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FRDA Report: Black Cottonwood and Balsam Poplar Managers ...

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The main cottonwood-associated species that usually concern forest managers<br />

in␣ the Prince Rupert Forest Region because of their competition with conifers are<br />

as␣ follows:<br />

Major competing species<br />

accompanying cottonwood Seral origin<br />

Vegetation complex or balsam poplar of competing species1 <strong>Cottonwood</strong> on floodplains Lonicera involucrata BB/SB<br />

in ICH, SBS, <strong>and</strong> BWBS Zones Cornus sericea BB<br />

Sambucus racemosa BB/SB<br />

Rubus parviflorus BB/SB<br />

Alnus crispa ssp. sinuata BB/SR<br />

Alnus tenuifolia BB/SR<br />

Calamagrostis canadensis BB/SR<br />

Cinna latifolia BB/SB?/SR?<br />

Symphoricarpos albus BB/SB<br />

<strong>Cottonwood</strong>-alder on Alnus rubra SR/SB(BB)<br />

floodplains in CWH Zone Rubus spectabilis BB/SB<br />

Cornus sericea BB/SB<br />

Oplopanax horridus BB<br />

Sambucus racemosa BB/SB<br />

Rubus parviflorus BB/SB<br />

Lonicera involucrata BB/SB<br />

Ribes bracteosum SB(BB)<br />

Mixed broadleaf species on Populus tremuloides SR/BB<br />

various sites in ICH, SBS, Betula papyrifera BB/SR<br />

<strong>and</strong> BWBS Zones Salix spp. BB/SR<br />

Alnus tenuifolia BB/SR<br />

Alnus crispa ssp. sinuata BB/SR<br />

1 SR = seed rain; BB = bud bank; SB = seed bank<br />

In the vegetation complex examples shown above, each complex consists of a list<br />

of species that tend to be associated with one another on ecologically similar sites. The<br />

lists have a silvicultural bias <strong>and</strong> thus emphasize species that may affect crop tree growth<br />

positively or negatively. The complexes are also useful for assessing potential wildlife<br />

forage values. Not all the species in each complex should always be expected to occur<br />

together on a given site. Depending on local conditions, species dominance varies␣ considerably.<br />

Although foresters can gain some insights from pre-harvest <strong>and</strong> adjacent st<strong>and</strong><br />

conditions to predict species composition of post-harvest competing species, they are<br />

still hampered by elements of chance when trying to predict seral vegetation trends.<br />

That element of chance can be reduced somewhat if there is a knowledge of available<br />

propagules, in the form of bud banks, seed banks in the soil, or seed rain from nearby<br />

ecosystems. For a given type of disturbance in a given area, the experience <strong>and</strong><br />

observation of local forest managers are the best predictors of successional stages,<br />

especially if that experience <strong>and</strong> observation are recorded in a way that can be used<br />

<strong>and</strong> integrated by others.<br />

Calamagrostis canadensis <strong>and</strong> Epilobium angustifolium are pioneer species that<br />

rapidly invade disturbed sites, especially in the boreal mixedwood region. Silviculturally<br />

they are important because they limit establishment of coniferous seedlings. However,<br />

on sites where balsam poplar is present, heavy competition from poplar root suckers<br />

after logging can decrease the normally very aggressive post-disturbance development<br />

of Calamagrostis.<br />

39

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