THE GENUS BRYUM (BRYACEAE, MUSCI) IN MIDDLE ... - Arctoa

200
V. I. ZOLOTOV
1 mm
Fig. 86. Bryum pallescens Schleich. ex Schwaegr. (from Perm Prov., Bezgodov & Selivanov, 228, MW): leaves.
Scale bar 1 mm for all.
Bryotheca Fennica 148 (.1908), MW; 3-4) [Central Europe],
anonymous, MW; 5) Caucasus, Teberda Reserve,
2050 m elev., Ignatova 2.VIII.1986, MHA; 6) same, 1300
m, 1.VIII.1986, MHA.
Bryum lonchocaulon C. Muell.,
?Bryum cirrhatum Hoppe et Hornsch., Flora
2(6): 90. 1819. Fig. 87-88
Polyoicous: shoots partly synoicous, partly male
and rarely female in the same tuft; in synoicous
plants anteridia few, (1-)2-7, red; paraphyses yellow
to orangish, longer than antheridia.
Plants in wide, compact or dense tufts, 1.0-2.5
cm high, green, yellowish or brownish-green, slightly
glossy, tomentose, rhizoids ferrugineous, +papillose.
Stem puprle, 1.0-1.5 cm long, densely foliate,
comose, subterminal shoots 1-3, to 0.5-1.5 cm long,
more loosely foliate, comose. Lower leaves loosely
arranged, ovate-lanceolate to ovate, to 2.0 mm
long, 0.6 mm wide. Upper leaves spreading when
wet, densely imbricate and slightly spirally twisted
around stem when dry, 2.0-3.0(-3.8) mm long,
0.7-0.9(-1.2) mm wide, widest at 1/5-1/3 of
leaf length, lanceolate or broadly lanceolate, longly
and gradually acuminate, non-decurrent, red at
base; margin recurved or revolute from base to
acumen, entire or serrulate; border distinct, (4-
)6(-7)-rowed, composed of thick-walled linear cells,
not differentiated in color; costa stout, red at
base, yellow above, brownish in acumen, excurrent
in long, (0.24-)0.4-0.65(-0.8) mm, straight or slightly
flexuose, serrulate or rarely smooth, rather thick
awn. Laminal cells with sligthly incrassate cell
walls, slightly porose, in acumen narrow, to 70(-
80) x 8-10 μm, in upper leaf rhombic-hexagonal
to rombic, (50-)60(-80) x (12-)16(-18) μm, in midleaf
(40-)50-60(-70) x(12-)16-20(-22) μm, at base
rectangular, to 70 x 36(-40) μm, in leaf corners
subquadrate, to 50 μm wide.
Perichaetial leaves ovate-lanceolate, margin narrowly
recurved to plane in innermost leaves; costa
excurrent to percurrent and ending below leaf apex
in innermost leaves. Seta (2.5-)3.5-4(-6) cm, cupperreddish
to fuscous-red, thin, slightly twisted when dry,
arcuate below capsule. Capsule pendulous, light-fuscous
to brown, 2.0-3.0(-4.5) mm long, 1.0-1.5 mm
thick; cylindric, symmetric and contracted below
mouth to obovate or pyriform and not contracted
below mouth, neck as long as urn or shorter. Operculum
small, slightly to highly convex, shortly beaked,
brown to dark red-brown, glossy. Annulus broad, to
140 μm, 2-3-rowed. Peristome inserted at urn mouth,
500 μm high. Exostome teeth at base 120 μm wide,
gradually tapered, below yellow, in uppermost part
hyaline; border 12-18 μm wide; median line zig-zag-