Butterfly Tour Report Ghana 26 Okt-7 Nov Larsen Sáfián BCGHANA

RECORDS FROM THE “BUTTERFLY CONSERVATION
SOCIETY, GHANA” TOUR THROUGH GHANA DURING
OCT/NOV 2008
Photo: The Ghost – Pseudopontia paradoxa by David Dennis
Torben B. Larsen & Szabolcs Sáfián

Tour programme
The present tour was organized mainly for participants wishing to observe and to photograph
a wide selection of the almost 950 butterflies known from Ghana. There were ten participants,
with Szabolcs Sáfián as the tour leader, and with Torben B. Larsen as accompanying
commentator. It began in the Shai Hills Guinea savannah – part of the Dahomey Gap – and
then moved north to the Aburi Botanical Gardens, a tiny remnant of the rainforests that
clothed the Akwapim Scarp just 50 years ago. From there the tour went to the magnificent,
and very special, Upland Evergreen forests of the Atewa Range in the Kibi area, and then to
the Bobiri Butterfly Sanctuary in Moist Semi-Deciduous forest. From there we went south to
Cape Coast to visit the now famous Kakum National Park in the Moist Evergreen forest zone,
and then east to Ankasa on the border with Côte d’Ivoire, in the wettest rainforest habitat in
Ghana, the Wet Evergreen Forest of Ankasa National Park. Time constraints did not allow us
to approach the Guinea savannahs of northern Ghana, but more than 80% of Ghana’s
butterflies are essentially forest species – and the Shai Hills savannah had already given a
graphic impression of this very different habitat.
Itinerary and the localities
26 th October – Arrival to Accra and Shai Hills Resort
27 th October – Shai Hills Nature Reserve
including the bat cave, and the tropical rainstorm which soaked us and the bus driver who
abandoned us.
28 th October – Aburi Botanical Garden
Nice and gentle day with lots of interesting butterfly species. We have all learnt how furry a
Lycaenidae larva can be.
29 th October – Atewa Forest Reserve
Gentle climbing from the main road, meeting with illegal loggers, the rain caught us again,
after the rain some of us discovered the farmland above Asiakwa.
30 th October – Atewa Forest Reserve
We were transported to the heights of Atewa Range.
31 st October – Bobiri Butterfly Sanctuary
Walk around the compound, we had a very late but very delicious Red Red.
1 st November – Bobiri Butterfly Sanctuary, accommodation at Bobiri Guesthouse
Full day in Bobiri, Torben’s group got lost. Motto: UBA LUBA! The children peed on Sáfi’s
tent.
2 nd November – Travel to Kakum National Park
Fantastic food at Hans Cottage Botel with Nile Crocodiles, good moth night at Kakum
Guesthouse

3 rd November – Kakum National Park
Early morning Canopy Walkway, than visiting Antiakwa which was a kind of
disappointment, since we couldn’t find a trail into the forest. We’ve lost Mike taking a trail to
the boundary line of the National Park.
4 th November – ‘The Forest’ Ecotoursim Conservation Area, (Simon’s compound)
Good but a bit muddy butterfly walk, with Forest Elephant dung. We have found lots of
rewarding species later on.
5 th November – Ankasa National Park, accommodation at Axim Beach Hotel
Rainy afternoon around the gate of Ankasa NP. No success of finding 3 species of Oboronia
for David.
6 th November – Ankasa National Park, accommodation at Axim Beach Hotel
Some members stayed in the Hotel for whale watching. The others went to Nkwanta Camp,
where we found a 15 metres wide motorway into the forest.
7 th November – Travel to Accra, evening flight to the UK
Cultural visit at the slave fort of Cape Caslte, last shopping in Accra. Farewell.
Summary of results
During the tour the weather was not as clement as we normally expect at the time. On two
occasions most of us probably got more thoroughly drenched by sudden thundershowers than
we have ever been before. There is a good reason why the forests of Ghana are called
rainforests! We hope that everyone on the team took such incidents with good spirits, though
there might have been a bit of envy to a few members who had foreseen the rain and repaired
to a “spot” in a neighbouring village, where shelter and cold beer was at hand!
However, in all about 350 species were recorded during the relatively brief trip – that is
approaching the total number of species in all of Western Europe. One in three of all Ghana
butterflies were recorded. Five or six additional species will be found when genitalia are
examined. That is pretty impressive by any standards, not least taking the indifferent weather
conditions into consideration – and also the fact that some 150 strict savannah species and
Volta Region endemics were not available. However, none was a new country record for
Ghana – which is an indication that the country has now been quite well researched. We are
getting only one or two new country records a year and a species new to science only every
few years. On the tour with “A Brush with Nature” in 2006 (20 Oct – 4 Nov) 439 species
were recorded, but this was a longer trip, under better weather conditions, and with more
collectors rather than photographers. In August 1996 a team of 15 very experienced American
collectors recorded 450 species during 13 days in the field under the guidance of Larsen –
essentially to all the same localities.
There were about 40 new locality records from the sites visited – and all of these are among
those forests being kept under close scrutiny since 1993. That might not seem like a lot … but
it actually is. We have kept the places visited during the tour under close observation since
1993/1994 and every new locality record is to be treasured. The reason that there are
relatively few new records is that we now have a really good “take” on these Ghanaian
forests. Nowhere else in the African rainforest zone do we have forests that have been so
assiduously monitored, except for Kakamega Forest in Kenya where huge numbers of
butterflies are available in Nairobi.

Table 1 summarizes the species recorded from individual localities during the trip and
compares them to the recorded totals of these. We have not yet re-calculated the estimated
totals for the localities in question. However, Atewa probably has about 700 species, Kakum
650, Bobiri and Ankasa 630, Aburi more than 400, and Shai Hills a bit more than 150. The
lacking positive records are all the rarest and most difficult to catch, so at this point further
progress becomes a very slow process.
Table 1. Total species recorded from the localities visited by the tour,
number of species recorded by the team, and new records to the individual
localities (a few additional species have been added since).
__________________________________________________________
Locality Current Recorded % of New
total on tour known total records
__________________________________________________________
Shai Hills 114 56 49% 14
Aburi Bot. Gardens 316 134 42% 8
Atewa Range 604 183 30% 1
Bobiri Forest 469 196 42% 9
Kakum NP 509 138 27% 5
Ankasa NP** 405 76 19% 2
Totals Ghana* 935 345 37% 0
__________________________________________________________
* Collins, de Jong, Boersma, and local collectors continued collecting till late
November, reaching a total of 410+ species for the entire month starting 27 October.
** Larsen was not present for most of the Ankasa visit and weather was indifferent,
resulting in a low total.
In the Excel appendix the species recorded during the tour from different localities are given
in red, compared with the total list of known species. Also included are the results from the
field trip following the Butterfly Conference, given in green for comparison. At Bobiri Steve
Collins and two local collectors for the African Butterfly Research Institute, Nairobi recorded
no less than 250 species during 2.5 days (our team recorded almost 200 during 1.5 days).
The most interesting records from the trip were:
Cellular Harlequin - Mimeresia cellularis was found on Atewa, the first time Larsen ever
saw the species west of the Dahomey Gap. There are a few records from the Volta Region and
just one previous from Atewa. Its close relative, M. moyambina was caught only three times
during the 20 th century, the type from Sierra Leone, three from Cote d’Ivoire, and a few from
Atewa. Both are readily identifiable as unusual butterflies and it is amazing that they are seen
so rarely, even in Ghana’s best exploited locality.
Likpe Sapphire - Iolaus likpe was found dead on the forest floor close to the guesthouse in
Bobiri Butterfly Sanctuary. This is only the second specimen from west of the Dahomey Gap,
the first one was caught by Sáfi (which was also the first recorded female) just two years ago.
The male holotype of this extremely rare species was bred by Father Theodore Maessen in

Like Mate. The larva was collected from the mistletoe infested citrus trees around his house in
the 1970s.
Black-Spotted Ciliate Blue - Cupidesthes mimetica from Kakum (Simon’ compound). This
species [perhaps more than one] is poorly known, though widely distributed. Only good series
from single localities will eventually bring this species to “heel”. Safi previously caught a
few. Larsen did not find it during a follow-up visit.
Bright Bush Blue - Cacyreus audeoudi was brought for inspection in a pill-box at Bobiri.
Larsen initially dismissed it as the very common C. lingeus, but Safi insisted on closer
investigation. We have not seen this from Ghana before, and there are few reliable records.
The colour of the upperside is much deeper violet than in the common species and was
immediately diagnostic. We make a practice of examining a few specimens from any locality
visited and feel sure we have not seen it before. Our surprise at later finding it later also in
Kakum was complete!
Western Blue Beauty - Protogoniomorpha cytora has no records of early stages. A larva was
seen on Atewa on a plant on which Larsen saw a female ovipositing many years ago. We
hope that one or more of the participants have good photos [as they indeed were].
Small Palm Nightfighter - Zophopetes ganda was collected by Sáfi as new to Kakum at light
on the wall of the guesthouse at 5 a.m. along with few Monza cretacea. It probably visited the
flowers of a nearby papaya plant. The species was later also found at Aburi. It is one of very
few Ghana records and the first within the system of protected areas (National Parks)
Scarce Forest Swift - Melphina tarace. Surprisingly, 3 specimens of this rare species were
found during our 2 days visit in Bobiri. The first one came to moth light, the second one has
got trapped in Torben’s balcony while he was reading there and the third one visited a
Bougainville flower at the guesthouse. The species wasn’t recorded before the trip, despite the
intensive activities during the last few years.
White-Spotted Forest Swift - Melphina statira was photographed as new to Bobiri. Neither
of us has seen this species in Ghana and there are just a handful of old records.
Photography versus collecting
Participants on the tour were not butterfly collectors … they were observers and
photographers. Many superb photos were taken. But as the tour went ahead, most participants
realized that as scientists we needed as many records as possible. They started taking
“voucher photographs” – if they saw something they did not remember seeing before, they
would snap a photo from a distance, and then perhaps try to get a really nice shot later. We sat
through several thousand snaps and found many interesting species. We had never seen the
rare skipper, Melphina statira, in Ghana, from where there are but a handful of old records …
but there it was, a poor specimen in frightful condition on a wonky photograph from Bobiri,
yet still readily identifiable and a new locality record. All in all maybe 20 of the 345 species
were recorded only on pictures.
The concept of “voucher photography” needs further development and incorporation into
future photographic tours on a more systematic basis. This is particularly important when no
specialists are present at the time and when areas where lists of species are not available.

Conclusion
The weather was unfortunately not helpful. Though the number of species recorded was quite
satisfactory, the number of individuals on the wing was rather low. Late October/early
November is usually a good period but the timing of rainfall has been increasingly erratic in
recent years. However, overall the trip should have given a good impression of the butterflies
of the West African rainforest zone, with the day in the Shai Hills savannah showing the
amazing contrast with the rainforest zone.
Most participants have placed their pictures at the disposal of Butterfly Conservation, Ghana
and they will gradually be integrated into the website on Ghana butterflies
(ghanabutterflies.uw.hu), as well as being used in publicity work and in popular science
articles. A number of excellent photographs of Hesperiidae, that really capture the “essence”
of some of these interesting butterflies, will be figured in Larsen’s proposed book on the
“Skipper Butterflies of the Afrotropical Region”, probably not this one…
Photo: David Dennis
And last but not least Torben B. Larsen and Szabolcs Sáfián would like to thank all
participants the enthusiasm for butterflies and the genuine interest in their conservation they
have expressed during the trip. It is very encouraging that butterfly-based eco-tourism might
prove to be a practical tool to save rainforests and other important habitats in tropical Africa.

1. Common Commodore - Precis pelarga; 2. Hewitson's Cream Pentila - Pentila hewitsonii; 3. Common
Pathfinder - Catuna crithea; 4. Common Orange Sprite - Celaenorrhinus galenus; 5. Common Grass
Yellow - Eurema hecabe; 6. Large Fairy Hairstreak - Hypolycaena antifaunus; 7. Demon Charaxes -
Charaxes etheocles; 8. Abadima Acraea - Acraea pseudegina (all photos by David Dennis)

BUTTERFLIES RECORDED DURING THE 2008 BUTTERFLY TOUR ORGANIZED
BY BUTTERFLY CONSERVATION, GHANA
Torben B. Larsen, November, 2008
Numbers: The numbering refers to the book Butterflies of West Africa – origins, natural history, diversity, conservation –
Missing numbers belong to West African species not found in Ghana.
Legends: Black columns are the present status of the visited localities in Larsen database,
including records from the present trip
SHH denotes actual records; shh denotes almost certain to occur; ooo denotes might possibly occur;
–– denotes very unlikely to occur in the habitat
SHH = Shai Hills Resource Reserve TOTAL LIST
SHA = tour group in Shai Hills 27.x
SHI = subsequent Butterfly conference visit 7.xi
ABU = Aburi Botanical Gardens TOTAL LIST
ABG = tour group in Aburi 28.x
ABB = subsequent Butterfly conference visit 8.xi
ATE = Atewa Range Forest Reserve TOTAL LIST
ATW = tour group on Atewa 29/30.x
BOB = Bobiri Butterfly Sanctuary TOTAL LIST
BBI = tour group in Bobiri 31.x & 1.xi
BBF = subsequent Butterfly conference visit 10-12.xi
KAK = Kakum National Park TOTAL LIST
KAK = tour group in Kakum (KAA = Kakum Abrafo HQ: KAS = Simon's compound) 2/3.xi
ANK = Ankasa/Nini Suhien TOTAL LIST
ANN = tour group in Ankasa 4/5.xi
$ = recorded by tour members; £ = recorded subsequently to the tour group departure
SHA new locality to the Ghana recording scheme

The following three-letter codes are used for rough ecological classifications
WEF implies that the species is centered on Wet Evergreen Forest
MEF implies that the species is centered on Moist Forests
implies that the species is centered on Drier Semi-deciduous and marginal
DRF
forests
ALF implies that the species is found in any type of forest
GUI implies that the species is centered on the Guinea Savannah
SUD implies that the species is centered on the Sudan Savannah
SPE implies that the species is found in special habitats
species that are practically ubiquitous through all habitats in most of
UBQ
Africa
The species are roughly graded by rarity, though this is always a difficult call to make. Very rare species may one day be numerous
in a single locality. Very common butterflies are sometimes absent. However, the following notations are used:
VC = very common – species that are usually found on any visit to a suitable locality
CO = common – species that are usually found on 75% of visits to most suitable localities
NR = not rare – met with frequently but often not common
RA = rare – species that are usually found on less than 25% of visits to most suitable localities
VR = very rare – species that are usually found on less than 5% of visits to most suitable localities
tbl = species collected by T.B. Larsen in Ghana col = species collected by colleagues since 1993
# = known from the protected areas system ~ = not known from protected areas system
NOTE Subspecies are hidden in column C: authors and dates of description are found in Larsen (2005).

LOCALITIES ALL SHH TOUR CON ABU TOUR CON ATE TOUR BOB TOUR BBF KAK TOUR TOUR ANK TOUR
PAPILIONIDAE
PAPILIO
1 antimachus –– –– ATE BOBd KAK? ooo
2 zalmoxis –– –– ooo –– ooo ank
4 dardanus $ SHH SHA SHI ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
5 phorcas –– ABU ABG ATE ooo KAK ank
7 horribilis –– –– ATE ATW –– KAK ANK
9 chrapkowskoides –– ABU ATE BOBd BBI BBF KAK ANK
10 sosia $ SHH8 SHA SHI ABU ATE ATW BOBd BBI KAK ANK
11 nireus $ SHH SHA SHI ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
12 menestheus $ –– SHA SHI ABU ABG ATE ATW BOB5 BBI BBF KAK KAS ANK ANN
13 demodocus $ SHH SHA SHI ABU ABG ABB ATE BOB5 BBI BBF KAK KAA KAS ANK ANN
15 cyproeofila $ –– ABU ATE ATW BOB5 BBI BBF KAK KAS ANK ANN
16 zenobia $ –– –– ATE ATW BOB5 BBI KAK ANK
17 nobicea –– –– –– –– –– ––
18 cynorta $ –– ABU ABG ATE ATW BOBd BBF KAK ANK
GRAPHIUM
20 angolanus $ SHH ABU+ ATE BOB7 KAK ooo ANN
22 tynderaeus $ –– –– ATE –– KAK ANK ANN
23 latreillianus $ –– –– ATE ATW ooo KAK ANK
24 almansor –– –– –– –– –– ––
25 adamastor shh ABU ooo ooo –– ––
26 agamedes –– ABUj –– ooo –– ––
28 rileyi –– –– ATE –– ooo ank
29 leonidas $ SHH SHA SHI ABU ATE ATW BOB5 KAK ANK ANN
30 illyris $ –– –– ATE ATW –– KAK ank
31 policenes $ SHH SHA SHI ABU ABG ATE ATW BOBo BBI BBF KAK KAA KAS ANK ANN
32 liponesco £ –– abu ate BOB5 BBF KAK ANK
34 antheus $ SHH8 SHA SHI ABU ATE BOBo KAK ank
PIERIDAE
PSEUDOPONTIINAE
PSEUDOPONTIA
35 paradoxa $ –– –– ooo –– kak ANK ANN

COLIADINAE
CATOPSILIA
36 florella $ SHH SHA SHI ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
EUREMA
38 senegalensis $ –– ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAS ANK ANN
39 hecabe $ SHH SHI ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAS ANK ANN
40 floricola $ –– ABU+ ABG ATE ATW BOB5 BBI KAK ANK
41 hapale –– –– ate BOBd kak ank
42 desjardinsii SHH ABU ate bob KAK ank
43 brigitta $ SHH SHA SHI ABU ABG ATE BOB5 BBI KAK KAA KAS ANK
PIERINAE
PINACOPTERYX
44 eriphia –– –– –– –– –– ––
NEPHERONIA
45 argia $ ooo ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAA KAS ANK
46 thalassina $ shh ABU+ ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK
47 pharis $ –– ABU+ ATE ATW BOB5 BBF KAK KAS ANK
COLOTIS
54 vesta –– –– –– –– –– ––
57 celimene –– –– –– –– –– ––
58 ione –– –– –– –– –– ––
60 danae –– –– –– –– –– ––
61 aurora ooo –– –– –– –– ––
62 antevippe SHH –– ooo ooo –– ––
63 euippe $ SHH SHA SHI ABU ABG ATE ATW BOB5 BBI BBF KAK ANK
65 evagore $ SHH SHA SHI ooo ooo ooo –– ––
BELENOIS
68 aurota shh abu ooo BOB6 kak ank
69 creona $ SHH ABU ABG ate bob KAK ank
70 gidica SHH –– ooo –– –– ––
72 subeida –– –– –– –– –– ––
73 calypso $ SHH SHA SHI ABU ABG ATE ATW BOB5 BBI BBF KAK KAS ANK
74 theora –– ABU ATE BOB5 BBI KAK ANK
76 hedyle $ SHH8 SHA SHI ooo ATE bob KAK ooo

DIXEIA
78 doxo –– –– –– –– –– ––
79 orbona –– –– –– –– –– ––
80 cebron £ –– abu ooo BOB5 BBF KAK ooo
81 capricornus £ –– ooo ooo BOB5 BBF kak ooo
APPIAS
84 sylvia $ SHH SHI ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
85 phaola –– –– ATE bob KAK KAS ANK
86 sabina $ –– ABU ATE ATW BOBo BBF KAK ANK
87 epaphia $ SHH SHI ABU ABG ate BOB6 BBF KAK ANK
LEPTOSIA
88 alcesta $ SHH SHI ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAA KAS ANK ANN
90 hybrida $ SHH+ ABU+ ABG ATE ATW BOBo BBI BBF KAK KAS ANK
91 medusa $ –– ABU ABG ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
92 marginea $ –– –– ATE ATW bob KAK ANK
93 wigginsi $ SHH SHA abu ate BOB6 kak ANK
MYLOTHRIS
95 chloris $ SHH SHA SHI ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAS ank
100 dimidiata $ –– –– ATE ATW ooo KAK ANK
103 aburi –– ABU ABG ABB –– –– –– ––
106 poppea $ SHH ABU ABG ABB ATE ATW BOB5 BBF KAK KAA KAS ANK
107 spica –– ABU ABG ABB ATE BOBd BBF KAK ank
109 rhodope $ SHH ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAS ANK
110 jaopura $ –– ABU ABG ABB ATE ATW BOBd BBF KAK KAS ANK
111 schumanni $ –– ABU ATE BOB6 BBI BBF KAK ANK
112 atewa –– –– ATE –– –– ––
LYCAENIDAE
MILETINAE
EULIPHYRA
114 hewitsoni –– –– ate BOB6 kak ANK
115 mirifica –– –– ate –– KAK ––
116 leucyania –– ABUi ate BOB7 kak ank
ASLAUGA
117 ernesti –– –– –– –– –– ––
118 marginalis $ –– ABU ATE+ BOB6 BBF KAK ANK8 ANN

121 lamborni –– –– ATE –– kak ank
121a cf prouvosti 1 ––
121b cf prouvosti21 ––
124 imitans –– –– –– –– –– ––
MEGALOPALPUS
127 zymna $ –– –– ATE BOB+ BBI BBF KAK ANK
129 metaleucus –– –– ATE BOB5 BBI BBF KAK ANK
SPALGIS
130 lemolea $ SHH8 SHA SHI ABU ABG ATE BOB5 BBI BBF KAK ank
LACHNOCNEMA
131 vuattouxi ooo abu ATE bob KAK ank
133 emperanus –– abu ate bob kak ANK
135 disrupta –– –– ??? ooo ooo ooo
136 reutlingeri –– –– ATE –– ooo ank
137 luna –– –– –– –– –– ––
139 albimacula –– –– ??? –– ooo ooo
LIPTENINAE
PTELINA
141 carnuta –– –– ATE BOB5 KAK ANK
PENTILA
142 pauli $ –– ABU ABG ABB ATE BOBs BBI BBF KAK ooo
144 petreoides –– –– ATE –– –– ooo
147 petreia $ –– ABU+ ABG ABB ATE ATW BOB+ BBF KAK ANK ANN
148 condamini –– –– ooo –– kk ank
152 picena $ –– –– ATE ATW BOB+ BBF KAK ANK
155 phidia –– ooo ATE BOB5 KAK ooo
157 hewitsonii $ –– –– ATE ATW BOB+ kak ank
TELIPNA
159 acraea –– –– ATE bob KAK ANK
160 semirufa £ –– –– ATE BOB6 BBF KAK ANK
161 maesseni –– –– –– –– –– ––
ORNIPHOLIDOTOS
168a maesseni –– –– ate –– ooo ooo
170 nigeriae now sylviae –– –– ATE bob kak ank
171 onitshae –– –– ATE bob KAK ank

172 irwini –– –– ATE bob kak ank
173 issia $ –– –– ATE ??? ooo ??? kak ank
174 tiassale £ –– –– ATE BOB8 BBF KAK ANK
175 nympha –– –– ATE bob KAK ank
TORBENIA
177 wojtusiaki $ –– –– ATE BOB+ BBI BBF KAK ANK
MIMACRAEA
179 neurata $ –– ABU8 ABG ATE BOB8 BBI BBF KAK KAS ank
181 darwinia –– –– ATE bob KAK ank
182 maesseni –– –– –– –– –– ––
MIMERESIA
184 libentina $ –– ABU ABG ATE BOB5 BBI KAK KAA KAS ANK
185 moyambina –– –– ATE ooo kak ank
186 debora –– –– ooo ooo ooo ooo
187 semirufa –– –– ATE BOB+ BBF KAK ANK
190 cellularis $ –– –– ATE ATW ooo kak ank
191 issia –– –– ATE ooo KAK ANK
PSEUDERESIA
192 eleaza –– –– ATE BOB+ KAK ank
ERESIOMERA
193 bicolor $ –– ABU ABG ABB ATE BOB5 BBI BBF KAK ANK ANN
194 isca $ –– ooo ATE BOBs BBI KAK ank
195 jacksoni –– ooo ate –– kak ank
197 petersi –– –– ATE –– KAK ANK
CITRINOPHILA
199 marginalis $ –– ABU ABB ATE ATW BOB7 KAK ANK
200 similis $ –– ABU ATE ATW BOB5 BBI BBF KAK ANK
202 erastus $ –– –– ATE ATW BOBs BBI BBF kak ank
ERESINA
204 maesseni abu ate BOB6 kak ank
206 pseudofusca $ –– ABU ABG ooo bob kak ank
210 saundersi –– ooo ooo ooo kak ank
212 theodori $ –– ABU ABG ABB ate BOB+ BBI kak ank
ARGYROCHEILA
213 undifera –– ABU ATE bob KAK ANK

LIPTENA
216 submacula $ –– ooo ATE ATW BOB5 BBI BBF KAK TBL ANK
217 griveaudi –– –– ATE –– ooo ank
218 simplicia $ –– ABU ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
222 tiassale £ –– ABU ooo –– ooo ooo
224 albicans –– –– ATE ooo kak ank
225 alluaudi $ –– –– ATE ATW BOB5 BBI BBF KAK KAS KAS ank
226 fatima –– –– ooo ooo ooo ooo
227 pearmani –– –– –– –– –– ––
229 ferrymani –– –– –– –– –– ––
231 septistrigata $ –– ABU ABG ABB ATE BOB6 BBF –– ––
232 evanescens –– –– ate ooo kak ank
234 xanthostola £ –– –– ATE BOB7 BBF KAK ANK
236 bia –– –– ATE ooo KAK ooo
237 flavicans £ –– ooo ATE ooo kak ank
239 seyboui –– –– ooo –– ooo ank
240 similis £ –– –– ATE ooo kak ank
242 helena $ –– –– ATE ATW bob KAK ANK
243 catalina –– –– ATE bob KAK ANK
KAKUMIA
246 otlauga –– –– ATE BOB8 BBF KAK ANK
FALCUNA
249 leonensis $ –– ooo ATE ATW BOB5 BBI BBF KAK KAS ANK
252 campimus –– –– ATE BOB7 KAK ANK ANN
TETRARHANIS
254 symplocus $ –– ABU+ ATE ATW BOB5 BBI BBF KAK KAA KAS ANK
255 baralingam $ –– –– ATE ATW ooo KAK KAA ANK ANN
260 stempfferi –– –– ATE bob ooo ooo
LARINOPODA
264 aspidos –– –– –– –– –– ––
265 eurema $ –– ABU+ ATE ATW BOB5 BBI BBF KAK KAA KAS ANK
MICROPENTILA
266 adelgitha $ –– –– ATE ATW bob KAK ANK ANN
267 adelgunda –– –– ate bob kak ank
268 dorothea –– –– ATE bob kak ank

270 brunnea –– –– ATE bob KAK ANK
270a sp nov cf katerae ANK
275 mamfe –– ooo ooo –– ooo ank
IRIDANA
278 incredibilis £ –– ABU ate bob kak ank
279 ghanana –– –– –– –– –– ––
280 exquisuta –– –– ate ooo kak ANK
281 nigeriana £ –– ABU ABB ate ooo ooo ank
282 hypocala –– –– ooo –– ––
HEWITSONIA
283 boisduvalii $ –– ABU+ ATE BOBs BBI BBF KAK ANK
284 occidentalis –– ABU ate bob kak ANK
286 inexpectata $ –– ABU ABG ABB ATE BOBs BBI BBF KAK6 KAS ANK
CERAUTOLA
289 crowleyi $ –– ABU ABG ate bob kak ank
291 ceraunia $ –– ABU ate BOB6 BBI BBF KAK ank
293 miranda $ –– ABU ABG ABB ate BOB6 BBI kak ank
EPITOLA
294 posthumus $ –– ABU ATE ATW BOBs kak ank
295 uranoides £ –– ooo ATE BOB5 BBF KAK ank
296 urania –– ooo ATE BOB6 kak ank
CEPHETOLA
297 cephena £ –– abu ate BOB8 BBF kak ank
299 pinodes –– ooo ate ooo kak ank
300 subcoerulea –– ABU ATE+ ooo ooo ooo
301 nigra –– –– ate 2008 ooo ank
302 mercedes –– ABUs ATEs –– ooo ank
303 obscura –– ABU ATE bob KAK ank
305 sublustris £ –– ABU ooo bob ooo ooo
306 maesseni –– –– ooo –– –– ––
307 collinsi £ –– ABU ABB –– ooo ooo ank
HYPOPHYTALA
308 hyettoides $ –– ABU ABG ABB ate bob KAK ank
310 hyettina –– ooo ATE BOB6 kak ank
311 henleyi –– –– ate BOB6 kak ank

312 benitensis –– –– ate BOB6 kak BIA
PHYTALA
314 elais –– –– ATE bob KAK ank
GERITOLA NEW albomaculata ATE
315 gerina $ –– ABU ooo BOB8 BBI KAK ank
320 virginea $ –– ABU ate BOB8 BBI KAKs ank
STEMPFFERIA
322 cercene –– –– ate BOBs KAK ank
324 moyambina £ –– –– ATE bob KAK ank
326 dorothea $ –– ooo ate BOBs BBI kak ank
330 leonina $ –– abu ate BOB5 BBI KAK ank
334 ciconia £ –– –– ATE BOB8 ?BBF kak ank
335 zelza –– –– ate ooo kak ooo
340 michelae $ –– abu ATE ATW BOBs TBL KAK ank
340a cf michelae bob KAK
342 kholifa –– –– ate BOB+ kak ank
344 staudingeri $ –– ABU ATE BOB8 BBI kak ank
AETHIOPANA
346 honorius $ –– abu ATE ATW BOB6 BBI KAK ANK
EPITOLINA
347 dispar $ –– abu ATE ATW BOB5 BBF KAK ANK
348 melissa $ –– ooo ATE ATW bob KAK KAS ANK
350 catori £ –– ABU ATE BOBs BBF kak ank
NEAVEIA
352 lamborni $ –– ABU ABG ABB ATE@ bob kak ank
THECLINAE
MYRINA
354 silenus SHH abu ooo bob KAK ANK
355 subornata shh –– –– –– –– ––
OXYLIDES
356 faunus $ –– ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAS ANK ANN
DAPIDODIGMA
359 hymen $ shh abu ate BOB6 BBF KAK KAS ANK ANN
360 demeter –– ooo ATE BOBh kak ANK

APHNAEUS
361 orcas £ –– abu ate BOBs BBF KAK ANK
362 argyrocyclus –– –– ooo ooo KAK ank
363 asterius –– ABU ATE ooo KAK ank
364 brahami shh –– –– –– –– ––
365 jefferyi –– –– ooo –– kak ank
366 charboneli –– –– ooo –– kak ank
367 gilloni –– –– ooo –– ooo ooo
APHARITIS
368 nilus shh –– –– –– –– ––
SPINDASIS
369 mozambica SHH abu ate bob KAK ank
370 avriko ooo –– –– –– –– ––
371 crustaria –– –– –– –– –– ––
372 iza –– ABU ATE BOB7 kak ank
373 menelas –– –– ate ooo ooo ank
ZERITIS
374 neriene ooo ooo ooo ooo ooo ooo
AXIOCERSES
375 harpax $ SHH ABU ATE bob KAK KAS ANK
377 amanga ooo –– –– –– –– ––
LIPAPHNAEUS
378 leonina –– –– ATE bob KAK ank
379 aderna –– abu ooo ooo ooo ooo
PSEUDALETIS
380 agrippina –– ABU7 ATE@ ooo ooo ank
386 zebra –– –– –– –– ooo ooo
388 jolyana –– –– ATE@ bob ooo ooo
391 leonis –– ABUj ate BOB6 KAK ank
IOLAUS
Subgenus Iolaus
392 eurisus $ –– ABU ATE bob KAK KAS ANK
Subgenus Iolaphilus
393 menas ooo –– –– –– –– ––
395 carolinae –– ooo ate ooo ooo ooo

397 iulus $ –– ABU ABG ABB ATE BOB7 KAK ank
Subgenus Philiolaus
398 ismenias shh abu ooo –– –– ––
400 alcibiades £ –– ABU ABB ate bob 2008 ank
401 parasilanus $ –– ABU8 ABG ABB –– ooo –– ––
402 paneperata –– –– ATE BOB kak ank
403 lukabas –– –– ate ooo ooo ooo
404 mane –– –– ATE bob ooo ank
405 theodori –– –– –– –– –– ––
406 likpe $ –– –– –– BOB6 BBI –– ––
407 calisto –– ABU ate bob KAK ank
408 laonides –– –– ooo ooo kak ank
Subgenus Tanuetheira
410 timon –– –– ATE BOB7 kak ank
Subgenus Epamera
411 alienus –– –– –– –– –– ––
414 scintillans ooo –– –– –– –– ––
415 laon £ –– abu ABB ooo bob kak ank
418 banco –– –– –– –– ooo ank
426 sappirus –– –– ooo BOB7 kak ANK
428 bellina –– abu ate BOB+ kak ank
432 fontainei –– –– –– –– –– ––
434 aethria $ –– ooo ABB ATE ATW BOBh KAK8 KAS ank
435 farquharsoni –– –– ate ooo kak ank
436 iasis £ SHH ABU ate BOB7 BBF KAK bia
437 maesa –– ABU ate BOBh ooo ooo
ETESIOLAUS
439 catori –– –– ate bob kak ANK
440 kyabobo –– –– ooo ooo ooo ooo
STUGETA
441 marmoreus shh –– –– –– –– ––
HYPOLYCAENA
443 philippus $ SHH8 SHI ABU ATE bob kak ANK
444 kadiskos –– –– ATE ooo ooo ooo
445 liara –– abu ATE BOB6 kak ank

446 lebona $ –– abu ATE BOB+ BBF KAK KAS ank
447 clenchi $ –– ooo ATE ATW BOBo KAK ANK
449 scintillans $ –– ABU ABG ABB ATE ATW BOB+ BBI KAK KAS ANK
450 dubia $ –– ABU ABG ATE BOB+ BBI KAK KAS ANK
451 kakumi £ –– ooo ATE BOB+ KAK TBL ANK
452 antifaunus $ –– ABU ATE BOBd BBI BBF KAK ank
453 hatita £ –– ABU ATE BOB5 BBF KAK ANK
455 nigra –– ABU ATE BOBd KAK ank
PILODEUDORIX
457 camerona $ SHH* ABU ABB ATE BOB+ BBI BBF KAK KAS ank
458 diyllus –– abu ATE bob kak ank
460 caerulea shh ABU ate bob kak ank
461 zela –– abu ATE ooo kak ank
462 catori –– abu ooo ooo kak ank
467 otraeda –– –– ATE bob KAK ANK
468 leonina –– –– ATE bob KAK ank
469 virgata –– –– ATE ooo KAK ANK
473 deritas –– –– ate ooo ooo ooo
474 aucta –– –– –– –– –– ––
475 pseudoderitas –– ooo ate bob kak ank
476 laticlavia –– –– ATE ooo kak ank
477 aurivilliusi –– ABU ATE ooo kak ank
478 kiellandi –– –– ATE ooo kak ank
479 corruscans –– –– ooo –– KAK ank
480 violetta –– ooo ATE bob KAK ank
481 fumata –– –– ooo –– ooo ooo
PARADEUDORIX
484 eleala £ –– ABU ATE BOB+ BBI BBF KAK ANK
485 petersi
487 moyambina –– –– ATE ooo kak ank
HYPOMYRINA
491 mimetica –– –– ate ooo kak ooo
492 nomion SHH ABU ate bob kak ooo
DEUDORIX
494 antalus $ SHH ABU ATE BOB8 BBI KAK ank

495 livia ooo –– –– –– –– ––
496 lorisona $ SHH8 SHA ABU ATE ATW BOB5 BBF KAK ank
497 kayonza –– –– ATE bob KAK ank
498 dinochares ooo ooo ooo bob ooo ooo
499 dinomenes –– –– ate BOB7 kak ank
500 odana –– ooo ATE bob KAK ank
501 galathea –– ABU ATE bob KAK ANK
502 caliginosa –– –– ATE ooo kak ank
CAPYS
506 vorgasi –– –– –– –– –– ––
POLYOMMATINAE
ANTHENE
507 rubricinctus $ –– ooo ATE BOB+ BBI BBF KAK KAA ANK ANN
508 ligures –– –– ATE+ BOB+ kak ank
510 sylvanus –– ABU+ ATE BOB5 KAK ANK
512 liodes $ –– abu ATE BOB+ BBI BBF KAK KAA ANK
513 definita ooo ooo ATE ooo KAK ooo
514 princeps ooo ABU ATE BOB7 kak ooo
515 starki ooo ooo –– –– –– ––
516 amarah shh –– ooo –– ooo ooo
517 lunulata $ shh abu ATE bob KAK ooo
518 kikuyu shh ooo ooo –– –– ––
519 talboti ooo –– –– –– –– ––
520 wilsoni –– –– –– –– –– ––
521 levis –– ABU ate bob KAK ank
522 irumu –– ABU ate BOB+ kak ank
523 larydas $ ooo ABU ABG ATE ATW BOB5 BBI BBF KAK KAA KAS ANK
524 crawshayi $ shh ooo ABG ATE ooo ooo ooo
525 lachares $ –– –– ATE BOBd BBI BBF KAK ank
527 lysicles £ –– ABU+ ATE BOBs BBF KAK ANK
530 atewa –– –– ATE ooo kak ANKs
532 radiata –– –– ATE –– kak ank
534 locuples £ –– –– ate BOBh BBF kak ank
535 mahota

537 scintillula –– –– ATE ooo ooo ank
538 helpsi –– –– ATE –– –– ––
539 juba –– –– ATE ooo KAK ANK
NEURYPEXINA
540 lyzanius –– –– ATE BOB6 BBF KAK ANK
NEURELLIPES
542 lusones $ –– –– ATE ooo KAK ANK ANN
543 chryseostictus $ –– –– ATE ATW BOB8 BBF KAK ank
544 fulvus –– –– ATE –– KAK ANK
545 staudingeri –– –– ate –– ooo ank
546 gemmifera –– –– ooo ooo ooo ooo
TRICLEMA
547 rufoplagata –– –– ooo –– ooo ank
548 lucretilis –– –– ATE BOB7 kak ank
549 lamias –– abu ate BOB6 kak ank
550 fasciatus –– –– ate bob KAK ank
551 obscura –– –– ate –– kak ank
552 inconspicua –– –– ate ooo KAK ank
554 hades ooo abu ATE BOB+ kak ank
555 phoenicis shh ooo ooo BOB7 –– ––
556 nigeriae shh ABU+ ATE bob ooo ooo
CUPIDESTHES
560 jacksoni –– –– ATE ooo KAK ANK
561 mimetica $ –– –– ??? –– KAKs KAS ooo
562 lithas $ –– ABU ATE ATW BOB5 BBF KAK KAS ANK
564 leonina –– –– ATE BOBs KAK ank
564 pungusei –– –– ooo –– KAK ooo
PSEUDONACADUBA
565 sichela $ SHH8 SHI –– ATE BOB+ BBI BBF KAK ANK
LAMPIDES
567 boeticus SHH SHA SHI abu ate bob KAK ANK
URANOTHAUMA
568 falkensteini $ ooo abu ATE ATW BOB5 BBI BBF KAK KAA KAS ANK

PHLYARIA
574 cyara $ –– –– ATE BOB6 KAK ANK8 ANN
CACYREUS
575 lingeus $ SHH ABU ABG ABB ATE BOB5 KAK KAA KAS ank
577 audeoudi $ –– –– ate BOB8 BBI KAK8 KAA KAS ank
LEPTOTES
578 pirithous $ SHH SHA SHI ABU ABG ATE BOB5 BBI BBF KAK ANK
579 babaulti shh abu ate BOB@ KAK ank
580 jeanneli shh abu ate bob kak ank
581 brevidentatus SHH abu ate bob kak ank
582 pulchra ooo –– ooo ooo ooo ooo
TUXENTIUS
583 cretosus ooo –– –– –– –– ––
584 carana $ –– abu ATE ATW BOB5 BBI BBF KAK ANK
TARUCUS
586 ungemachi shh –– –– –– –– ––
588 rosacea –– –– –– –– –– ––
ACTIZERA
592 lucida –– –– –– ooo ooo ooo
EICOCHRYSOPS
593 hippocrates $ SHH SHA SHI ABU ATE BOB6 KAK KAA KAS ANK
594 dudgeoni ooo –– –– –– –– ––
CUPIDOPSIS
595 jobates SHH+ ooo –– –– –– ––
596 cissus shh ooo ATE ooo ooo ANK
EUCHRYSOPS
598 albistriata shh abu ooo ooo KAK ooo
600 reducta ooo –– –– –– –– ––
601 malathana $ SHH SHA SHI ABU+ ABG ABB ATE BOB5 KAK KAA ANK
604 osiris $ SHH SHA abu ATE BOB6 KAK ank
605 barkeri ooo –– ooo BOBo –– ––
606 sahelianus ooo –– –– –– –– ––
LEPIDOCHRYSOPS
607 victoriae shh –– –– –– –– ––
608 parsimon –– ooo –– bob ooo ooo

611 synchrematiza –– –– –– ooo ooo ooo
615 quassi –– –– ooo ooo ooo ooo
THERMONIPHAS
617 micylus $ –– ABU ATE BOB+ BBI BBF KAK KAS ANK ANN
OBORONIA
622 punctatus –– ooo ATE BOB6 KAK ANK
623 liberiana $ –– –– –– –– KAK ANK ANN
624 pseudopunctatus £ –– –– –– –– BBF –– ––
625 guessfeldti $ –– ABU8 ABG ABB ATE ATW BOB5 BBI KAK ooo
626 ornata $ –– ooo ATE BOB5 BBI BBF KAK KAA KAS ANK
AZANUS
627 ubaldus –– –– –– –– –– ––
628 jesous shh ooo –– –– –– ––
629 moriqua –– ooo –– –– –– ––
630 mirza $ shh ABU ATE BOB5 BBI KAK KAA KAS ANK
631 natalensis –– ooo –– ooo –– ––
632 isis –– abu ATE bob KAK ANK
CHILADES
633 eleusis –– –– –– –– –– ––
634 trochylus $ SHH SHA SHI abu ooo –– ooo ooo
ZIZEERIA
635 knysna $ SHH SHA SHI ABU ABG ABB ATE BOB5 BBF KAK KAA ANK
ZIZINA
636 antanossa $ SHH8 SHA ABUs ABG ate bob KAK ANK
ZIZULA
637 hylax £ shh ABU ATE+ BOB5 BBF KAK ANK
RIODINIDAE
ABISARA
638 intermedia –– –– ate –– ooo ooo
639 tantalus –– –– ate –– ooo ooo
642 gerontes –– –– ATE bob ooo ooo

NYMPHALIDAE
LIBYTHEINAE
LIBYTHEA
646 labdaca $ –– ABU ATE ATW BOB5 BBI BBF KAK ANK
DANAINAE
DANAUS
647 chrysippus $ SHH SHA SHI ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
TIRUMALA
648 petiverana $ SHH ABU ATE ATW BOB5 BBI BBF KAK TBL ank
AMAURIS
650 niavius $ shh ABU ATE ATW BOBo BBI BBF KAK ANK
651 tartarea $ SHH ABU ATE ATW BOB5 BBF KAK KAA ANK
652 hecate $ ooo ABU ABG ATE BOB5 BBF kak ANK
653 damocles $ shh ABU ATE BOBd BBI BBF KAK ANK
SATYRINAE
GNOPHODES
656 betsimena $ –– ABU ABB ATE ATW BOBj BBI BBF KAK KAA KAS ank
657 chelys $ –– ABU ATE BOBj BBI BBF KAK KAS ANK
MELANITIS
658 leda $ shh SHA ABU ATE ATW BOBj BBF KAK ANK
659 libya ooo ABU+ ABG ate BOBj BBI KAK KAS ANK
ELYMNIOPSIS
661 bammakoo –– ABU ABB ATE BOBj KAK TBL ANK
BICYCLUS
663 xeneas $ –– ooo ATE ATW BOBj BBI BBF KAK ANK
665 evadne $ –– –– ATE bob KAK KAS ANK ANN
669 ephorus –– –– ATE ooo KAK ank
672 italus –– –– –– –– –– ––
673 zinebi –– abu ATE BOBj KAK ANK
674 uniformis £ –– –– ATE BOB+ KAK8 GAB ank
678 procora –– –– ATE BOBj KAK ANK
679 pavonis –– –– –– –– –– ––
680 milyas –– –– –– –– –– ––
681 trilophus –– –– ATE ooo ooo ANK
682 ignobilis £ –– –– ATE BOBo kak ANK

683 maesseni £ –– –– ATE BOBs BBF ooo ooo
684 nobilis $ –– –– ATE ATW ooo kak ANK ANN
687 taenias $ –– ABU ABG ABB ATE ATW BOBj BBI BBF KAK TBL ANK
690 vulgaris shh ABU ATE BOBj KAK ANK
691 dorothea $ –– ABU ABG ABB ATE ATW BOBj BBI BBF KAK KAA KAS ANK ANN
692 sandace $ –– ABU ABG ABB ATE ATW BOBj BBI BBF KAK KAA KAS ANK
693 sambulos –– –– ATE BOB+ KAK ANK
694 sangmelinae $ –– –– ATE ATW BOBj BBI BBF KAK ANK
695 mandanes –– ooo ATE ooo KAK ooo
696 auricruda $ –– ooo ATE+ ATW BOBk kak ank
697 campa –– –– –– –– –– ––
698 angulosa shh –– –– BOB6 –– ––
699 sylvicolus –– –– ATEk –– KAK ––
700 abnormis –– –– ATE BOBj KAK ank
701 safitza $ SHH SHA SHA ABU+ ABG 2008 BOBj BBI KAK8 GAB ooo
702 funebris $ shh ABU ATE ATW BOBj BBI BBF KAK ank
704 dekeyseri –– –– ATE –– KAK ank
705 istaris –– ABU ATE BOB6 ooo ooo
707 madetes $ –– ABU ABG ABB ATE ATW BOBj BBI BBF KAK KAS ANK
709 martius $ –– ABU ABG ATE ATW BOBj BBI BBF KAK ANK ANN
HALLELESIS
712 halyma –– –– ATE BOBj KAK ANK
HENOTESIA
713 elisi –– –– –– –– –– ––
HETEROPSIS
714 peitho $ –– ABU ATE ATW –– ooo ank
YPTHIMA
715 asterope ooo –– –– –– –– ––
716 condamini $ SHH SHA SHI –– –– –– –– ––
717 antennata shh –– –– –– –– ––
718 vuattouxi ooo abu ate ooo –– ––
719 doleta $ ooo ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
721 pupillaris ooo –– –– –– –– ––
722 impura ooo –– –– –– –– ––

YPTHIMOMORPHA
724 itonia $ SHH8 SHA SHI –– –– BOBj BBF kak ank
CHARAXINAE
CHARAXES
725 varanes $ SHH SHI ABU ATE BOB5 KAK KAS ANK
726 fulvescens $ –– ABU ABG ABB ATE ATW BOBy BBF KAK ANK
728 candiope ooo abu ATE ooo 2008 ooo
729 protoclea $ SHH ABU8 ABG ABB ATE ATW BOBy BBI BBF KAK ANK
730 boueti $ –– ABU ATE BOB5 KAK ANK ANN
731 cynthia $ –– ABU ABG ATE ATW BOBy BBI BBF KAK ANK
732 lucretius $ –– ABU ABG ATE BOBy BBI KAK ANK ANN
733 lactetinctus ooo –– –– –– –– ––
734 epijasius £ SHH SHI –– ATE ooo –– ––
736 castor $ SHH abu ATE BOB5 KAK ANK
737 brutus $ SHH ABU ABG ATE BOBy BBI BBF KAK ANK
738 pollux –– –– ATE –– ooo ooo
740 eudoxus –– –– ooo –– ooo ooo
741 tiridates $ SHH SHA SHI ABU ABB ATE BOBy BBF KAK KAA KAS ANK
742 bipunctatus $ –– –– ATE BOBy BBI KAK ANK
743 numenes $ SHH ABU ATE BOBy BBI BBF KAK KAS ANK
744 smaragdalis £ –– –– ATE bob kak ANK
745 imperialis –– –– ATE ooo kak ANK
746 ameliae $ –– –– ATE ATW BOBo KAK ANK
747 pythodoris –– –– ooo –– ooo ooo
748 hadrianus $ –– –– –– –– ooo ANK ANN
750 nobilis –– –– ATE –– –– ank
752 fournierae –– –– ATE –– –– ooo
753 zingha $ –– ABU ATE ATW BOB5 KAK ANK
754 etesipe $ –– ABU+ ABG ABB ATE BOBy BBI BBF KAK KAA KAS ANK
755 achaemenes SHH ABU ATE ooo –– ––
756 eupale $ –– ABU ATE ATW BOBy BBI BBF KAK ANK ANN
757 subornatus £ –– –– ATE BOB5 BBI BBF kak ank
758 anticlea –– ooo ATE BOBy KAK ank
759 hildebrandti $ –– –– ATE BOBy BBI kak ank

760 etheocles $ SHH ABU ATE BOBy BBI BBF KAK ANK
762 petersi –– –– ATE ooo kak ank
765 bocqueti –– –– ATE ooo kak ank
767 virilis £ –– ooo ATE BOB5 BBF KAK ank
768 cedreatis $ SHH ABU ATE BOBy BBI KAK ANK
769 plantroui –– –– ATE –– –– ––
770 viola ooo –– –– BOBh –– ––
771 northcotti ooo –– –– –– –– ––
772 pleione $ –– ABU ATE BOBy BBI BBF KAK KAS ANK
773 paphianus £ –– –– ATE BOBy BBF KAK ank
774 nichetes –– –– ATE bob kak ANK
775 porthos –– –– ATE BOBd kak ank
776 zelica £ –– –– ATE BOB5 BBF kak ANK
777 lycurgus $ –– –– ATE BOBy BBI BBF KAK ANK
778 mycerina –– –– ATE BOBy kak ANK
779 doubledayi –– –– ATE BOB5 kak ank
EUXANTHE
780 eurinome $ SHH ABU ABG ABB ATE BOBy KAK KAS ANK
PALLA
783 violinitens $ –– ooo ATE BOBy BBI KAK ank
784 decius £ –– ABU ABB ATE BOBy BBF KAK ank
785 ussheri $ –– abu ATE ATW BOBy BBF KAK ANK
786 publius –– ooo ATE BOBy KAK ank
APATURINAE
APATUROPSIS
786a cleochares –– –– ATE BOB+ KAK ank
NYMPHALINAE
KALLIMOIDES
787 rumia $ –– ooo ATE ATW BOBj BBI BBF KAK KAA ANK
VANESSULA
788 milca $ –– –– ATE ATW –– –– ––
ANTANARTIA
789 delius $ –– –– ATE ATW BOB5 BBI BBF KAK ANK ANN

VANESSA
791 cardui shh abu ate bob KAK ANK
PRECIS
792 octavia $ SHH8 SHA SHI ABU ABG ABB ate bob BBF kak ank
793 antilope $ shh SHI ABU8 ABG ate bob kak ooo
796 ceryne –– –– ooo ooo kak ooo
797 pelarga $ SHH SHA SHI ABU ABG ABB ATE ATW BOBd BBF KAK KAS ANK
798 sinuata –– –– ATE ooo KAK ANK
799 milonia –– –– ATE –– –– ––
HYPOLIMNAS
801 misippus $ SHH SHA SHI ABU ABG ABB ATE BOB5 BBF KAK KAA KAS ooo ANN
802 anthedon $ ooo ABU ABG ABB ATE ATW BOBj BBI BBF KAK KAA KAS ANK
803 dinarcha –– –– ATE bob KAK ANK
806 salmacis $ –– ABU ABG ABB ATE ATW BOBj BBI BBF KAK KAA KAS ANK ANN
SALAMIS
808 cacta $ –– ABU ABG ATE BOB5 BBF KAK ANK
PROTOGONIOMORPHA
809 cytora $ –– abu ATE ATW BOB5 BBI BBF KAK KAA ANK
811 parhassus $ –– ABU+ ABG ABB ATE ATW BOB5 BBI BBF KAK KAA TBL ANK ANN
812 anacardii $ –– SHI ABU ABG ABB ATE+ BOB+ KAK ooo
JUNONIA
813 orithya SHH SHI –– ooo ooo ooo ooo
814 oenone $ SHH ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA ANK
815 hierta SHH abu ooo ooo ooo ooo
816 cymodoce $ –– ABU ATE BOB5 BBI KAK ANK
817 westermanni –– –– ATE ooo ooo ank
818 hadrope –– –– –– –– –– ––
819 sophia $ SHH ABU8 ABG ABB ATE ATW BOBo BBI BBF KAK ANK ANN
820 stygia $ –– –– ATE ATW BOBo BBI BBF KAK KAA KAS ANK ANN
822 chorimene $ SHH SHI ABU ABG ABB ooo BOB6 BBF ooo KAA KAS ooo
823 terea $ SHH ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAA KAS ANK ANN
CATACROPTERA
824 cloanthe $ SHH8 SHA ABU ate ooo ooo ooo

CYRESTINAE
CYRESTIS
825 camillus $ –– ABU+ ATE ATW BOB5 BBF KAK ANK
BIBLIDINAE
BYBLIA
826 anvatara $ SHH SHA DFI ABU+ ATE ATW BOBo BBI BBF KAK ANK
827 ilithyia –– –– –– –– –– ––
MESOXANTHA
828 ethosea $ –– –– ATE ATW BOB5 BBI KAK ANK
ARIADNE
829 enotrea $ –– ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
830 albifascia $ –– ABU ABG ATE BOBb KAK ANK
NEPTIDOPSIS
833 ophione $ –– ABU ABG ABB ATE ATW BOBo BBI BBF KAK ANK
EURYTELA
834 dryope $ SHH SHA SHI ABU ABG ABB ATE ATW BOBj BBF KAK ANK
836 hiarbas $ –– ABU ATE ATW BOB5 BBI BBF KAK ANK
SEVENIA
837 occidentalium –– ooo ATE BOB7 KAK ANK
838 boisduvali –– ooo ATE ooo kak ank
839 umbrina –– ooo –– –– –– ––
841 amulia BOB7
LIMENITIDINAE
HARMA
843 theobene $ –– ABU ATE ATW BOBj BBI BBF KAK KAS ANK
CYMOTHOE
846 fumana –– –– ATE BOBj BBF KAK KAA KAS ANK
851 egesta $ –– –– ATE ATW BOB5 BBF KAK ANK ANN
853 lurida –– –– ATE BOBd ooo ank
858 aubergeri £ –– –– –– –– KAK TBL ooo
859 herminia –– –– ATE –– ooo ANK
860 weymeri –– –– ATE –– ooo ank
863 caenis $ –– ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAS ANK

866 althea –– –– ooo ooo KAK ANK
868 jodutta $ –– –– ATE bob KAK KAA KAS ANK ANN
872 coccinata –– –– ATE ooo ooo ooo
873 mabillei $ –– abu ATE ATW BOB5 BBF KAK KAA KAS ank
878 ‘sangaris’ $ –– –– ATE ATW BOB7 ooo ank
PSEUDONEPTIS
879 bugandensis $ –– ABU+ ABG ATE BOBo BBI BBF KAK KAA KAS ANK ANN
PSEUDACRAEA
880 eurytus $ –– ABU ABG ABB ATE BOBo BBF KAK KAS ANK
884 boisduvalii $ SHH8 SHI ABU ABG ABB ate BOB5 BBF kak ANK
887 lucretia $ –– ABU ABG ABB ATE BOBo BBF KAK ANK
888 warburgi $ –– –– ATE ATW BOBo BBF KAK ooo
889 hostilia –– –– ATE ooo ooo ANK
900 semire $ –– abu ATE ATW BOBo KAK ANK
NEPTIS
901 nemetes $ –– ABU ABG ATE ATW BOB5 BBI KAK ANK
903 metella $ –– ABU ABG ATE ATW BOB5 BBI BBF KAK ANK
905 serena $ SHH8 SHA ABU ATE ATW BOBh BBI KAK ooo
906 kiriakoffi SHH SHI abu ate ooo kak ANK
907 morosa $ SHH SHA SHI ABU ate BOBh KAK KAS ank
908 loma –– –– ATE ooo ooo ooo
910 angusta –– –– –– BOB7 –– ––
911 alta $ –– ABU ATE BOBo BBI KAK ANK
912 seeldrayersi –– –– ATE BOBo kak ank
913 puella –– –– ATE BOB6 kak ANK
914 conspicua –– –– ate ooo kak ooo
915 najo –– abu ate BOB7 KAK ank
916 metanira –– –– ate ooo kak ANK
917 continuata –– –– ate ooo kak ooo
918 nysiades $ –– abu ATE BOB5 BBI BBF KAK ANK
921 nicomedes –– ooo ATE bob KAK ANK
922 quintilla $ –– ooo ATE BOB6 BBI KAK ANK
926 paula –– –– ATE BOBh KAK ank
927 strigata –– –– ATE BOB5 KAK ank
929 nicoteles –– ABU ATE BOB+ KAK ANK

930 nicobule $ –– –– ATE ATW BOB8 BBF* KAK ANK
931 mixophyes –– –– ATE BOB5 KAK ANK
933 nebrodes $ –– –– ATE BOB+ KAK ANK
934 trigonophora $ –– ooo ATE BOB+ BBI KAK ank
936 agouale $ –– ABU ABB ATE BOB5 BBI BBF KAK ANK
937 melicerta $ –– ABU ATE BOBd BBI BBF KAK KAS ank
938 troundi $ –– ABU+ ATE ATW BOB5 BBF KAK ANK
CATUNA
941 crithea $ –– ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAS ANK ANN
942 niji –– –– –– ooo –– ANK
943 oberthueri $ –– ooo ATE ATW BOBo BBI BBF KAK ANK ANN
944 angustatum $ –– ABU ABG ABB ATE ATW BOB+ BBI BBF KAK ooo
EURYPHURA
946 togoensis –– –– ATE BOB7 KAK ank
948 chalcis $ –– –– ATE BOB5 BBI BBF KAK ANK
HAMANUMIDA
951 daedalus $ SHH SHA SHI ABU ABG ATE BOBs BBI KAK KAA ANK
ATERICA
953 galene $ –– ABU ABG ABB ATE ATW BOBj BBI BBF KAK KAS ANK ANN
CYNANDRA
954 opis $ –– ooo ATE ATW bob KAK ANK
EURIPHENE
959 incerta –– –– ATE ooo KAK ANK
960 barombina $ –– ABU+ ABG ATE ATW BOBo BBI BBF KAK KAS ANK
961 veronica –– –– –– –– –– ANK
964 grosesmithi –– –– ooo –– KAK ank
968 simplex $ –– –– ATE ATW BOBo BBI BBF KAK ANK
974 amicia $ –– –– ATE ATW BOBj BBI KAK ank
976 aridatha $ –– ooo ATE ATW BOBo KAK ANK
978 coerulea –– –– ATE BOBj KAK ANK
985 ernestibaumanni –– –– ooo ooo ooo ooo
986 gambiae $ –– ooo ATE ATW BOBj BBF KAK KAA TBL ANK
987 ampedusa $ –– ABU ABG ABB ATE ATW BOBj BBF KAK ANK ANN
988 leonis –– –– –– –– ooo ooo
989 atossa –– –– ATE BOBk KAK ANK

990 doriclea –– –– ATE bob KAK ANK
BEBEARIA
994 lucayensis –– –– ATE BOB7 KAK ANK
994a languida BOB7 ???
995 tentyris $ –– ABU8 ABG ATE ATW BOBo BBI BBF KAK KAS ANK
996 osyris –– –– ATE ooo KAK KAS ANK
998 carshena –– –– ATE BOBo KAK ank
999 absolon $ –– ABU ABB ATE ATW BOBj BBI BBF KAK ANK
1001 zonara $ –– ooo ATE BOBo BBI BBF KAK ooo
1002 mandinga $ –– ABU ABG ATE BOBj BBI BBF KAK ANK
1003 oxione $ –– ABU ATE ATW BOBj KAK ANK ANN
1004 abesa $ –– –– ATE BOBj BBI KAK ank
1006 barce –– –– ATE bobJ8 KAK ANK
1008 mardania $ –– ABU ABG ATE BOBj BBI BBF KAK KAA KAS ANK ANN
1011 cocalia –– ABU ATE BOBj BBF KAK KAA ANK
1012 paludicola $ –– –– ATE BOBu BBI BBF KAK ANK
1014 sophus $ –– ABU ABB ATE BOBj BBI BBF KAK ANK
1017 arcadius –– –– ATE –– ooo ANK
1021 laetitia $ –– –– ATE ATW bob KAK ANK
1027 phantasina $ –– ooo ATE ATW BOBo BBI BBF KAK ANK
1029 demetra –– –– ate BOBj kak ank
1033 maledicta –– –– ATE ooo kak ank
1035 ashantina –– –– ATE BOBo KAK ank
1037 cutteri –– –– ATE ooo KAK ANK
EUPHAEDRA
Subgenus Medoniana
1046 medon $ SHH SHA SHI ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAA KAS ANK
Subgenus Gausapia
1047 gausape –– –– ATE BOBj KAK ANK
1047 mariaechristinae –– –– ATE bob KAK ANK
Subgenus Xypetana
1055 xypete $ –– ooo ATE ATW BOBj BBI BBF KAK ANK
1057 hebes $ –– –– ATE BOBj KAK KAS ANK
1059 diffusa £ –– abu ATE BOBo BBF ooo ––
1060 crossei –– –– –– –– –– ––

1061 crockeri $ –– –– ATE ATW ooo KAK ooo
Subgenus Radia
1062 eusemoides –– –– ATE –– –– ooo
Subgenus Euphaedra
1064 cyparissa –– –– ATE BOBj KAK ank
1065 sarcoptera –– ABUj ATE BOB5 KAK ooo
Subgenus Euphaedrana
1066 themis $ –– ABU ABG ABB ATE BOBj KAK ANK
1067 laboureana –– –– ATE bob KAK ANK
1071 minuta –– –– ooo BOB6 KAK ANK
1072 modesta $ –– –– ATE BOB6 KAK ANK ANN
1075 janetta $ –– ABU ABB ATE ATW BOBj BBF KAK ANK
1076 splendens –– –– ATE BOBk ooo ooo
1077 aberrans –– –– ooo –– ooo ooo
1078 vetusta –– –– ooo –– KAK ooo
1083 ceres $ SHH8 SHA SHI ABU ABG ABB ATE ATW BOBo BBI BBF KAK ANK
1085 phaethusa $ –– ABU ABG ATE ATW BOBo BBI BBF KAK KAA KAS ANK ANN
1086 inanum £ –– ooo ATE BOBj BBF KAK ooo
1096 ignota –– –– ATE ooo KAK ooo
1103 velutina ooo KAK ANK
1106 francina $ –– –– ATE ATW –– KAK KAA ANK ANN
1108 eleus $ –– –– ATE ATW BOB5 KAK KAA ANK
1112 zampa $ –– –– ATE ATW –– KAK DAB ANK
1115 edwardsii $ –– ABU ABG ATE BOBj BBF KAK ANK
1116 ruspina –– –– –– –– –– ––
1117 perseis –– –– ATE BOBj KAK ANK
1118 harpalyce $ SHH8 SHA ABU ABG ABB ATE ATW BOBj BBF KAK KAS ANK ANN
1119 eupalus $ –– –– ATE ATW BOBo KAK ANK
EUPTERA
1121 crowleyi –– –– ate ooo kak ank
1122 elabontas £ –– –– ate bob kak ank
1123 dorothea –– –– –– –– –– ANK
1124 zowa –– ABU+ ATE+ bob kak ank
PSEUDATHYMA
1133 falcata –– –– ATE BOBh KAK ank

1134 sibyllina –– –– ATE BOBh KAK ank
1138 martini –– –– ate ooo ooo ank
HELICONIINAE
ACRAEA
Subgenus Actinote
1139 perenna $ –– –– ATE BOB+ BBI KAK8 KAA KAS ooo
1144 circeis $ SHH8 SHA SHI ABU ABG ABB ATE ATW BOBy BBI BBF KAK KAS ANK
1147 translucida –– –– ATE –– –– ––
1148 peneleos $ –– ABU ABG ABB ATE ATW BOBo BBI kak ANK
1149 parrhasia –– ABU ATE bob KAK ANK
1150 orina $ –– –– ATE BOB+ BBI kak ANK
1152 pharsalus –– ABU ABG ATE BOBy BBF KAK ANK
1153 encedon $ SHH ABU ATE ATW BOBo BBF KAK ANK
1154 encedana $ SHH SHA ABU8 ABB ooo ooo kak ooo
1155 alciope $ SHH* ABU ABG ABB ATE ATW BOBo BBI BBF KAK KAA KAS ANK ANN
1156 aurivillii –– –– ATE bob KAK ANK
1157 jodutta $ SHH ABU ABG ABB ATE ATW BOBo BBF KAK KAS ANK
1158 lycoa $ SHH SHI ABU ABG ABB ATE ATW BOB+ BBI BBF KAK KAA KAS ANK
1159 serena $ SHH ABU ABG ABB ATE BOBy BBI BBF KAK KAA KAS ANK
1160 acerata $ SHH8 SHA SHI ABU ATE BOB8 BBF KAK ank
1161 pseudepaea £ –– –– ATE BOBo kak ank
1165 bonasia shh ABU ATE BOB5 KAK ANK
1167 orestia SHH abu ATE BOB5 kak ANK
1168 polis $ –– ABU ATE ATW BOBy KAK KAS ANK ANN
1169 vesperalis £ –– –– ATE BOB6 BBF ooo ooo
Subgenus Acraea
1172 kraka –– –– ATE –– –– ––
1173 rogersi $ –– ABU ATE BOBo KAK KAS ank
1174 abdera $ –– ABU ate BOB8 BBI ooo ank
1176 egina $ SHH SHA ABU ABB ATE ATW BOBy BBI BBF KAK ANK
1178 pseudegina ooo ABU ATE BOB+ KAK KAS ANK
1179 caecilia ooo ooo ooo BOB6 –– ––
1180 zetes $ SHH SHA SHI ABU ABB ATE BOB6 BBF KAK ANK
1181 endoscota –– –– ATE BOB7 KAK ANK

1182 leucographa –– –– ATE BOB6 KAK ank
1184 quirina $ –– ABU ATE ATW BOBd BBF KAK KAS ANK
1185 neobule SHH SHI ABU ATE bob KAK KAS ANK
1186 eugenia –– –– –– –– –– ––
1187 camaena –– ooo –– ooo ooo ––
1188 vestalis $ –– ABU ABB ATE ATW BOBy BBF KAK ANK
1189 macaria $ –– –– ATE ooo KAK ANK ANN
1190 umbra $ –– ABU ABB ATE ATW BOB+ BBF KAK ANK ANN
1191 alcinoe $ SHH8 SHI ABU+ ABG ABB ATE ATW BOB+ BBF KAK KAS ANK
1192 consanguinea –– –– ooo ooo kak ANK
1196 epaea $ SHH SHA ABU ABG ABB ATE ATW BOB+ BBI BBF KAK KAS ANK ANN
LACHNOPTERA
1199 anticlia $ –– –– ABG ATE ATW BOBo BBI KAK ANK ANN
PHALANTA
1200 phalantha $ SHH SHA SHI ABU ABG ATE ATW BOB5 BBI KAK ooo
1201 eurytis $ –– ooo ATE BOBd BBI BBF KAK KAA KAS ANK ANN
HESPERIIDAE
COLIADINAE
COELIADES
1203 chalybe $ –– ABU ATE ATW BOB5 BBF KAK KAS ANK
1204 bixana –– ABU ate bob kak ank
1206 libeon –– abu ATE ooo kak ank
1207 forestan $ –– ABU ABB ATE BOB5 BBI BBF KAK KAS ANK
1208 pisistratus £ SHH ABU ABB ATE BOB8 BBF KAK ANK
1209 hanno –– ABU ATE bob KAK ank
PYRRHIADES
1210 lucagus SHH ABU –– –– –– ––
PYRRHOCHALCIA
1211 iphis $ SHH SHI ABU ABG ABB ATE bob KAK KAS ANK
PYRGINAE
LOXOLEXIS
1212 holocausta –– –– ATE ooo ooo ANK
1213 dimidia –– –– ooo –– ooo ank

1214 hollandi –– –– ATE BOB6 kak ANK
KATREUS
1215 johnstonii –– –– ATE ooo ooo ANK
CELAENORRHINUS
1216 rutilans –– –– ATE ooo ooo ank
1217 sagamase –– –– ATE ooo KAK ank
1219 leona £ –– –– ATE BOBo BBF ooo ank
1223 ankasa $ –– –– ATE ATW BOBj BBF KAK ANK
1224 galenus $ –– ABU ABG ABB ATE ATW BOB5 BBI BBF KAK ANK
1225 cf galenus –– –– ATE ooo kak ank
1226 meditrina –– –– ATE bob kak ank
1227 ovalis –– –– ATE ooo KAK ank
1230 proxima $ –– ABU ABB ATE BOB6 BBI BBF KAK ank
1231 plagiatus –– –– ATE BOBs KAK ANK
TAGIADES
1232 flesus $ SHH SHA ABU ABB ATE ATW BOB5 BBI BBF KAK KAA KAS ANK ANN
EAGRIS
1233 denuba $ –– ABU ABG ATE ATW BOB5 BBF KAK ANK
1234 decastigma –– –– ATE BOBs ooo ank
1235 tigris –– –– ATE BOBs ooo ank
1236 subalbida $ –– ooo ATE ATW BOB5 BBI BBF KAK ANK
1237 hereus £ –– –– ATE bob BBF KAK ANK
1238 tetrastigma –– ooo ATE BOB5 KAK ank
CALLEAGRIS
1239 lacteus –– –– ate BOBs kak ANK
PROCAMPTA
1241 rara –– ABU ATE BOB+ KAK ank
ERETIS
1242 lugens ooo abu ATE –– –– ––
1243 plistonicus £ –– ABU ATE BOB+ BBF KAK ANK
1244 melania –– ABU ate BOB+ ooo ooo
SARANGESA
1245 laelius $ –– –– –– BOB8 BBI –– ––
1246 phidyle –– –– –– –– –– ––
1247 tertullianus –– ooo ate BOB6 KAK ANK

1248 majorella –– abu ate bob kak ank
1249 tricerata –– ABU ooo bob ooo ooo
1250 thecla $ –– ABU ABG ABB ATE BOB5 BBI KAK KAS ANK
1251 bouvieri $ –– ABU ABG ABB ATE BOB5 BBI BBF KAK ANK
1252 brigida £ –– ABU ABB ATE BOB7 KAK ANK
CAPRONA
1253 adelica shh ooo –– –– –– ––
1254 pillaana shh –– –– –– –– ––
NETROBALANE
1255 canopus shh ooo –– –– –– ––
ABANTIS
1256 bismarcki –– –– –– –– –– ––
1257 leucogaster –– –– ATE BOB+ kak ank
1258 nigeriana SHH ooo ooo ooo ooo ooo
1259 pseudonigeriana –– –– –– –– –– ––
1261 lucretia –– –– ATE ooo kak ank
1262 elegantula –– –– ATE ooo kak ank
1263 ja –– –– ATE –– ooo kak
1263 tanobia –– –– ate –– ooo ank
SPIALIA
1265 spio shh ooo ooo ooo ooo ooo
1267 diomus shh ooo ooo ooo ooo ooo
1268 dromus shh ooo ooo ooo ooo ooo
1269 ploetzi $ –– abu ATE BOB8 BBI KAK ank
GOMALIA
1270 elma $ SHH SHI ABU ABG ooo ooo ooo ooo
HESPERIINAE
ASTICTOPTERUS
1276 anomoeus –– ABU ATE ooo ooo ooo
1277 abjecta £ SHH* ooo ooo –– –– ––
PROSOPALPUS
1278 debilis –– abu ATE ooo kak ANK
1279 styla –– ooo ate BOB6 ooo ooo
1280 saga $ –– –– ATE+ ATW ooo kak ANK

KEDESTES
1282 protensa –– –– –– –– –– ––
GORGYRA
1284 aretina –– ABU ATE BOB7 KAK ANK
1285 heterochrus –– –– ate bob KAK ANK
1286 mocquerysii –– ABU ATE bob KAK ank
1287 aburae £ –– ABU ABB ATE ooo kak ank
1289 bina –– ABU ATE bob KAK ank
1290 sola –– –– ooo ooo kak ank
1291 afikpo –– –– ATE ooo kak ank
1292 diversata –– ABU ate bob kak ANK
1293 bule –– ooo ooo bob kak ooo
1294 minima shh abu ooo ooo kak ooo
1295 sara –– ABU+ ATE BOB7 KAK ANK
1296 subfacatus –– ABU ATE bob KAK ank
1297 pali £ –– –– ATE BOB8 ?BBF kak ANK
GYROGRA
1299 subnotata –– abu ATE bob KAK ANK
CERATRICHIA
1301 phocion $ –– ooo ATE ATW BOBo BBI KAK ANK ANN
1302 semilutea –– abu ATE ooo KAK ank
1303 clara $ –– –– ATE ATW BOB6 BBI BBF KAK ank ANN
1305 crowleyi –– –– –– –– –– ANK ANN
1306 nothus $ –– –– ATE+ ATW BOB6 KAK ANK
1307 argyrosticta $ –– ABU ATE BOB+ BBI BBF kak ANK
1308 maesseni $ –– –– ATE BOB7 BBI BBF KAK ANK
TENIORHINUS
1309 watsoni –– ooo ate bob kak ANK
1310 ignita –– ooo ooo BOB8 BBF ooo ooo
PARDALEODES
1311 incerta $ SHH SHA ABU ooo bob ooo ooo
1312 edipus $ –– ABU ABG ABB ATE ATW BOB5 BBI BBF KAK KAS ANK ANN
1313 sator $ –– ABU ATE ATW BOB5 BBF KAK KAS ANK
1314 tibullus $ –– ABU ATE ATW BOB5 KAK ANK ANN
1315 xanthopeplus –– –– ATE ooo ooo ank

XANTHODISCA
1317 rega $ –– ABU ABG ATE8 ATW bob KAK ank
1318 astrape $ –– ooo ATE BOB+ BBI BBF KAK ank
PAROSMODES
1320 morantii SHH –– –– –– –– ––
1321 lentiginosa –– ABUj ATE BOB6 ooo ooo
RHABDOMANTIS
1322 galatia –– ooo ATE BOB6 KAK ank
1323 sosia $ –– ABU ATE BOB6 KAK KAS ank
OSMODES
1324 laronia –– abu ATE BOB5 KAK ANK
1325 omar –– ooo ate BOB7 KAK ANK
1326 lux £ –– –– ATE BOB6 BBF KAK ANK
1328 thora $ –– ABU ATE ATW BOB5 BBF KAK ANK
1329 distincta –– –– ATE ooo kak ank
1330 adon –– –– ATE bob KAK ank
1332 adosus –– –– ATE bob KAK ank
1333 lindseyi –– ooo ATE BOB+ kak ank
1334 costatus –– –– ATE BOB7 KAK ank
1335 banghaasi –– –– ATE BOB7 KAK ANK
OSPHANTES
1336 ogowena –– –– ate ooo KAK ank
PARACLEROS
1337 placidus –– ABU ATE+ bob KAK ank
1338 biguttulus $ –– ABU ABG ATE BOB+ BBI KAK KAS ANK
1339 substrigata £ –– –– ate BOB6 BBF KAK ank
1340 maesseni –– –– ATE bob KAK ank
ACLEROS
1341 ploetzi $ ABU ABB ATE BOB5 BBI BBF KAK ANK
1342 mackenii $ –– ABU ATE BOBs BBI BBF KAK ANK
1343 nigrapex $ –– abu ATE ATW BOB6 BBF KAK ANK
1344 bala £ –– –– ate BOB7 BBF KAK ooo
SEMALEA
1345 pulvina –– ABU ATE bob KAK ANK
1346 sextilis £ –– ABU ATE bob BBF KAK ANK

1347 atrio –– –– ATE ooo KAK ank
1349 arela –– ABU ATE BOBs KAK ooo
HYPOLEUCIS
1350 ophiusa –– abu ATE BOB5 KAK ANK
1351 tripunctata –– –– ATE bob kak ank
1352 sophia –– –– ate ooo KAK ank
MEZA
1353 indusiata –– ooo ate bob KAK ANK
1354 meza $ –– ABU ABB ATE ATW BOB5 BBI BBF KAK KAS ANK ANN
1355 mabea –– –– ooo BOBs ooo ooo
1356 leucophaea $ –– ooo ATE bob KAK KAS ANK
1357 elba –– abu ATE bob KAK ank
1358 mabillei –– –– ATE BOB@ kak ANK
1359 cybeutes –– ooo ATE bob KAK ANK
PARONYMUS
1361 xanthias –– –– ATE bob KAK ANK
1363 ligora £ –– abu ATE BOBs BBF KAK ank
1364 nevea –– –– ATE@ –– ooo ANK
ANDRONYMUS
1365 neander –– ABU+ ATE bob kak ank
1367 caesar $ –– ABU+ ABB ATE BOB5 BBI BBF KAK ANK
1368 hero $ –– ooo ATE BOB5 BBF KAK KAS ank
1369 helles –– ooo ATE BOB6 KAK ANK
1370 evander –– ooo ATE BOB5 KAK ANK
ZOPHOPETES
1373 ganda –– ABU8 ABB ooo –– KAK8 KAA ooo
1374 cerymica –– ABU8 ABB ATE BOB7 KAK KAA ank
1376 quaternata –– ooo ooo ooo ooo ooo
GAMIA
1377 buchholzi £ –– ABU ABB ATE BOB7 BBF kak ANK
1378 shelleyi –– ooo ate BOBs KAK ANK
ARTITROPA
1379 comus $ –– ABU ABB ATE BOB7 BBI BBF kak ank
MOPALA
1380 orma £ –– –– ate BOB5 BBF KAK ank

GRETNA
1381 waga –– ABU ate BOB5 KAK ANK
1382 carmen –– –– ate BOB6 ooo ooo
1383 cylinda $ –– abu ate BOB5 BBI BBF KAK ANK
1386 balenge –– ABU ate BOB5 KAK ank
PTEROTEINON
1387 laufella $ –– ABU ABG ABB ATE BOB5 BBI BBF KAK KAA KAS ANK
1388 iricolor –– –– ATE BOB+ KAK ANK
1389 laterculus –– –– ate bob KAK ank
1390 capronnieri –– –– ooo –– ooo ANK
1391 caenira $ –– ABU ABG ATE BOB6 BBF KAK KAA KAS ANK
1392 ceucaenira –– abu ATE bob KAK ANK
1393 concaenira –– –– ATE ooo KAK ANK
1394 pruna –– –– ate BOBs KAK ank
LEONA
1395 binoevatus –– –– ate bob KAK ank
1397 lota –– –– ooo –– ooo ANK
1399 leonora –– –– ate bob kak ank
1401 stoehri –– –– ate ooo KAK ank
1402 meloui –– –– ate ooo kak ank
1403 halma –– ??? ??? ??? ??? ???
1404 lissa –– –– ATE –– ooo ooo
1405 luehderi –– ABU ate ooo ooo ank
CAENIDES
1406 soritia –– ooo ATE BOB6 kak ANK
1407 kangvensis £ –– ABU+ ATE BOBd BBF KAK ANK
1408 xychus £ –– –– ate BOB+ BBF KAK ank
1409 benga –– –– ATEs BOB+ kak ank
1410 otilia –– –– ate ooo kak ANK
1411 dacenilla –– –– ate ooo kak ANK
1412 dacela £ –– ABU ABB ATE BOB7 BBF KAK ANK
1413 hidarioides –– –– ATE BOB6 KAK ank
1414 dacena –– ABUi ATE BOB6 KAK ANK
MONZA
1415 alberti $ –– ABU ABB ATE ATW BOB+ BBF KAK ANK

1416 cretacea $ SHH+ ABU ATE ATW BOB5 BBI BBF KAK KAA ANK
MELPHINA
1417 noctula –– –– ATE+ ooo KAK ANK
1419 unistriga $ –– ABU ATE BOB8 BBF KAK ank
1420 tarace $ –– ABU8 ABG ATE BOB7 BBI BBF ooo ank
1421 flavina –– ABU ate ooo kak ank
1422 statirides –– ooo ATE BOB8 BBF kak ank
1423 statira $ –– ooo ooo BOB8 BBI ooo ank
1425 malthina –– –– ate bob KAK ANK
1426 maximiliani –– –– ooo –– ooo ooo
FRESNA
1427 netopha £ –– ABU ATE BOB6 kak ooo
1428 maesseni –– –– ate BOB6 kak ank
1429 nyassae –– abu ATE BOB6 kak ooo
1430 cojo £ –– abu ATE BOB8 BBF kak ank
1431 carlo £ –– –– ate BOB8 BBF KAK ank
PLATYLESCHES
1432 galesa $ ooo abu ATE ATW bob kak ANK
1434 moritili shh abu ate ooo ooo ooo
1435 rossi –– ooo ooo –– ooo ooo
1437 picanini –– abu ATE bob KAK ANK
1438 lamba –– –– ooo ooo ooo ooo
1439 affinissima ooo ooo ooo ooo ooo ooo
1440 chamaeleon ooo ooo ooo ooo ooo ooo
1441 batangae ooo –– ooo –– –– ––
PELOPIDAS
1444 mathias $ SHH SHA SHI ABU ATE BOB6 BBI kak ANK
1445 thrax $ SHH8 SHA SHI abu ATE bob kak ank
BORBO
1446 fallax SHH abu ooo ooo kak ooo
1447 fanta £ SHH8 SHI ABU ABB ate BOB5 BBF KAK ANK
1448 perobscura £ SHH ABU ATE BOB6 BBF KAK ank
1449 micans shh ooo ATE bob kak ank
1450 borbonica shh ooo ate ooo ooo ooo
1451 gemella SHH+ abu ooo ooo kak ooo

1452 binga –– ABU+ ooo bob KAK ank
1453 fatuellus $ SHH SHI ABU ABB ATE BOB5 BBF KAK KAA ANK
1454 holtzi shh ABU ooo ooo ooo ––
PARNARA
1456 monasi –– ooo ATE bob kak ank
GEGENES
1457 ‘pumilio’ ooo –– –– –– –– ––
1459 niso ooo abu ooo ooo KAK ooo
1460 hottentota ooo ooo ATE+ ooo ooo ooo
$ Tour Group 349 118 56 62 316 134 120 604 183 484 196 262 509 68 119 405 76
£ Subsequent 61 joint 138
TOTAL 2008 410