An annotated checklist of brachyuran crabs from Taiwan (Crustacea ...

Ng el al. : Checklist of crabs from Taiwan
and Tung, 2000 [Fig. 8f] [The records of H. tridens (not Ocypode ( Helice) tridens De Haan,
1835) by Terao (1916), Sato (1936c, d), Horikawa (1940), Wu, Ling, Shieh and Wang
(1962), Chiu (1964), Sakai (1940, 1976), Su and Lue (1984), Shih, Lue and Wang (1991)
and Wu (1992) cannot be accurately ascertained, many probably being mixed species records including
H. formosensis and/or Helice sp. (see later). Similarly, records of H. latimera (not
Helice tridens latimera Parisi, 1918) from Taiwan by Maki and Tsuchiya (1923), Sakai (1939,
1940, 1976), Wu, Ling, Shieh and Wang (1962) and Wang and Liu (1996c, from Kinmen) are
probably also H. formosensis (M. Tiirkay, personal communication to P. K. L. Ng). See also
comments for Helice sp. later on ]
Helice subquadrata (Dana, 1851) (= Helice leachii Hess, 1865) — Sakai, 1939; Sakai, 1976;
Dai, Yang, Song and Chen, 1986; Fukui, Wada and Wang, 1989; Dai and Yang, 1991; Shih,
Lue and Wang, 1991; Huang, 1994; Hsueh, 1996; Yu, Jeng, Chan, Ho and Shy, 1996; Jeng,
1997; Lee and Tung, 2000 [Fig. 8g. ] [The old records of H. leachii from many parts of the
Indo-West Pacific are actually H. subquadrata according to M. Tiirkay (personal communication
to P. K. L. Ng), and this species will also be referred to a new genus when the revision of this
group is completed, see comments for Helice sp. later on]
Helice sp. (= Helice tridens aut. part; Helice wuana nut. part) — Sakai, 1939, 1940, 1976; Sakai
and Yatsuzuka, 1980; Dai, Yang, Song and Chen, 1986; Fukui, Wada and Wang, 1989; Dai
and Yang, 1991; Shih, Lue and Wang, 1991; Huang, 1994; Hsueh, 1996; Wang and Liu,
1996a, b; Ho and Hung, 1997; Jeng, Lin, Wang and Jan, 1998; Lin, 19988; Wang and Liu,
1998; Jeng and Wang, 2000; Lee and Tung, 2000 [Fig. 8h] [The specimens reported by various
authors as Helice wuana (not Helice tridens wuana Rathbun, 1931) from Taiwan will be referred
to a new species in a soon to be finished revision of the genera Chasmagnathus and Helice by K.
Sakai, M. Tiirkay and S.-L. Yang (M. Tiirkay, personal communication to P. K. L. Ng). He
also comments that the records of H. sheni Sakai, 1939, by Sakai (1939, 1976) (including
records from Taiwan) are also H. wuana s. str., the two names being subjective synonyms. Dai
and Yang's (1991) and Hsueh's (1995, 1996) records of H. japonica (not Helice {Helicana)
japonica K. Sakai and Yatsuzuka, 1980) are also likely to belong to the present new species. Helice
wuana (and the present new species) was referred to the subgenus Helicana Sakai and Yatsuzuka,
1980, by these authors and in the upcoming revision, Helicana will be regarded as a distinct
genus. ]
Hemigrapsus penicillatus (De Haan, 1835 ) — Sakai, 1939, 1940; Wu, Ling, Shieh and Wang, . ^ , ..„„
1962; Sakai, 1976; Dai, Yang, Song and Chen, 1986; Fukui, Wada and Wang, 1989; Dai and ^ '
Yang, 1991; Shih, Lue and Wang. 1991; Hsueh, 1996; Wang and Liu, 1996a, b; Ho and
Hung, 1997; Jeng, Lin, Wang and Jan, 1998; Jeng, Shao, Tzeng, Feng and Wu, 1998; Wang
and Liu, 1998; Chou, Lai and Fang, 1999; Jeng and Wang, 2000; Lee and Tung, 2000
Hemigrapsus sanguineus (DeHaan, 1835) — Balss, 1922c; Sakai, 1939; Sakai, 1976; Dai, Yang,
Song and Chen, 1986; Fukui, Wada and Wang, 1989; Dai and Yang, 1991; Hsueh, 1996; + ****'
Jeng, Jan, Tzeng, Feng and Yang, 1996; Wang and Liu, 1996a, b; Ho and Hung, 1997; Jeng,
Jan, Tzeng and Feng, 1997; Jeng, Lin, Wang and Jan, 1998; Jeng, Shao, Tzeng, Feng and
Wu, 1998; Wang and Liu, 1998; Jeng and Wang, 2000
Metaphtx elegans De Man, 1888 — Fukui, Wada and Wang, 1989; Shih, Lue and Wang, 1991;
45