Bulletin of the European Association of Coleopterology vol. VIII ...

Bulletin of the European Association of Coleopterology
vol. VIII
Boletin de la Asociaci6n Europea de Coleopterologia
vol. VIII
Barcelona 1994

ELYTRON. 1994. Vol 8: 63-72 ISSN: 02 14- 1353
TWO NEW SPECIES OF
MICROACMAEODERA COBOS, 1966 FROM
THAILAND (COLEOPTERA: BUPRESTIDAE)
M. G. Volkovitsh
Zoological Institute
Russian Academy of Sciences
199034 ST. PETERSBURG. RUSSIA
C. L. Bellamy
Coleoptera Department
Transvaal Museum
PO Box 413
PRETORIA 0001. SOUTH AFRICA
ABSTRACT
Two New species of Microacmaeodera Cobos, 1966 from Thailand (Coleoptera: Ruprestidae).
Mi(./-oc~c~nrucocle~r-u (S~IIUIII~~~I-~U~~III~IC~~~I~~I-~I)
XII/XIIII and M. (Mie~r.ouc.nruco~/c~~c~) tl~uil~~~c/i(~u
spp. n. arc described I'roni Thailand. illuslralcd and contraslcd from thcir congcncrs in both
tabularly and in a kcy.
Key words: Colcoplcra. Uuprcslidac, Acmacodcrinac, Mi[./-otrt~rircrcoclrr-(~, Thailand.
INTRODUCTION
The buprestid genus Mi[*]-oac.n~aeode~-a was originally described by COBOS
( 1966) as a subgenus for Acmaeodel-u (Mic.1-ocic.n~ueode~-a) 1ongic.ol-11i.s Cobos
from eastern Afghanistan. Later the status of Mic.~-oac.n~ucode~-a was elevated to
the generic rank by VOLKOVITSH ( 1986), and two subgenera, Mic.1-oat-n~aeodel-a
(s. .st]-.) and M. (Squarnic.~-oac.n~aeode~-a) were separated.
In terms of extant distribution and zoogeography, the species of Mic.1-oacmacodel-a
arc from the Indo-Oriental region: M. (M.) 1ongico1-nis (type species)
and M. (M.) ~7ittn1c.1-i Vol kovitsh are partly geographically sympatric in the
western Himalayas; M. (S.) helli (Kerremans) (type species) is from India and
M. (S.) a]-ue~~sis ('l'hiry) is from Aru lsland and perhaps Borneo. Recenlly one
more species, M. (S.) rncicg~-ego]-i Bellamy & Volkovitsh ( 1992) was described
from Luzon Is., Philippines.
The discovery of two new species, collected by Czech coleopterists in
Thailand during 1991-92, has prompted a return to the taxonomy of this genus
once again. The locality data is recorded herein as presented on the specimen
labels. We are grateful to our colleague Ing. Vitezslav Kuban, Brno, Czech
Republic for rhe loan of material for this study; his collection will be
abbreviated VKBC.

M. G. VOLKOVITSH & C. L. BELLAMY
Microacmaeodera (Squamicroacmaeodera) kubani sp. n. (Figs. 1, 2, 6)
Diagnosis
Body small, length 3.4 mm, width 1.2 mm, short, broad, robust, weakly
convex, with marked dorsal curvature (Fig. 1); black with coal-like sheen;
covered with thin white hyaline scales.
Description
Head broad, flat above; front weakly depressed in lower part, with sides
nearly straight and diverging slightly to vertex. Vertex without longitudinal
keel, 1.90~ as wide as transverse diameter of eye and 1.06~ as wide as front
above antenna1 depressions. Clypeus narrow, with small, shallow, arcuate
median emargination anteriorly. Front with sculpture reticulate, formed by
rounded superficial umbilicate punctures with marked granules and centrally
placed dots; intervals forming fine transverse rugae in lower part; covered with
short, fine decumbent scales. Antennae short (female), 1.41~ as long as height
of eye; expanded from antennomere 5; antennomere 2 ovoid, robust; 3 and 4
similar, short, feebly expanded apically; 5 triangular; 6-10 strongly transverse,
almost 2x as wide as long; I I oblong, irregular. Pronotum evenly convex,
without basal cavities and median depression; slightly wider than long, basal
width 1.41~ length, widest in anterior to base; sides feebly arcuately converging
anteriorly; anterior margin projecting angularly, slightly bisinuate; basal margin
straight. Lateral carina narrow, distinct, entire, straight. Sides with reticulaterugose
sculpture, marked concentric rugae and fragments of umbilicate punctures
with distinct dots placed between them; disc with simple punctate sculpture of
dense large punctures in the middle; intervals smaller than diameter of punctures,
shining. Vestiture consisting of short, barely visible, decumbent, fine scales.
Anterior prothoracic margin slightly bisinuate, bordered by a deep groove;
prothoracic sculpture consisting of small, elongate, asperate umbilicate punctures;
sculpture of meso-, metathorax and metacoxae consisting of similar punctures.
Hypomera with reticulate-rugose sculpture of marked longitudinal rugae with
shagreened intervals and hardly visible vestiges of umbilicate punctures. Elytra
broad, convex; 2.13~ as long as wide at base; sides slightly converging
posterior to humeri, feebly diverging arcuately toward midpoint, and firstly
evenly arcuate, then strongly, almost straightly converging to widely jointly
rounded, nearly truncate apices. Incisure posterior to humeri shallow but
marked, extending into shallow curvature at the level of hind coxae; denticles
of epipleura small, hardly visible only in apical 114. Stria1 punctures fine,
fusing together along the entire length of elytron and forming grooves on
posterior 113; striae distinct along entire length. Intervals flat, nearly similar,
3-4x as wide as striae; 9th not elevated, without serration; covered with large,
superficial, uniseriate punctures on finely rugulose background, especially on
basal 113, and with evenly uniseriate, narrowly lanceolate, hyaline, short scales
approximately 112 interval width; surface shining, without pattern. Legs black;
metacoxae with posterior margin straight, slight emarginate laterally but without
lateral tooth. Tibiae slightly expanded apically, straight. Femora with thin
scales; tibiae with short inconspicuous setae; metatibiae bearing one row of
white bristle-like setae on external margin. Tarsomeres nearly similar; 5th thin,
expanded apically; tarsal pulvilli present, well developed on 1-4, each larger
toward distal end; basal tarsomeres with markedly longer setae laterally. Claws
short, broad, curved, and with broad, blunt tooth before middle of internal

TWO NEW SPECIES OF MICROACMAEODERA
Figs. I ant1 4: Dol.s;~l habitus of: Mic~rocrcn~creodcru (Sq~ron~ic~-oac~ni~~codcrc~) Xuhot~i. sp. n. ( I );
Mic.r.onc~niurur1er.a (s. sir..) thoilandica, sp. n. (4).
margin. Abdomen completely black; with reticulate sculpture, 1st and anal
sternites with reticulate-rugose sculpture with marked longitudinal rugae; disc
with partly obliterated reticulate sculpture of elongate umbilicate punctures;
covered with fine decumbent scales. Anal sternite of female short, broadly
rounded, feebly deflexed apically.
Male. Unknown.
Female
Ovipositor of tubular type (Figs. 2, 6), rather elongate, approximately 3 .5~
as long as enlarged part; rather deeply emarginate apically; with short parallelsided
styli, separated from each other by 2x their length, bearing apical bristles
1.5~ as long as length of stylus. Hemisternites thin, slightly curved; apical
sclerotization of dorsal ones poorly marked, undifferentiated, without recurved
branches (Fig. 6).

66 M. G. VOLKOVITSH & C. L. BELLAMY
Specimens examined
Holotype, female: Thailand, Thimonghta, 350m, 15.02N, 98.335E, 9- 13.-
IV.1991, Vit. Kuban Igt.; Thailand '91, Thanon Thong Chai, D. Kral et V.
Kuban. Holotype deposited in VKBC.
Comments
Microacmaeoder-a (S.) kuhani comes closest to M. (S.) aruensis, differing
by the small body, lateral keel of pronotum entire, and sculpture of hypomera
and abdomen markedly reticulate-rugose. From this last character it differs
from all other species of M. (Squamicroacmaeodera). The four species of this
taxon are contrasted in Table 1.
Etymology
This new species is obviously named for the collector of the unique type
specimen, our friend and colleague Vita Kuban.
Microacmaeodera (Microacmaeodera) thailandica sp. n. (Figs. 3, 4, 5)
Diagnosis
Body small, length 3.2 mm, width 1.0 mm, elongate, relatively broad,
convex, with poorly marked dorsal curvature (Fig. 4); black with coal sheen;
covered with white hyaline decumbent setae.
Figs. 2 and 5: Ovipositor, dorsal aspect: Microacniaeodera (Squamicroacn~aeoder'a) kuhani, sp. n.
(2); Microacmaeodera (s. str.) thailandii~a, sp. n. (5).

TWO NEW SPECIES OF MICROACMAEODERA
Description
Head broad, convex from above, feebly flattened in middle; front weakly
depressed in middle, convex in dorsal portion, sides nearly straight and
strongly converging to vertex; vertex without longitudinal keel, 2. 10x as wide
as transverse diameter of eye and 0.95~ as wide as front above antenna1
depressions. Clypeus very narrow. nearly straight, with weak, broad, arcuate
median emargination anteriorly. Front with sculpture punctate, formed by fine,
simple, sparse (particularly on vertex) punctures; intervals smooth, shining,
wider than diameter of punctures; covered with short. fine, decumbent setae.
Antennae relatively short (female). 2 .0~ as long as height of eye: expanded
from antennomere 5: antennomere 2 ovoid, robust; 3 and 4 similar, short,
feebly expanded apically; 5- 10 triangular, nearly as wide as long, with arcuate
inner margin; 11 elongate with rounded apex. Pronotum evenly convex,
without basal cavities and median depression; slightly wider than long. basal
width 1.25~ length, widest between middle and posterior 113; sides arcuately,
more strongly converging anteriorly than posteriorly; anterior margin feebly
projecting anteriorly, nearly straight; basal margin straight. Lateral carina
narrow. distinct, entire, nearly straight. reaching anterior margin. Sides with
pseudoalveolate culpture; disc with simple punctate sculpture of sparse punctures;
intervals larger than diameter of punctures. shining. Vestiture consisting of
short, decumbent. similar, hyaline setae. Anterior prothoracic margin slightly
bisinuate. bordered by a fine groove; prosternum evenly convex with small,
sparse, deep, simple punctures; sculpture of meso-, metathorax and metacoxae
consisting of similar, but larger punctures. Hypomera with large, deep, simple
punctures on finely shagreened background. Elytra relatively broad, convex;
2.30~ as long as wide at base; sides slightly converging posterior to humeri,
then very feebly diverging to midway between middle and posterior 113. and
weakly arcuate, converging to widely jointly rounded apices. Incisure posterior
to humeri very shallow but marked; denticles of epipleura small, serrate.
visible only in apical 114. Stria1 punctures deep, rounded or slightly elongate,
isolated on anterior 112 and parlly fusing posteriorly; forming shallow grooves
on posterior 112; striae distinct along entire length. Intervals subconvex, nearly
similar, narrow, 1.5-2.5x, at most 3x as wide as striae; 9th not elevated,
without serration; covered with fine, marked, mostly uniseriate punctures on
finely rugulose. shining background, and with short, uniseriate, hyaline,
posteriorly-pointing setae, hardly shorter than interval width: surface shining.
without pattern. Legs blackish-brown; metacoxae with posterior margin slightly
emarginate without lateral tooth. Tibiae thin, very feebly expanded apically,
weakly curved. Legs covered with light setae and bristles; metatibiae bearing a
row of yellowish bristles on external margin. First tarsomere slightly longer
than 2 and 3 together; 5th thin, expanded apically, as long as three proximal
tarsomeres together; tarsal pulvilli small, poorly developed, each larger toward
distal end. Claws short, curved, with large, acute, blunt tooth before middle of
internal margin. Abdomen completely black; with simple punctate sculpture,
sides with more dense and coarse punctures, disc with sparse, fine, slight
asperate punctures; covered with fine decumbent scales. Anal sternite of
female short, broadly rounded, bordered with fine groove.

68 M. G. VOLKOVITSH & C. L. BELLAMY
Male. Unknown.
Female
Ovipositor of tubular type (Figs. 3, 5), relatively short, approximately 2.5~
as long as enlarged part; deeply triangularly emarginate apically; styli separated
from each other by 2x their length, bearing apical bristles 1.5~ as long as
length of stylus. Hemisternites thin, slightly curved; apical sclerotization of
dorsal ones well marked, differentiated, with long, bent recurved branches
(Fig. 3).
Specimens examined
Holotype, female: NW Thailand, Mae Hong Son, 7-14.V. 1992, P. Pacholatko
lgt. Holotype deposited in VKBC.
Figs. 3 and 6: Apical portion of ovipositor, dorsal aspect: Microacmarodrra (s. str.) thailandica,
sp. n.(3); Micrc~acmaeodrra (Squamicroac~maeodera) kuhani, sp. n.(6).

TWO NEW SPECIES OF MICROACMAEODERA
Comments
Microacmaeodera (s. str.) thailandica differs from all others of its
congeners from M. (s. str.) by having the feebly elongate, rather broad body
and relatively short antennae. With these characters it bears a superficial
resemblance to species of M. (Squamicroacmaeodera); however, it can be
separated into the nominate subgenus by virtue of the punctate sculpture and
setose vestiture. Moreover, the female antennae of M. (s. str.) thailandica are
significantly longer than those of M. (Squamicroacmaeodera) species, which
are characterized by an antenna1 length to eye height index varying from 1.32
(M. kuhani) to 1.57 (M. macgregori); the female of M. aruensis was not
studied. The three species of Microacmaeodera (s. str.) are contrasted in
Table 2.
Etymology
This species is named, obviously, for the country of origin.
KEY TO THE SUBGENERA AND SPECIES OF MZCROACMAEODERA
I. Body broad and robust, elytra not longer than 2.3~ basal width (generally less than 2.2);
covered with thin transparent squamae or squamose setae. Antennae normal, no more than
twice as long as height of eye in both sexes. Sides of pronotum with alveolate, reticulate or
reticulate-rugose sculpture of umbilicate punctures; disc with asperate or simple punctate
sculpture. Elytral intervals broad, no less than 2.5~ as wide as striaes ...............................
................................ Subgenus Squamicroacmaeodera Volkovitsh ................................ 2
- Body elongate and narrow, elytra no less than 2.3~ basal width (generally 2.44-2.90);
covered with simple setae. Antennae longer, in female no less than twice as long as height
of eye, in male up to half length of body. Pronotum completely covered with simple
punctation, occasionally more dense'and pseudoalveolate on the sides. Elytral intervals
equal to or not more than twice as wide as striae ..............................................................
....................................... Subgenus Microacmaeodero Cobos ....................................... 5
2. Punctate striae distinct on entire length of elytra. Hind coxa without tooth .................... 3
- Punctate striae visible only on posterior ID of elytra, anteriorly fused with punctures of
intervals. Hind coxa with large rectangular lateral tooth. Front parallel-sided, not widening
to vertex. Length 4.2 mm. India ................................................ M. (S.) helli (Kerremans)
3. Front distinctly widening to vertex. Width of pronotum at base 1.41-1.46~ length; with
greatest width at middle or posterior to middle; sides of pronotum with reticulate or
reticulate-rugose sculpture, disc with simply punctate sculpture. Elytral epipleura with
distinct incisure behind humeri ....................................................................................... 4
- Front parallel-sided, not widening to vertex. Width of pronotum at base 1.59-1.64~ length;
with greatest width just anterior to base; pronotal sculpture with sides alveolate, disc
asperate. Elytral epipleura with only feeble incisure posterior to humeri. Median lobe of d
genitalia without lamina. 5.4-5.9 mm. Philippines (Luzon Isl.) ..........................................
....................................................................... M. (S.) rnacgregori Bellamy & Volkovitsh
4. Lateral keel of pronotum well developed only at posterolateral angles. Hypomera with
round umbilicate punctures. Median lobe of d genitalia with lamina divided, consisting of
2 sclerites. 5.5-5.7 mm. AN Isl., Borneo? ................................... M. (S.) oruensis (Thtry)
- Lateral keel of pronotum distinct, entire. Hypomera with reticulate-rugose sculpture of
longitudinal rugae. d unknown. 3.4 mm. Thailand ........................... M. (S.) kuhani sp. n.

M. (S.) belli M. (S.) aruensis
Front parallel-sided,
with reticulate sculp-
ture of umbilicate
punctures with irregu-
lar smaller punctures
on surface.
Pronotum widest at
middle; width at base
1.33~ length; lateral
keel distinct, entire; sides
with reticulaterugose
sculpture, disc
with simply punctures.
Hypomera with large,
elongate, umbilicate
punctures with rugose
surface.
Elytral epipleura with
shallow but distinct in-
cisure posterior to hu-
meri.
Elytral striae visible
only on posterior 112.
Hind coxae with large
rectangular tooth.
Abdomen with simple
or asperate punctures,
except on sternite I
laterally.
Male genitalia unk-
nown.
Body length 4.2 mm.
width 1.4 mm.
M. G. VOLKOVITSH & C. L. BELLAMY
Front with sides di-
verging to vertex, with
ocellate sculpture of
umbilicate punctures
bearing punctures in
middle.
Pronotum widest posterior
to middle; width
at base 1.44 (1.43-
1.46)~ length: lateral
keel not distinct, well
developed only at posterolateral
angles; sides
with reticulate or
reticulate-rugose sculpture;
disc with simple
punctate sculpture.
Hypomera with round
umbilicate punctures
with granulate surface.
Elytral epipleura with
shallow but distinct in-
cisure posterior to hu-
meri.
Elytral striae distinct
along entire length.
Hind coxae without la-
teral tooth.
Abdomen entirely co-
vered with fine reticu-
late sculpture of umhi-
licate punctures.
Medium lobe of d ge-
nitalia with lamina di-
vided into two scleri-
tes.
Body length 5.6 (5.5-
5.7) mm; width 2.0
(2.0-2.10) mm.
M. (S.) kubani M. (S.) macgregor-i
Front with sides di-
verging to vertex, with
reticulate sculpture of
umbilicate punctures
bearing punctures in
their middles.
Pronotum widest at
middle, width at base
1.41 x length; lateral
keel distinct, entire; si-
des with reticulate-
rugose sculpture; disc
with simple punctures.
Hypomera with reticu-
late-rugose sculpture
of longitudinal rugae.
Elytral epipleura with
shallow but distinct in-
cisure posterior to hu-
meri.
Elytral striae distinct
along entire length.
Hind coxae without la-
teral tooth.
Abdomen covered with
reticulate and reticula-
te-rugose sculpture of
umbilicate punctures,
partly obliterated in
middle.
Male genitalia unk-
nown.
Body length 3.4 mm,
width 1.4 mm.
Front nearly parallel-
sided, with ocellate to
reticulate sculpture of
umbilicate punctures
with irregular smaller
punctures on surface.
Pronotum widest at
posterior 113 or just
anterior to base; width
at base 1.62 (1.59-
1.64)~ length; lateral
keel distinct, entire; sides
with alveolate
sculpture; disc with asperate
sculpture.
Hypomera with round
umbilicate punctures
with granulate.
Elytral epipleura with
very feeble incisure
posterior to humeri.
Elytral striae distinct
along entire length.
Hind coxae without la-
teral tooth.
Abdomen with aspe-
rate punctures except
laterally on sternite 1.
Medium lobe of d ge-
nitalia without lamina.
Body length 5.5 (5.4-
5.9) mm; width 2.0
(1.9-2.1) mm.
Table I. Diagnostic differences between four species of Microacntaeoderu (Squanticroacn~aeoderu~.

TWO NEW SPECIES OF MICROACMAEODERA
M. (M.) lon~icomis M. (M.) wittrneri M. (M.) thailandica
Black with bluish sheen.
Strongly or moderately elon-
gate, length of elytron 2.72
(2.44-2.90) x width at base.
Vertex 1.05 (1.00-1017) x as
wide as front above antennal
depressions; front parallel-sided
or slightly wider to vertex.
Length of antenna in d 5.26
(5.18-5.3 1) x, in 9 3.07 (2.88-
3.33) x height of eye.
Width of pronotum at base 1.25
(1.20- 1.30) x length; sides wit-
hout tubercles laterally; sculp-
ture completely even.
Median lobe of d genitalia with
entire lamina displaced to base
of apophyses.
Ovipositor greatly elongate. to-
tal length 5 x widest part.
Black with aeneous sheen.
Strongly or moderately elon-
gate, length of elytron 2.61
(2.48-2.74) x width at base.
Vertex 0.97 (0.96-0.97) x as
wide as front above antennal
depressions; front slightly na-
rrowed to vertex.
Length of antenna in d 3.54 x,
in Q 2.25 x height of eye.
Width of pronotum at 1.09
(1.07-1.10) x length; sides with
weak lateral tubercles at midd-
le; sculpture completely even.
Median lobe of Q genitalia with
lamina displaced to anterior por-
tion, divided, consisting of two
sclerites.
Ovipositor relatively short, total
length 3 x widest part.
Black with coal sheen.
Feebly elongate, length of ely-
tron 2.30 x width at base.
Vertex 0.95 x as wide as front
above antennal depressions;
front slight narrowed to vertex.
Length of antenna in 9 2.00 x
eye height.
Width of pronotum at base 1.25
x length; sides without lateral
tubercles; punctation very den-
se laterally, sparse on disc.
Male genitalia unknown.
Ovipositor relatively short, total
length 2.5 x widest part (Figs.
3, 5).
Table 2. Diagnostic differences between three species of Microacmaeodcra (s. str.)

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