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40 J.E. Caracuel et al. / Geobios 39 (2006) 25–42<br />

Fig. 7. Syn<strong>the</strong>tic <strong>Jurassic</strong> successions at Sierra Espuña, with indication <strong>of</strong> bi<strong>of</strong>acies, micr<strong>of</strong>acies <strong>and</strong> <strong>paleoenvironmental</strong> interpretation for <strong>the</strong> main lith<strong>of</strong>acies.<br />

M (mudstones), W (wackestones), P (packstones), G (grainstones). Legend for bioclasts in Fig. 2.<br />

Fig. 7. Succession synthétique du Jurassique de la Sierra Espuña, avec indication des bi<strong>of</strong>aciès, micr<strong>of</strong>aciès et interprétation paléoenvironnementale du lith<strong>of</strong>aciès<br />

principal. M (mudstones), W (wackestones), P (packstones), G (grainstones). Légende des bioclastes Fig. 2.<br />

inner moulds, mostly reworked; frequent loss <strong>of</strong> bodychambers<br />

<strong>and</strong> truncations incompatible with <strong>the</strong> stratification.<br />

In <strong>the</strong> nodular-brecciate facies, inner moulds <strong>of</strong> ammonites<br />

are <strong>of</strong>ten fragmented <strong>and</strong> imbricate.<br />

Finally, at <strong>the</strong> end <strong>of</strong> <strong>the</strong> <strong>Jurassic</strong> <strong>the</strong> generalized <strong>the</strong>rmal<br />

subsidence tends to deepen <strong>the</strong> paleomargin, while <strong>the</strong> huge<br />

relief caused by <strong>the</strong> horst <strong>and</strong> graben system is smoo<strong>the</strong>d.<br />

Consequently, <strong>the</strong> area evolved to a basin in Lower Berriasian<br />

(probably already from <strong>the</strong> uppermost Tithonian) with<br />

deposition <strong>of</strong> periplatform limestones toge<strong>the</strong>r with <strong>the</strong> local<br />

sedimentation by planktonic micr<strong>of</strong>ossils (calpionellids, foraminifers,<br />

radiolarians, as well as algae).<br />

7. Conclusions<br />

A multidisciplinary study has enabled greater precision in<br />

updating <strong>the</strong> biostratigraphic framework <strong>and</strong> <strong>the</strong> <strong>paleoenvironmental</strong><br />

interpretation <strong>of</strong> <strong>the</strong> <strong>Jurassic</strong> succession at Sierra<br />

Espuña. This area, which is one <strong>of</strong> <strong>the</strong> bigger, better exposed<br />

<strong>and</strong> more fossiliferous <strong>Jurassic</strong> outcrops <strong>of</strong> <strong>the</strong> Malaguide<br />

domain, can be considered a clue area to analyze <strong>the</strong> <strong>evolution</strong><br />

<strong>of</strong> <strong>the</strong> Internal Zones <strong>of</strong> <strong>the</strong> Betic Cordillera.<br />

As a whole, ammonite biostratigraphic data from <strong>the</strong> Internal<br />

Zones are scanty, <strong>and</strong> related only to <strong>the</strong> “Dorsal” <strong>and</strong><br />

Malaguide domain such as <strong>the</strong> Sierra Espuña area. Particularly<br />

for this area, <strong>the</strong> previous biostratigraphic data, which<br />

comes from <strong>the</strong> 1960s <strong>and</strong> 1970s, need to be updated <strong>and</strong><br />

revised. Thus, three previously studied <strong>and</strong> two new <strong>Jurassic</strong><br />

sections at Sierra Espuña were sampled, leading to a more<br />

precise biostratigraphic ammonite framework. Ammonite<br />

assemblages have enabled <strong>the</strong> recognition <strong>of</strong> <strong>the</strong> Domerian,<br />

Lavinianum (Cornacaldense Subzone), Algovianum (Ragazzoni,<br />

Bertr<strong>and</strong>i, Accuratum <strong>and</strong> Levidorsatum Subzones) <strong>and</strong><br />

Emaciatum (Solare <strong>and</strong> Elisa Subzones) Zones, <strong>the</strong> Lower<br />

Toarcian, Polymorphum <strong>and</strong> Serpentinum Zones, <strong>the</strong> Middle<br />

Toarcian, Bifrons <strong>and</strong> Gradata Zones, <strong>the</strong> Upper Toarcian,<br />

Reynesi Zone, <strong>the</strong> uppermost Lower/Upper Bajocian, <strong>the</strong><br />

Lower Callovian (Bullatus <strong>and</strong> Gracilis Zones), <strong>the</strong> Middle<br />

<strong>and</strong> Upper Oxfordian (Transversarium, Bifurcatus, Bimammatum<br />

<strong>and</strong> Planula Zones) <strong>and</strong> <strong>the</strong> Lower <strong>and</strong> Upper Kimmeridgian<br />

(Platynota, Strombecki, Divisum <strong>and</strong> Beckeri<br />

Zones). Some <strong>of</strong> <strong>the</strong>se zones <strong>and</strong> subzones are recognized, or<br />

documented with reported faunas, for <strong>the</strong> first time.<br />

As a whole, benthic assemblages dominated during <strong>the</strong><br />

Lower <strong>Jurassic</strong>, while benthonic/planktonic assemblages

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