An annotated checklist of brachyuran crabs from Taiwan (Crustacea ...

Ng el al. : Checklist of crabs from Taiwan 31
and Jeng, 1999b
Permanotus purpureus (Gordon, 1934) ( = Harrovia purpurea) — Chia and Ng, 1998; Ng and
Jeng, 1999b
Tiaramedon spinosum (Miers, 1879) ( = Ceratocarcinus spinosuum) — Hwang and Yu, 1980; Ng
and Jeng, 1999b
Subfamily Galeninae A. Milne Edwards, 1862
Galene bispinosa (Herbst, 1783) (= Galene granulata Miers, 1884)—Sakai, 1939; Lin, 1947,
1949; Sakai, 1976; Dai, Yang, Song and Chen, 1986; Dai and Yang, 1991; Huang, 1994;
Jeng, Jan, Tzeng and Feng, 1997; Ng, 1998 [Lin (1947) reported Galene granulata as a recent
fossil record from Taiwan, but this taxon was originally described as an extant species. Galene
granulosa Miers, 1884, however, is clearly a junior synonym of G. bispinosa (Herbst, 1783).
The first author has examined a large series of specimens of this species from various parts of the
Indo-West Pacific and G. granulosa is nothing more than a juvenile of G. bispinosa. Small specimens
tend to have more granular carapaces, becoming distinctly smoother with increasing size. As
such, both names must be synonymised. ]
Subfamily Halimedinae Alcock, 1898
Halimede fragifer (De Haan, 1835)—Lin, 1949; Huang, 1994 [ Halimede tyche (Herbst, 1801) is
often regarded as a junior synonym of M. fragifer, but Galil (2000) recently argued that they were
two distinct species. ]
Subfamily Rhizopinae Stimpson, 1858
Pseudolitochira Integra (Miers, 1884) (= Heteropilumnus integer)—Jeng, 1997
Typhlocarcinus villosus Stimpson, 1858 — Lin, 1949; Huang, 1994
Subfamily Tanaochelinae Ng and Clark, 2000
Tanaocheles bidentata (Nobili, 1906) (= Chlorodiella bidentata)— Suzuki, 1985; Ng and Clark,
2000
Family Trapeziidae Miers, 1886
Subfamily Trapeziinae Miers, 1886
Quadrella maculosa Alcock, 1898 — Shin and Mok, 1996
Tetralia glaberrima (Herbst, 1790)—Miyake, 1938; Horikawa, 1940; Jeng, Jan, Tzeng, Feng and
Yang, 1994; 1996; Yu, Jeng, Chan, Ho and Shy, 1996; Jeng, 1997; Jeng, Jan, Tzeng and
Feng, 1997; Jeng, Shao, Tzeng, Feng and Wu, 1998
Trapezia areolata Dana, 1852 — Horikawa, 1940; Lin, 1949; Suzuki, 1985; Tzeng and Chen,
1992; Jeng, Jan, Tzeng, Feng and Yang, 1994, 1996; Jeng, Jan, Tzeng and Feng, 1997; Jeng,
Shao, Tzeng, Feng and Wu, 1998
Trapezia cheni Galil, 1983 — Galil, 1983; Yu, Jeng, Chan, Ho and Shy, 1996; Jeng, 1997
Trapezia cymodoce (Herbst, 1799) — Horikawa, 1940; Lin, 1949; Hwang and Yu, 1980; Suzuki,
1985; Jeng, Jan, Tzeng, Feng and Yang, 1994, 1996; Yu, Jeng, Chan, Ho and Shy, 1996;