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A survey of the vegetation of Ben Nevis - Scottish Natural Heritage

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<strong>Scottish</strong> <strong>Natural</strong> <strong>Heritage</strong> Commissioned Report No. 090 (ROAME No. F02LD01)<br />

species such as Nar<strong>the</strong>cium ossifragum, Polygala serpyllifolia and Pedicularis sylvatica. Under <strong>the</strong> vascular<br />

sward is a thin weft <strong>of</strong> bryophytes such as Calliergonella cuspidata, Hylocomium splendens, Sphagnum<br />

fallax, Scleropodium purum and Lophocolea bidentata.<br />

M27 Filipendula ulmaria-Angelica sylvestris tall-herb fen<br />

This type <strong>of</strong> fen <strong>vegetation</strong> occurs around Achriabhach in Glen <strong>Nevis</strong>. It consists <strong>of</strong> tall, leafy swards <strong>of</strong><br />

Filipendula ulmaria, <strong>the</strong>ir stems topped in summer with <strong>the</strong> fragrant pale cream flowers, and an assemblage<br />

<strong>of</strong> o<strong>the</strong>r species including Athyrium filix-femina, Cirsium palustre, Epilobium palustre, Juncus effusus, Galium<br />

saxatile, Rumex acetosa, R. obtusifolius and Cardamine pratensis. There is little space for bryophytes under<br />

<strong>the</strong> dense herbage. There are just a few wisps <strong>of</strong> Calliergonella cuspidata, Rhytidiadelphus squarrosus and<br />

Lophocolea bidentata. This <strong>vegetation</strong> does not fit comfortably into any <strong>of</strong> <strong>the</strong> three sub-communities <strong>of</strong> M27.<br />

Ecologically it is probably closest to <strong>the</strong> Urtica dioica-Vicia cracca sub-community M27b, a lowland<br />

<strong>vegetation</strong> type <strong>of</strong> disturbed soils, as it occurs down-slope <strong>of</strong> <strong>the</strong> sheep fanks and farm track and is likely to<br />

be flushed with nitrogen-rich water.<br />

M31 An<strong>the</strong>lia julacea-Sphagnum denticulatum spring<br />

These small springs – generally though not always associated with snow-beds – are common at higher<br />

altitudes in <strong>the</strong> <strong>survey</strong> area. Individually <strong>the</strong>y are small, but on wet slopes where water seeps out diffusely<br />

<strong>the</strong>y can cover quite a lot <strong>of</strong> ground in total. They occur where <strong>the</strong> flow <strong>of</strong> water is slow or even intermittent,<br />

and commonly fill <strong>the</strong> meltwater channels down-slope from places where snow lies late. They are very distinctive.<br />

They consist <strong>of</strong> grey-green swollen mats and patches made up <strong>of</strong> dense masses <strong>of</strong> <strong>the</strong> tiny cylindrical shoots<br />

<strong>of</strong> <strong>the</strong> liverwort An<strong>the</strong>lia julacea. These mats are studded with <strong>the</strong> red-gold shoots <strong>of</strong> Sphagnum denticulatum,<br />

<strong>the</strong> flat purplish shoots <strong>of</strong> Scapania undulata and with a variety <strong>of</strong> small plants such as Saxifraga stellaris,<br />

Deschampsia cespitosa ssp. alpina, Gnaphalium supinum, Viola palustris, Huperzia selago, Carex bigelowii,<br />

Epilobium anagallidifolium and Armeria maritima. Although usually found where <strong>the</strong> irrigating water is acid<br />

as well as very cold, a few base-tolerant species can grow here, for example Cochlearia pyrenaica ssp.<br />

alpina, Selaginella selaginoides, Carex saxatilis, Thalictrum alpinum and Persicaria vivipara. These springs<br />

are also home to a number <strong>of</strong> scarce montane bryophytes such as Kiaeria starkei, Philonotis seriata,<br />

Pohlia ludwigii, Polytrichum sexangulare, Pleurocladula albescens and Scapania uliginosa.<br />

M32a Philonotis fontana-Saxifraga stellaris spring, Sphagnum denticulatum sub-community<br />

This is <strong>the</strong> less species-rich and diverse <strong>of</strong> <strong>the</strong> two forms <strong>of</strong> this type <strong>of</strong> spring-head <strong>vegetation</strong>. It is <strong>the</strong> more<br />

common <strong>of</strong> <strong>the</strong> two where <strong>the</strong> spring water emerges through acidic, peaty soil, but also occurs on richer<br />

mineral soils where it forms mosaics with <strong>the</strong> Montia-Chrysosplenium sub-community (see below) as well as<br />

with a range <strong>of</strong> o<strong>the</strong>r springs and mires. M32a consists <strong>of</strong> rich orange-gold spreads <strong>of</strong> <strong>the</strong> moss Sphagnum<br />

denticulatum, patched with species such as Polytrichum commune, Philonotis fontana and Scapania undulata.<br />

Rooted in <strong>the</strong> mats <strong>of</strong> mosses are plants such as Viola palustris, Saxifraga stellaris, Huperzia selago,<br />

Juncus squarrosus and Eriophorum angustifolium, and grasses such as Deschampsia cespitosa, Agrostis canina<br />

and Anthoxanthum odoratum. M32a springs are common throughout <strong>the</strong> study area. In <strong>the</strong> upper parts <strong>of</strong><br />

Coire a’ Mhadaidh and and Choire Lèi<strong>the</strong>ith <strong>the</strong>y form networks which make up a considerable proportion<br />

<strong>of</strong> <strong>the</strong> total <strong>vegetation</strong>.<br />

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