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608 University of California Publications in Botany [Vol. 8
43. Cymathaere J. Ag.
Holdfast discoid or with inconspicuous, rudimentary, simple hap-
teres; stipe short, flattened, persistent; blade long, narrow, wider at the
base and tapering upward, longitudinally grooved, or loosely folded,
folds deep at the base becoming nearly plane above ; mucilage ducts
abundant and large in both stipe and blade ; sori broad, on both sur-
faces, at the base of the blade, paraphyses unicellular with well
developed hyaline appendage (not as in Griggs, Ohio Naturalist, vol. 7,
1907, p. 92, as to absence of either mucilage ducts or hyaline tips to
the paraphyses)
.
J. G. Agardh, De Lamin., 1867, p. 29.
The genus Cymathaere resembles Pleurophycvs in having longi-
tudinal grooves or folds. It is also peculiar in having a discoid hold-
fast, although this is not a primitive structure as Griggs (1907) sup-
poses. Griggs is, also, in error in denying mucilage ducts and hyaline
tips to the paraphyses on C. triplicata, the only known species.
Cymathaere triplicata (Post, and Rupr.) J. Ag.
Holdfast discoid, 6-15 mm. broad ; stipe short and stout, 5-25 cm.
long, cylindrical below, flattened above; blade undivided, narrowly
linear, tapering upward, with acuminate apex and rounded or cuneate,
rarely cordate base, 1.5-4 m. long, 8-18 cm. wide, thick and coriaceous
at the base, thinner above, possessing three characteristic longitudinal
folds extending throughout the length of the blade ;
sori broad, occupy-
ing the basal portion only ; paraphyses linear with hyaline appendages ;
color yellowish brown.
Growing on rocks in the upper sublittoral belt. From Bering Sea
to Puget Sound.
J. G. Agardh, De Lamin., 1867, p. 30; Saunders, Alg. Harriman
Exp., 1901, p. 430; Setchell and Gardner, Alg. N.W. Amer., 1903.
p. 264 ; Griggs, Cymathere, 1907, pp. 89-96, pi. 7 ; Collins, Holden and
Setchell, Phyc. Bor.-Amer. (Exsicc.), no. XXXIII a, b; Tilden, Amer.
Alg. (Exsicc), no. 343; Larniina/ria triplicata Postels and Ruprecht.
Illus. Alg., 1840, p. 10, pi. 10.
The type and only species of Cymathaere resembles in appearance,
color, texture, etc., Costa ria costata more nearly than any other alga.
In pressed specimens, the longitudinal folds are often obscured or even