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Download Frederic Clements' Book on Plant Succession in PDF ...

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CONVERSION OF FOSEST<br />

bulk of the material is c<strong>on</strong>verted <strong>in</strong>to humus, which gradually compacts itself<br />

<strong>in</strong>to a firm layer. Heath formati<strong>on</strong> <strong>on</strong> such a soil is <strong>in</strong>terestmg for the reas<strong>on</strong><br />

that it may occur without the leach<strong>in</strong>g of the upper layer, and mdeed may be<br />

159<br />

found <strong>on</strong> heavy loam or clay. Grebe describes the acti<strong>on</strong> of the raw humus<br />

up<strong>on</strong> vegetati<strong>on</strong> as follows :<br />

'<br />

(1) The raw humus cuts off the lower soil almost<br />

completely from its air supply. (2) It h<strong>in</strong>ders the circulati<strong>on</strong> of water m the<br />

soil. It prevents the evaporati<strong>on</strong> of superfluous moisture <strong>in</strong> w<strong>in</strong>ter and sprmg,<br />

and <strong>in</strong> summer it h<strong>in</strong>ders the penetrati<strong>on</strong> of light ra<strong>in</strong>s and of dew. (3) It is<br />

probable that the soil beneath the layer of heath-felt passes out of the stage<br />

of oxidati<strong>on</strong> <strong>in</strong>to that of stagnati<strong>on</strong> and reducti<strong>on</strong>. (4) The upper soil layer<br />

is relatively poor <strong>in</strong> dissolved m<strong>in</strong>eral salts, the middle and lower relatively<br />

rich. (5) While the raw humus of the heath is as rich as the humus of the<br />

beech woods and p<strong>in</strong>e woods, it is so firmly comb<strong>in</strong>ed as a c<strong>on</strong>sequence of its<br />

peaty nature that it can not be used by the trees.'<br />

"Grebe has been correct <strong>in</strong> his assumpti<strong>on</strong> that the aerati<strong>on</strong> of the soil is<br />

ialmost completely prevented by the raw hmnus. Accord<strong>in</strong>g to my op<strong>in</strong>i<strong>on</strong>,<br />

this factor sufiices almost entirely al<strong>on</strong>e to make the proper growth of trees<br />

impossible and to call forth sickness, stunt<strong>in</strong>g, or death accord<strong>in</strong>g to the <strong>in</strong>tensity<br />

of its acti<strong>on</strong>."<br />

C<strong>on</strong>versi<strong>on</strong> of forest <strong>in</strong>to moor.— "It is generally recognised that the heath<br />

moor differs from the meadow moor <strong>in</strong> that it is not level but c<strong>on</strong>vex. It<br />

grows upward not <strong>on</strong>ly <strong>in</strong> the middle, but also, even though slowly, at the<br />

marg<strong>in</strong>. Now if such a moor arises <strong>in</strong> a shallow depressi<strong>on</strong>, it slowly pushes<br />

its edges up the slopes. Thus it f<strong>in</strong>ally reaches a gap <strong>in</strong> the surround<strong>in</strong>g hUls,<br />

and it then extends a t<strong>on</strong>gue through the gap <strong>in</strong>to further levels. Thus it<br />

comes about that a lively movement of water is noticed, when the t<strong>on</strong>gue of the<br />

moor lies up<strong>on</strong> slop<strong>in</strong>g ground. S<strong>in</strong>ce the t<strong>on</strong>gue lies lower than the surface<br />

of the moor and the Sphagnum holds the water so firmly that the surplus can<br />

soak <strong>in</strong>to the soil but slowly, the t<strong>on</strong>gue is c<strong>on</strong>stantly dripp<strong>in</strong>g with water,<br />

and <strong>in</strong> most cases a quantity of water flows away from it, as at Kolbermoor.<br />

If the soil of the slope and adjacent lower areas is not especially pernutrient at<br />

its surface, the formati<strong>on</strong> of heath moor proceeds rapidly. The cushi<strong>on</strong>s of<br />

Sphagnum spread more and more widely till they reach the bottom of the low<br />

area always fed with water from above. The bottom <strong>on</strong>ce reached, the c<strong>on</strong>stant<br />

flow furnishes abundant water for further development, unless, as is<br />

frequently the case, a col<strong>on</strong>y of Sphagnum has already occupied the bottom as<br />

a c<strong>on</strong>sequence of the accumulat<strong>in</strong>g water, <strong>in</strong> which event the two masses unite.<br />

Whenever the hollow or the slope and gap are covered with forest, the soil is<br />

c<strong>on</strong>verted <strong>in</strong>to swamp by the Sphagnum and the air is driven out as a result.<br />

The physiological effect up<strong>on</strong> the growth of trees is the same as <strong>in</strong> the formati<strong>on</strong><br />

of raw humus up<strong>on</strong> the forest floor. It is a peculiarly desolate picture<br />

that is formed by the countless dead stand<strong>in</strong>g trunks <strong>in</strong> a young moor. One<br />

trunk after another falls, and so<strong>on</strong> they are all buried <strong>in</strong> the moor, and noth<strong>in</strong>g<br />

visible rema<strong>in</strong>s to rem<strong>in</strong>d <strong>on</strong>e of the former forest.<br />

"In order to exhibit the entire process of the swamp<strong>in</strong>g of a forest, I have<br />

purposely chosen cases <strong>in</strong> which the moor must pass over a small elevati<strong>on</strong>,<br />

s<strong>in</strong>ce the important events <strong>in</strong> the water movement are much clearer than <strong>in</strong><br />

the comm<strong>on</strong> <strong>in</strong>stances. For the most part, the formati<strong>on</strong> of heath moor up<strong>on</strong><br />

meadow moor, or also <strong>in</strong> lowland forest, takes place completely <strong>on</strong> the level<br />

and <strong>in</strong> the follow<strong>in</strong>g manner. The lowlands have become filled with meadow<br />

moors [swamps], as a result of the forland<strong>in</strong>g of p<strong>on</strong>ds and lakes, and the<br />

c<strong>on</strong>sequent development of swamps. The ground level of the swamp slowly<br />

grows upward because of the annual <strong>in</strong>crement of plant rema<strong>in</strong>s, but <strong>on</strong>ly to

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