 2 

Heteroptera 

mirmicoides 

habitats) 

Hemiptera: Nabidae Nabis ericetorum a nabid bug Widespread Locally 

F2 0 Heathland. Predatory 3 1 4 

Heteroptera 


amongst Heather. 

Hymenoptera Chrysididae Elampus panzeri a cuckoo Local Common F1 F111 Heathland and 

2 3 5 

wasp 

grassland. 

Cleptoparasite of 

Mimesa spp. 

Hymenoptera Chrysididae Hedychrum a cuckoo RDB3 Common F1 0 Heathland & 

2 3 5 

niemelai wasp 

grassland. 


Cerceris spp. 

Hymenoptera Crabronidae Astata boops a digger Local Common F1 0 Heathland & grassland. 2 3 5 

wasp 

Predates shieldbug 

nymphs. 

Hymenoptera Crabronidae Cerceris Ornate Widespread Common F1 0 Eurytopic. Predates 2 3 5 

rybyensis Tailed 

Digger 

Wasp 

solitary bees. 




Status 


Hymenoptera Crabronidae Diodontus a digger RDB3 Local F1 F111 Heathland & other 2 3 5 

insidiosus wasp 


Predates aphids. 

Hymenoptera Crabronidae Mimesa lutaria a digger Local Common F1 F111 Heathland. Predates 3 3 6 

wasp 

various leafhopper 

bugs. 

Hymenoptera Crabronidae Oxybelus Spiny Widespread Common F1 0 Heathland & other 2 3 5 

uniglumis Digger 


Wasp 

habitats. Predates 

various flies (Diptera), 

esp. Muscidae. 

Hymenoptera Crabronidae Tachysphex a digger Widespread Common F1 0 Heathland & acid 3 3 6 

pompiliformis wasp 

grassland. Predates 

grasshopper nymphs. 

Hymenoptera Formicidae Formica fusca an ant Widespread Common F1 0 Eurytopic (dry 

habitats) 

1 3 4 

Hymenoptera Formicidae Formica Slavemaker Nb Locally F1 0 Heathland 3 3 6 

sanguinea Ant 


Hymenoptera Formicidae Lasius niger Black Widespread Ubiquitous F1 0 Eurytopic (dry 

1 2 3 

Garden Ant 

habitats) 

Hymenoptera Formicidae Leptothorax an ant Widespread Locally 

F1 F111 Eurytopic - esp. dead 2 3 5 

acervorum 


wood, heaths and 

marginal areas of 

Formica nests. 

Hymenoptera Formicidae Myrmica sabuleti an ant Widespread Local F1 0 Heathland & dry 

grassland etc 

2 2 4 

Hymenoptera Formicidae Myrmica an ant Widespread Ubiquitous F2 0 Eurytopic (dry 

1 2 3 

scabrinodis 

habitats) 

Hymenoptera Formicidae Tetramorium 

caespitum 

an ant Local Local F1 F111 Heathland 3 3 6 

Hymenoptera Pompilidae Evagetes a spider- Widespread Common F1 0 Heathland & other 2 3 5 

crassicornis hunter wasp 


habitats. Predates nests 

of other Pompilidae. 

Hymenoptera Pompilidae Priocnemis a spider- Widespread Common F1 0 Open woodland, heaths 1 2 3 

perturbator hunter wasp 

etc. Predates spiders. 




Status 


Hymenoptera Pompilidae Priocnemis a spider- Local Common F1 0 Heathland & other 2 2 4 

pusilla 

hunter wasp 


Predates spiders. 

Hymenoptera Sphecidae Ammophila Red Banded Widespread Common F1 0 Heathland & dry 2 3 5 

sabulosa Sand Wasp 

grassland etc. Predates 

larvae of Noctuidae 

moths. 

Hymenoptera Tiphiidae Methocha a solitary Nb Local F1 F111 Heathland. Parasite 3 3 6 

articulata wasp 

of tiger beetle larvae 

(Cicindelinae) 

Hymenoptera Tiphiidae Myrmosa atra Black Widespread Common F1 0 Heathland & dry 2 3 5 

Headed 

grassland etc. Parasite 

Velvet Ant 

of various ground 

nesting Hymenoptera. 

Hymenoptera Andreninae Andrena a mining Na Locally F1 F111 Heathland. 

3 3 6 

(Apidae) 

argentata bee 

common 

Specialises in loose 

sand nesting, often on 

tracks 

Hymenoptera Andreninae Andrena fuscipes a mining bee Local Locally 

F1 F111 Heathland. Heather- 3 3 6 

(Apidae) 


specialist 

Hymenoptera Anthophorinae Anthophora a solitary Local Locally 

F1 0 Heathland & other 2 3 5 

(Apidae) 

bimaculata bee 



Hymenoptera Anthophorinae Nomada rufipes a cuckoo Widespread Common F1 F111 Heathland and clay 2 3 5 

(Apidae) 

bee 

woodland. 


Andrea fuscipes 

(heaths) & 

A.denticulata (woods). 

Hymenoptera 

(Apidae) 

Apinae Apis mellifera Honey Bee Ubiquitous Ubiquitous 0 0 Eurytopic 1 1 2 

Hymenoptera Halictinae Lasioglossum Slender Widespread Ubiquitous F1 0 Eurytopic (dry 

1 3 4 

(Apidae) 

calceatum Mining Bee 

habitats) 

Hymenoptera Halictinae Lasioglossum Brassy Widespread Ubiquitous F1 0 Eurytopic (dry 

1 3 4 

(Apidae) 

morio 

Mining Bee 

habitats) 

Hymenoptera Halictinae Lasioglossum Shaggy Widespread Ubiquitous F1 0 Eurytopic (dry 

1 3 4 

(Apidae) 

villosulum Mining Bee 

habitats) 




Status 


Hymenoptera Halictinae Sphecodes a cuckoo Widespread Common F1 0 Eurytopic. 

1 3 4 

(Apidae) 

geoffrellus bee 


various Lasioglossum 

spp. 

Hymenoptera Halictinae Sphecodes a cuckoo Local Locally 

F1 F111 Heathland & other 2 3 5 

(Apidae) 

pellucidus bee 



habitats. Cleptoparasite 

of Andrena barbilabris 

and Lasioglossum 

leucozonium. 

Isopoda Armadillidiidae Armadillidium Pill Widespread Ubiquitous 0 0 Eurytopic. Detritivore. 1 1 2 

vulgare 

Woodlouse 

Isopoda Philosciidae Philoscia Striped Widespread Ubiquitous 0 0 Eurytopic. Detritivore. 1 1 2 

muscorum Woodlouse 

Isopoda Porcellionidae Porcellio scaber Rough 

Woodlouse 

Widespread Ubiquitous 0 0 Eurytopic. Detritivore. 1 1 2 

Lepidoptera Arctiidae Diacrisia sannio Clouded Local Locally 

F2 0 Heathland & chalk 2 1 3 

Buff 


grassland, larvae feed 

on heather (heaths) and 

various other herbs 

(chalk). 

Lepidoptera Lycaenidae Lycaena phlaeas Small Widespread Locally 

F2 0 Heathland, grassland 1 1 2 

Copper 


etc. Larvae feed on 

Sorrell. 

Lepidoptera Lycaenidae Plebejus argus Silver- UKBAP / Nb Locally F2 0 Heathland. Larvae 3 3 6 

studded 


associated with Erica 

Blue 

spp. Heathers and 

Lasius spp. ants. 

Lepidoptera Noctuidae Anarta myrtilli Beautiful 

Yellow 

Underwing 

Widespread Ubiquitous F2 0 Eurytopic 1 1 2 

Lepidoptera Nymphalidae Polygonia calbum 

Comma Widespread Common F2 0 Eurytopic 1 1 2 

Lepidoptera Satyridae Hipparchia Grayling UKBAP Local F1 F111 Heathland (formerly 3 3 6 

semele 

also on downland in 

Surrey). Larvae feed 




Status 

Orthoptera Acrididae Chorthippus 

brunneus 

Orthoptera Acrididae Myrmeleotettix 

maculatus 

Orthoptera Phaneropteridae Leptophyes 

punctatissima 

Field 

Grasshopper 

Mottled 

Grasshopper 

Speckled 

Bush 

Cricket 

Orthoptera Tetrigidae Tetrix undulata Ground 

Hopper 

Orthoptera Tettigoniidae Metrioptera 

brachyptera 

Bog Bush 

Cricket 


on fine grasses. 

Widespread Common F2 0 Grassland etc 1 1 2 


F1 0 Heathland & acid 


grassland. 

Widespread Common F2 F212 Scrub edge, woodland 

etc 



0 0 Heathland, chalk 

grassland and 

open/cleared woodland 

supporting bare 

ground. 

Local Local W3 0 Heathland. Formerly 

Nb 



3 2 5 

1 1 2 

2 3 5 

3 1 4

7.3. Appendix 3: Annotated species list for Thursley Common: 

Notes: There are two status columns, the first - National Status - indicates either the published status (e.g. UKBAP, RDB, Nationally Scarce) 

or the status given in ISIS (i.e. Local or Widespread). Widespread is perhaps an arbitrary status for those species considered to be well 

distributed and fairly common within their British geographic range. Local species tend to be those restricted either geographically or by 

habitat requirements. Naturalised and Migrant species are also indicated. Surrey Statuses are derived from published and provisional Surrey 

atlases and checklists (where available. Published atlases/checklists include: Butterflies, Larger Moths, Smaller Moths, Ants, Bees, Wasps, 

Hoverflies, Orthoptera, Shieldbugs & Beetles [checklist only]. Provisional works were consulted to inform upon Larger Brachycera Flies 

and Spiders. Other groups are based on the authors knowledge). 

The BAT and SAT columns denote the assemblage categories applied to each species by the ISIS programme and can be cross referenced 

for relevant BATs & SATs with the ISIS results tables in Section 3 of this report. A ‘0’ indicates that there is no given category. A detailed 

explanation of ISIS is provided in Appendix 4. 

The Comment column provides a brief ecological account of each species and is by no means comprehensive. Species listed as ‘Eurytopic’ 

are those species found in a wide range of habitats and situations. 

The columns titled ‘Hth’ (Heathland) and ‘BGrd’ (Bare Ground) list the numeric values of a fidelity index; Low Fidelity = 1, Medium 

Fidelity = 2 & High Fidelity = 3 – Totals (Ttl) for each species are given in the final column. These data show which species recorded during 

the survey have the highest fidelity to the target habitat - dry, bare ground (i.e. exposed dry sand) on lowland heathland. It follows that a 

species scoring a maximum of 6 has a very high fidelity to the target habitat (stenotopic) and that a species scoring the minimum of 2 has no 

great affinity to either heathland nor bare ground. 


Status 


Arachnida: Araneae Agelenidae Tegenaria 

gigantea 

a spider Local Common 0 0 Eurytopic, including houses. 1 1 2 

Arachnida: Araneae Araneidae Agalenatea redii a spider Local Local F2 0 Heathland & other sand, 

chalk and clay habitats 

supporting rough grassland. 

2 1 3 

Arachnida: Araneae Araneidae Araneus a spider Widespread Local 0 0 Heathland & other sand and 2 1 3 

quadratus 

chalk habitats supporting 





Status 


Arachnida: Araneae Araneidae Hypsosinga a spider Local Rare F2 0 Heathland & other sand and 2 1 3 

albovittata 

chalk habitats supporting 


Arachnida: Araneae Araneidae Mangora a spider Widespread Common F2 0 Heathland & other sand, 2 1 3 

acalypha 


supporting rough grassland. 

Arachnida: Araneae Clubionidae Clubiona a spider Widespread Local F2 0 Heathland. Amongst Heather 3 1 4 

trivialis 

& Gorse. 

Arachnida: Araneae Dictynidae Dictyna latens a spider Local Locally F2 F222 Heathland and rough 

2 1 3 


grassland. 

Arachnida: Gnaphosidae Haplodrassus Heath UKBAP / Nb Very Local F1 F111 Heathland. Amongst 3 3 6 

Araneae 

dalmatensis Grasper 

Heather and under stones 

Spider 

etc. 

Arachnida: Araneae Linyphiidae Erigone atra a money 

spider 

Widespread Ubiquitous 0 0 Eurytopic 1 1 2 

Arachnida: Araneae Linyphiidae Erigone a money Widespread Ubiquitous 0 0 Eurytopic 1 1 2 

dentipalpis spider 

Arachnida: Araneae Linyphiidae Lepthyphantes a money Widespread Ubiquitous 0 0 Eurytopic 1 1 2 

tenuis 

spider 

Arachnida: Araneae Linyphiidae Ostearius a money Widespread Local 0 0 Eurytopic 1 1 2 

melanopygius spider 

Arachnida: Araneae Lycosidae Arctosa perita a wolf spider Local Local F1 F111 Heathland & other sandy 

habitats supporting bare 

ground. 

2 3 5 

Arachnida: Lycosidae Xerolycosa a wolf Nb Local F1 0 Heathland & other sand, 2 2 4 

Araneae 



supporting bare ground. 

Arachnida: Oxyopidae Oxyopes Lynx Spider RDB2 Locally F2 F222 Heathland (virtually 3 1 4 

Araneae 

heterophthalmus 


confined to the Surrey 

heaths) 

Arachnida: Philodromidae Philodromus a crab Nb Local A1 0 Associated with coniferous 2 1 3 

Araneae 

collinus spider 

trees, esp. Scot's Pine. 

Arachnida: Araneae Philodromidae Philodromus a crab spider Local Local F2 0 Heathlands. Amongst 3 1 4 

histrio 

Heather. 

Arachnida: Araneae Philodromidae Tibellus 

oblongus 

a crab spider Widespread Common F2 0 Grasslands 1 1 2 




Status 


Arachnida: Salticidae Aelurillus v- a jumping Nb Local F1 F111 Heathland. Disturbed areas 3 3 6 

Araneae 

insignitus spider 

with bare ground. 

Arachnida: Araneae Salticidae Heliophanus a jumping Widespread Common F2 0 Heathland, grassland etc. 2 1 3 

flavipes spider 

Arachnida: Araneae Theridiidae Anelosimus a spider Widespread Common A1 0 Eurytopic. Arboreal - esp. on 1 1 2 

vittatus 

Gorse 

Arachnida: Araneae Theridiidae Simitidion simile a spider Widespread Local F2 F222 Heathland. Amongst heather. 3 1 4 

Arachnida: 

Araneae 

Arachnida: 

Araneae 

Theridiidae Steatoda 

albomaculata 

Thomisidae Thomisus 

onustus 

Arachnida: Araneae Thomisidae Xysticus 

sabulosus 

Arachnida: 

Araneae 

Uloboridae Uloborus 

walckenaerius 

a spider Nb Rare F1 0 Heathland. Under heather 

adjacent to bare ground. 

a crab 

spider 

Nb Local F2 F222 

Ant predator. 

Heathland. On Heathers. 3 2 5 



3 3 6 

a spider Local Rare F1 F111 Heathland. On bare ground. 3 3 6 

a spider RDB3 Very Local F2 F222 Heathland. Webs amongst 

heather. 

3 1 4 

Arachnida: 

Opiliones 

Phalangiidae Mitopus morio a harvestman Widespread Common 0 0 Eurytopic 1 1 2 

Coleoptera Apionidae Apion 

Red Sorrel Widespread Local F2 0 Heathland & acid grassland. 2 2 4 

haematodes Weevil 

On Sheep's Sorrell. 

Coleoptera Byrrhidae Byrrhus Banded Pill Widespread Rare F2 0 Heathland & other sandy 2 2 4 

fasciatus Beetle 

habitats. 

Coleoptera Byrrhidae Byrrhus pilula Pill Beetle Widespread Local F1 0 Heathland & grassland. 2 2 4 

Coleoptera Byrrhidae Byrrhus 

pustulatus 

Pill Beetle Local Rare F2 0 Heathland. 3 2 5 

Coleoptera Carabidae Bradycellus a ground Widespread Local F2 0 Heathland & other sandy 2 2 4 

harpalinus beetle 

habitats. 

Coleoptera Carabidae Bradycellus a ground Widespread Local F2 0 Heathland & other sandy 2 2 4 

ruficollis beetle 

habitats. 

Coleoptera Carabidae Calathus erratus a ground Widespread Local F1 F111 Heathland & other sandy 2 3 5 

beetle 

habitats. 

Coleoptera Carabidae Cicindela Green Tiger Local Local F1 F111 Heathland & other sand, 2 3 5 

campestris Beetle 

chalk and clay habitats.


Status 


Coleoptera Carabidae Cicindela Heath Tiger UKBAP / Na Local F1 F111 Heathland. 3 3 6 

sylvatica Beetle 

Coleoptera Carabidae Harpalus 

rubripes 

Coleoptera Carabidae Harpalus 

rufipalpis 

Coleoptera Carabidae Notiophilus 

germinyi 

Coleoptera Carabidae Notiophilus 

substriatus 

Coleoptera Carabidae Olisthopus 

rotundatus 

a ground 

beetle 

a ground 

beetle 

a ground 

beetle 

a ground 

beetle 

a ground 

beetle 

Local Local F1 0 Heathland & other sand, 

chalk and clay habitats. 

Local Locally F1 F111 Heathland & other sandy 


habitats. 



2 2 4 

2 3 5 

Local Rare F2 0 Heathland & other sand, 


2 2 4 

Local Common F1 0 Eurytopic (dry habitats) 1 1 2 

Local Locally 


Nb Locally 




F1 0 Heathland & other sand, 


2 2 4 

F1 F111 Heathland 3 3 6 

Coleoptera Carabidae Poecilus lepidus a ground 

beetle 

Coleoptera Carabidae Syntomus a ground 

F1 0 Heathland & other sandy 2 2 4 

foveatus beetle 

habitats. 

Coleoptera Chrysomelidae Cryptocephalus a leaf beetle Local Local F1 F112 Heathland (on Sheep's 2 2 4 

fulvus 

Sorrel) & chalk etc (on 

Bird's-foot trefoil). 

Coleoptera Curculionidae Neliocarus sus Heather 

Weevil 

Local Local F2 0 Heathland. Heather-specialist 3 1 4 

Coleoptera Curculionidae Rhinoncus castor Sorrel Local Local F1 0 Heathland & acid grassland. 2 2 4 

Weevil 

On Sheep's Sorrell. 

Coleoptera Curculionidae Smicronyx a weevil Nb Very Local F2 F222 Heathland & chalk 

2 2 4 

jungermanniae 

grassland. On Dodder. 

Coleoptera Elateridae Athous 

haemorrhoidalis 

a click beetle Widespread Ubiquitous F2 0 Eurytopic 1 1 2 

Coleoptera Silphidae Silpha tristis a beetle Local Local? 0 0 Grassland etc. Snail predator 1 1 2 

Coleoptera Staphylinidae Ischnosoma 

splendidum 

Coleoptera Staphylinidae Ocypus 

aeneocephalus 

Coleoptera Staphylinidae Quedius 

persimilis 

a rove beetle Widespread Local F2 0 Uncertain ? ? ## 

a rove beetle Local Local F2 0 Heathland & other sandy 

habitats. Associated with 

rabbit burrows? 

a rove beetle Widespread Local 0 0 Heathland & other sand, 


2 2 4 

2 2 4


Status 


Coleoptera Tenebrionidae Melanimon a darkling Local Very Local F1 F111 Heathland & other sandy 2 3 5 

tibialis 

beetle 

habitats. 

Dictyoptera Blattellidae Ectobius Tawny Nb Locally 

0 0 Heathland & chalk 

2 2 4 

pallidus cockroach 


grassland. 

Diptera Syrphidae Scaeva pyrastri a hoverfly Migrant Common F1 0 Eurytopic 1 1 2 

Diptera Tabanidae Chrysops 

viduatus 

a deer fly Local Local W3 0 Heathland etc. 2 1 3 

Hemiptera: Cicadellidae Aphrodes a leafhopper Widespread Common F2 0 Eurytopic 1 1 2 


makarovi bug 

Hemiptera: Cicadellidae Ulopa reticulata Heather Widespread Locally F2 0 Heathland. Heather-specialist 3 1 4 


Leafhopper 


Hemiptera: Cicadellidae Zygina 

Small Nb Rare? F2 F222 Heathland. Heather- 3 1 4 


rubrovittata Heather 

Leafhopper 

specialist 

Hemiptera: Anthocoridae Orius niger a bug Widespread Locally F2 0 Heathland. Heather-specialist 3 1 4 

Heteroptera 


Hemiptera: Lygaeidae Kleidocerys a ground bug Widespread Locally F2 0 Heathland. Heather-specialist 3 1 4 

Heteroptera 

ericae 


Hemiptera: Lygaeidae Macrodema a ground bug Widespread Locally F2 0 Heathland & other sandy 2 2 4 

Heteroptera 

micropterum 


habitats. 

Hemiptera: Lygaeidae Nysius 

a ground RDB3 Rare F1 F111 Heathland. Associated with 3 3 6 

Heteroptera 

helveticus bug 

Bell Heather (Erica 

cinerea) 

Hemiptera: Lygaeidae Scolopostethus a ground bug Widespread Locally F2 0 Heathland. Heather-specialist 3 1 4 

Heteroptera 

decoratus 


Hemiptera: 

Heteroptera 

Miridae Calocoris stysi a plant bug Widespread Common F2 0 Eurytopic 1 1 2 

Hemiptera: Miridae Orthotylus a plant bug Widespread Locally F2 0 Heathland. Heather-specialist 3 1 4 

Heteroptera 

ericetorum 


Hemiptera: Nabidae Nabis ericetorum a nabid bug Widespread Locally F2 0 Heathland. Predatory 

3 1 4 

Heteroptera 


amongst Heather. 

Hemiptera: Pentatomidae Piezodorus Gorse Widespread Locally F2 0 Heathland etc. On Gorse etc 2 1 3 

Heteroptera 

lituratus Shieldbug 





Status 


Hemiptera: Reduviidae Coranus Heath Local Local F1 F111 Heathland and other sandy 3 3 6 

Heteroptera 

subapterus Assasin Bug 

habitats. 

Hymenoptera Crabronidae Crabro 

a digger Na Local F1 0 Heathland. Predator of 3 3 6 

scutellatus wasp 

Dolichopodidae (Diptera) 

Hymenoptera Crabronidae Crossocerus Wesmael's Widespread Common F1 F111 Heathland & other sandy 2 3 5 

wesmaeli Digger Wasp 

habitats. Predates small 

Diptera. 

Hymenoptera Crabronidae Mimesa lutaria a digger Local Common F1 F111 Heathland. Predates various 3 3 6 

wasp 

leafhopper bugs. 

Hymenoptera Crabronidae Philanthus Bee Wolf RDB2 Very common F1 0 Eurytopic. Honey Bee 2 3 5 

triangulum 

predator. Recent range 

increase. 

Hymenoptera Formicidae Formica Slavemaker Nb Locally F1 0 Heathland 3 3 6 

sanguinea Ant 

common 

Hymenoptera Formicidae Lasius flavus Yellow 

Meadow Ant 

Widespread Common F1 F112 Eurytopic 1 2 3 

Hymenoptera Formicidae Lasius 

meridionalis 


Hymenoptera Formicidae Lasius niger Black 

Garden Ant 


Hymenoptera Formicidae Lasius 

psammophilus 


Hymenoptera Formicidae Myrmica an ant Widespread Local F1 0 Heathland & dry grassland 2 2 4 

sabuleti 

etc 

Hymenoptera Formicidae Tetramorium 

caespitum 


Hymenoptera Pompilidae Anoplius viaticus Black Local Locally F1 F111 Heathland & other sandy 2 3 5 

Banded 

Spider Wasp 


habitats. Spider predator. 

Hymenoptera Pompilidae Evagetes dubius a spider- Nb Local F1 F111 Heathland. Predates nests 3 3 6 

hunter wasp 

of other Pompilidae spp. 

Hymenoptera Pompilidae Pompilus Leaden Widespread Local F1 F111 Heathland & other sandy 2 3 5 

cinereus Spider Wasp 

habitats. Spider predator. 

Hymenoptera Pompilidae Priocnemis a spider- Local Common F1 0 Heathland & dry grassland 2 2 4 

pusilla 

hunter wasp 

etc. Spider predator. 




Status 


Hymenoptera Sphecidae Ammophila Heath Sand Very Local Locally F1 F111 Heathland. Predates larvae of 3 3 6 

pubescens Wasp 

common 

heather-specialist moths. 

Hymenoptera Sphecidae Ammophila Red Banded Widespread Common F1 0 Heathland & dry grassland 2 3 5 

sabulosa Sand Wasp 

etc. Predates larvae of 

Noctuidae moths. 

Hymenoptera Tiphiidae Myrmosa atra Black Widespread Common F1 0 Heathland & dry grassland 2 3 5 

Headed 

etc. Parasite of various 

Velvet Ant 

ground nesting 

Hymenoptera. 

Hymenoptera Andreninae Andrena a mining Na Locally F1 F111 Heathland. Specialises in 3 3 6 

(Apidae) 

argentata bee 

common 

loose sand nesting, often on 

tracks 

Hymenoptera 

(Apidae) 

Andreninae Andrena ovatula a mining bee Widespread Local F1 0 Heathland & acid grassland 3 3 6 

Hymenoptera Anthophorinae Epeolus cruciger a solitary bee Local Locally F1 0 Heathland. Cleptoparasite of 3 3 6 

(Apidae) 

common 

Colletes spp. 

Hymenoptera Anthophorinae Nomada baccata a cuckoo Na Locally F1 F111 Heathland. Cleptoparasite 3 3 6 

(Apidae) 

bee 

common 

of Andrena argentata 

Hymenoptera Anthophorinae Nomada a cuckoo bee Widespread Local F1 0 Heathland, woodland etc. 2 3 5 

(Apidae) 

leucophthalma 

Cleptoparasite of several 

Andrena spp. 

Hymenoptera 

(Apidae) 

Apinae Apis mellifera Honey Bee Ubiquitous Ubiquitous 0 0 Eurytopic 1 1 2 

Hymenoptera Apinae Bombus jonellus Heath Widespread Locally F2 0 Heathland & grassland 2 1 3 

(Apidae) 

Bumble Bee 

common 

Hymenoptera Apinae Bombus Large Red Ubiquitous Ubiquitous F1 0 Eurytopic 1 1 2 

(Apidae) 

lapidarius Tailed 

Bumble Bee 

Hymenoptera Apinae Bombus lucorum White-tailed Ubiquitous Ubiquitous F1 0 Eurytopic 1 1 2 

(Apidae) 

Bumble Bee 

Hymenoptera Apinae Bombus Common Ubiquitous Ubiquitous F1 0 Eurytopic 1 1 2 

(Apidae) 

pascuorum Carder Bee 

Hymenoptera Colletinae Colletes a mining bee Widespread Locally F1 F111 Heathland. Heather-specialist 3 3 6 

(Apidae) 

succinctus 

common 




Status 


Hymenoptera Halictinae Halictus a mining RDB3 Very Local F1 F111 Heathland. Nests in 

3 3 6 

(Apidae) 

confusus bee 

aggregations. 

Hymenoptera 

(Apidae) 

Hymenoptera 

(Apidae) 

Halictinae Sphecodes 

ephippius 

Megachilinae Coelioxys 

conoidea 

a cuckoo bee Widespread Common F1 0 Eurytopic. Cleptoparasite of 

various Lasioglossum spp. 

1 3 4 

a cuckoo bee Local Very Local F1 F111 Heathland & other sandy 

habitats. Cleptoparasite of 

Megachile maritima 

(recorded at Thursley by 

SGD in 2011) 

2 3 5 

Widespread Ubiquitous 0 0 Eurytopic. Detritivore. 1 1 2 

Isopoda Porcellionidae Porcellio scaber Rough 

Woodlouse 

Lepidoptera Noctuidae Anarta myrtilli Beautiful 

Yellow 

Underwing 


Lepidoptera Noctuidae Autographa 

gamma 

Silver Y Migrant Ubiquitous 0 0 Eurytopic 1 1 2 

Lepidoptera Pyralidae Crambus a micro-moth Widespread Common 0 0 Grassland (and other habitats 1 1 2 

pascuella 

supporting at least some 

grasses!) 

Lepidoptera Satyridae Maniola jurtina Meadow Widespread Common F2 0 Grassland (and other habitats 1 1 2 

Brown 

supporting at least some 

grasses!) 

Lithobiomorpha Lithobiidae Lithobius 

forficatus 

a centipede Widespread Common 0 0 Eurytopic. 1 1 2 

Orthoptera Acrididae Myrmeleotettix Mottled Widespread Locally F1 0 Heathland & acid grassland. 3 2 5 

maculatus Grasshopper 




7.4. Appendix 4: Invertebrate Species-habitat Information System (ISIS, 2010) assemblage 

assessment. 

Species assemblage information has been compiled using the ISIS 2010 (Invertebrate Species-habitat 

Information System; Webb & Lott, 2006) computer programme developed by Natural England. ISIS 

applications are detailed in Drake et al. (2007a), with further developments discussed by Lott (2008). 

Lott (2008) describes the essence of ISIS as a database that can be used to recognise invertebrate assemblage 

types in species lists and evaluate their value for nature conservation. In broad laymans terms ISIS has been 

described as the equivalent to an ‘invertebrate National Vegetation Classification’. An explanation of the ISIS 

(2008) system based on Drake et al. (2007a) and Lott (2008) and adapted from the summaries given in Drake et 

al. (2007b) and Lott (2008) is presented below: 

Interprets species lists via recognition and conservation quality scoring of assemblage types present. 

Developed for Common Standards Monitoring (CSM) on Sites of Special Scientific Iinterest (SSSI) but 

other applications are possible at a range of geographic scales. 

Two levels of assemblage type are recognised (BAT & SAT) with a recently introduced (ISIS, 2008 - 2010) 

third resource based category which sits outside of the hierarchy of ISIS (2007): 

o 11 Broad Assemblage Types (BATs) characterised by more widespread species. 

o 17 Specific Assemblage types (SATs) characterise d by stenotopic species (i.e. those able to 

tolerate only a restricted range of habitats and ecological conditions) of intrinsic nature 

conservation value. 

o A further 8 assemblage types dependant upon important habitat resources or environmental 

gradients. I have provisionally called these largely Resource-based Assemblage Types (RATs). 

RATs are habitat specific and are therefore shown as SATs in the results section of ISIS (2010). 

Scores generated for BATs: 

o Representation score – measures relative importance of the BAT represented in the species list 

on a scale of 1 – 100. On a small scale (e.g. a management unit within a site) this can provide a 

crude measure of ecological change. On a larger scale it is merely representative of the habitats 

sampled. Using the visibility threshold options can refine this score to show only assemblages of 

significance. 

o Rarity score – the mean of all the individual species rarity scores present in the assemblage. 

Rarity scores are based upon conservation status and / or distribution at a 10km sq resolution, 

similar to Species Quality Index scoring systems. 

o BAT species richness – A count of species coded to BAT 

Scores generated for SATs: 

o Weighted species score – Equal to the number of species coded to a SAT, though species of 

particularly high fidelity may be weighted. Required for CSM favorable condition assessment 

(SSSI’s only). 

o No. spp. – A count of species coded to SAT. 

o % of national species pool – No. spp. divided by species index for SAT. 

o Related BAT rarity score – Rarity score of the parent BAT. 


7.5. Appendix 5: Map of sampling locations at Sheets Heath. 

7.6. Appendix 6: Map of sampling locations at Thursley Common. 


Based upon the Ordnance Survey 1:10 000 Map 

with the permission of The Controller of 

Her Majesty's Stationery Office. Crown Copyright. 

Licence Number AL100031669. 

C1811, January 2013 

 

 

 

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Appendix 6 

Map of sampling locations at Sheets Heath 

Aerial View 

 

Scale 1:3500 

Produced by: Surrey Wildlife Trust Consultancy, part of Surrey Wildlife Trust






 

 

 

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Appendix 6 

Map of sampling locations at Sheets Heath 

OS View 

 

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Appendix 6 

Map of sampling locations at Thursley Common 

Aerial View 

 

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Appendix 6 

Map of sampling locations at Thursley Common 

OS View 

 

Scale 1:3500

Thursley Common [Shrike Hill] (SU9041) & Sheets Heath (SU9457 ...

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