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A survey of the vegetation of Ben Nevis - Scottish Natural Heritage

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<strong>Scottish</strong> <strong>Natural</strong> <strong>Heritage</strong> Commissioned Report No. 090 (ROAME No. F02LD01)<br />

W9b Fraxinus excelsior-Sorbus aucuparia-Mercurialis perennis woodland, Crepis paludosa<br />

sub-community<br />

There are small, mostly linear patches <strong>of</strong> W9b in <strong>the</strong> gullies which cut down <strong>the</strong> western slope <strong>of</strong> Meall<br />

Cumhann, and which are flushed with base-rich water from <strong>the</strong> limestone above. There are also patches<br />

along <strong>the</strong> banks <strong>of</strong> <strong>the</strong> Allt Coire a’ Mhadaidh in <strong>the</strong> north-east <strong>of</strong> <strong>the</strong> study area. It is a woodland <strong>of</strong> damp,<br />

basic soils and has a canopy <strong>of</strong> ash Fraxinus excelsior, hazel Corylus avellana, birch Betula pubescens, elm<br />

Ulmus glabra, goat willow Salix caprea and rowan Sorbus aucuparia. Under <strong>the</strong> trees is a rich tangle <strong>of</strong><br />

tall herbs and grasses: a colourful spread <strong>of</strong> flowers in summer. The species here include Agrostis capillaris,<br />

Deschampsia cespitosa, Anthoxanthum odoratum, Geum urbanum, Geranium robertianum, G. sylvaticum,<br />

Filipendula ulmaria, Stachys sylvatica, Ranunculus acris, Prunella vulgaris, Sanicula europaea, Primula vulgaris,<br />

Fragaria vesca, Viola riviniana, Angelica sylvestris, Cardamine pratensis and <strong>the</strong> ferns Athyrium filix-femina,<br />

Dryopteris affinis, D. filix-mas and Blechnum spicant. The uncommon grass Melica nutans grows here in this<br />

type <strong>of</strong> <strong>vegetation</strong>. There is a deep carpet <strong>of</strong> bryophytes on <strong>the</strong> ground, over rocks and on <strong>the</strong> bases<br />

<strong>of</strong> trees, mostly made up <strong>of</strong> Thuidium tamariscinum, T. delicatulum, Iso<strong>the</strong>cium alopecuroides, I. myosuroides,<br />

Hylocomium splendens, H. brevirostre, Hypnum cupressiforme, Rhytidiadelphus loreus, R. triquetrus,<br />

Brachy<strong>the</strong>cium plumosum, Racomitrium aciculare, Atrichum undulatum, Plagiomnium undulatum and<br />

Eurhynchium praelongum. Some <strong>of</strong> <strong>the</strong> herb-rich grassland above <strong>the</strong> woodland on <strong>the</strong> slopes <strong>of</strong><br />

Meall Cumhann and Coire Choimhlidh might develop into this kind <strong>of</strong> woodland if grazing were less intense.<br />

W11a Quercus petraea-Betula pubescens-Oxalis acetosella woodland, Dryopteris dilatata<br />

sub-community<br />

This type <strong>of</strong> grassy, acid woodland has a canopy <strong>of</strong> birch Betula pubescens, rowan Sorbus aucuparia and<br />

hazel Corylus avellana. The ground layer is lightly grazed. The sward <strong>of</strong> species such as Holcus mollis,<br />

Agrostis capillaris, Potentilla erecta, Viola riviniana and Conopodium majus is punctuated by tall clumps <strong>of</strong><br />

Dryopteris filix-mas and D. dilatata. In some patches <strong>the</strong>re is also much Rubus fruticosus or R. idaeus, or trails<br />

<strong>of</strong> Lonicera periclymenum. The bryophyte layer is made up <strong>of</strong> species such as Hylocomium brevirostre,<br />

Thuidium tamariscinum, Hylocomium splendens, Pleurozium schreberi and Polytrichum commune. Large ferns<br />

and scrambling shrubs are rare in upland woods unless grazing is light. They are also a much more common<br />

component <strong>of</strong> <strong>the</strong> <strong>vegetation</strong> from <strong>the</strong> south-west Highlands southwards, where summers are warmer than<br />

<strong>the</strong>y are fur<strong>the</strong>r north. So W11a is quite a notable type <strong>of</strong> <strong>vegetation</strong> in <strong>the</strong> study area. It occurs in Glen <strong>Nevis</strong><br />

on islands in <strong>the</strong> river and on <strong>the</strong> very steep slopes <strong>of</strong> <strong>the</strong> gorge.<br />

W11b Quercus petraea-Betula pubescens-Oxalis acetosella woodland, Blechnum spicant<br />

sub-community<br />

This is <strong>the</strong> most extensive type <strong>of</strong> woodland on <strong>the</strong> steep slopes in Glen <strong>Nevis</strong>. It is also a common<br />

component <strong>of</strong> <strong>the</strong> mixed woodlands on steep, rocky slopes along <strong>the</strong> nor<strong>the</strong>rn edge <strong>of</strong> <strong>the</strong> study area. It is<br />

a grassy woodland <strong>of</strong> well-drained acid soils. It is so common in <strong>the</strong> uplands that people seldom realise that<br />

it is far from natural, and that <strong>the</strong> ground layer would be more heathy or more herb-rich were it less grazed.<br />

The canopy is <strong>of</strong> birch Betula pubescens, with smaller amounts <strong>of</strong> oak Quercus petraea, rowan Sorbus<br />

aucuparia, holly Ilex aquifolium, hazel Corylus avellana and even alder Alnus glutinosa and ash Fraxinus<br />

excelsior. Underneath <strong>the</strong> trees, <strong>the</strong> field layer is green and grassy and made up <strong>of</strong> Anthoxanthum odoratum,<br />

Agrostis capillaris, A. canina, Holcus mollis and H. lanatus. This is speckled with small herbs such as<br />

Potentilla erecta, Galium saxatile, Viola riviniana, Primula vulgaris, Blechnum spicant, Oxalis acetosella,<br />

Conopodium majus and Anemone nemorosa. Hyacinthoides non-scripta can be common, stippling <strong>the</strong><br />

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