1 CURRICULUM VITAE John Friedrich Barthell Deans - Biology ...

EDUCATION
1
CURRICULUM VITAE
John Friedrich Barthell
Deans’ Office, 208 Howell Hall
University of Central Oklahoma (UCO)
Edmond, OK 73034-5209
(405) 974-2481 - College of Mathematics & Science
(405) 974-3824 - College FAX
jbarthell@uco.edu
1992 PhD, Entomology, University of California, Berkeley
Dissertation Title: Heterogeneity, invaders, and the population dynamics of some oak
forest solitary bee communities.
1985 B.A., Zoology, University of California, Berkeley
PROFESSIONAL/ACADEMIC POSITIONS
2006-Present Dean, UCO College of Mathematics & Science
2005-2006 Interim Associate Dean, UCO College of Mathematics & Science
2004-2005 Professor, Univ. Central Oklahoma, Biology Dept.
Assistant Dean, Joe C. Jackson College of Graduate Studies & Research
2000-2004 Associate Professor, Univ. Central Oklahoma, Biology Dept.
1995-2000 Assistant Professor, Univ. Central Oklahoma, Biology Dept.
1992-1995 Post Doctoral Researcher, Univ. California, Davis, Entomology Dept.
1986-1992 Graduate R.A. and T.A., Univ. Cal., Berkeley, Entomology Dept.
1985-1986 Staff Research Associate II, Univ. Cal., Berkeley, Entomology Dept.
1981-1985 Laboratory Assistant I, Univ. Cal., Berkeley, Entomology Dept.

Dr. John Barthell 2
TEACHING EXPERIENCE (COURSES TAUGHT AT UCO - 15 YEARS)
“General Entomology” (with laboratory).
“Animal Biology” (includes laboratory).
“General Biology” (with separate laboratory).
“History of Biology” (now “History and Nature of Science”)
“Pollination Biology” (with Dr. Gloria Caddell)
SELECTED PROFESSIONAL PRESENTATIONS (Student Names Boldfaced)
Barthell, J. F., M. L. Clement, H. Wells, K. C. Crocker, E. C. Becker, K. D. Leavitt, B. T.
McCall, M. Mills-Novoa, C. M. Walker, and T. Petanidou. Foraging Patterns of Bees in
Response to Nectar Availability in Populations of the Invasive Thistle Species Centaurea
solstitialis L. in Native (Greece) and Non-Native (USA) Island Ecosystems. Annual Meeting
of the Society of Integrative and Comparative Biology, January 2009, Boston, MA, USA.
(Oral Presentation.)
Krum, C. J., A. N. Savitski, J. M. Hranitz, K. D. Brubaker, J. F. Barthell, and G. D. Yocum.
2008. Comparison of Stress Protein cDNA and Amino Acid Sequences Among Three
Megachilid Solitary Bees. Annual Meeting of the Society for Integrative and Comparative
Biology, January 2008, San Antonio TX, USA. (Poster Presentation.)
Halon, L.T., J.M. Hranitz, K.D. Brubaker, G.D. Yocum, and J.F. Barthell. 2007. Comparison of
stress protein cDNA and Protein Sequences Among Three Species of Bee in Different
Thermal Environments. Annual Meeting of the Society for Integrative and Comparative
Biology, January, 2007, Phoenix, Arizona, USA. (Poster Presentation.)
Barthell, J. F., J. M. Hranitz, R. W. Thorp, L. M. Overall and J. L. Griffith. Heat stress
responses of an invasive leafcutting bee species nesting in the Central Valley of California: a
case of risky maternal behavior? Annual Meeting of the Society for Integrative and
Comparative Biology, December, 2005, San Diego, California, USA.
Barthell, J. F., R. W. Thorp, J. M. Randall, A. M. Wenner, R. T. Reidenbaugh and K. D. Rice.
Positive and negative interactions with non-native insects affect yellow star-thistle
reproduction on Santa Cruz Island, California. 52nd Annual Meeting of the Entomological
Society of America. November, 2004, Salt Lake City, Utah, USA.

Dr. John Barthell 3
REFEREED JOURNAL PUBLICATIONS (Student Names Boldfaced)
Cakmak, I., D. S. Song, T. A. Mixson, E. Serrano, M. L. Clement, A. Savitski, G. Johnson, T.
Giray, C. I. Abramson, J. F. Barthell and H. Wells. 2009. Foraging response of Turkish
honey bee subspecies to flower color choices and reward consistency. Journal of
Comparative Psychology. (Accepted with Revision.)
Cakmak, I., C. Sanderson, T. D. Blocker, S. C. Checotah, L. L. Pham, B. K. Harader-Pate, A.
A. Cro, R. T. Reidenbaugh, P. Nenchev, J. F. Barthell and H. Wells. 2009. Different
Solutions by Bees to a Foraging Problem. Animal Behaviour. (In Press.)
Hranitz, J. M., J. F. Barthell, R. W. Thorp, L. M. Overall and J. L. Griffith. 2009. Nest site
selection influences mortality and heat stress responses in developmental stages of an invasive
leafcutting bee, Megachile apicalis Spinola (Hymenoptera: Megachilidae). Environmental
Entomology. (In Press.)
Amaya-Marquez, M., P. S. M. Hill, J. F. Barthell, L. L. Pham, D. R. Doty and H. Wells. 2008.
Learning and memory during foraging of the blue orchard bee, Osmia lignaria Say
(Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society. 81: 315-327.
Barthell, J. F., R. T. Reidenbaugh and J. L. Griffith. 2006. Persistent size and behavioral
variation among males of the large carpenter bee, Xylocopa virginica (Hymenoptera:
Apidae). Southwestern Entomologist. 31: 223-232.
Barthell, J. F., and T. A. Baird. 2004. Size variation and aggression among male Xylocopa
virginica (L.) (Hymenoptera: Apidae) at a nesting site in central Oklahoma. Journal of the
Kansas Entomological Society. 77: 10-20.
Hranitz, J. M., and J. F. Barthell. 2003. Levels of heat shock protein 70 during development of
the alfalfa leafcutting bee, Megachile rotundata (Hymenoptera: Megachilidae).
Southwestern Entomologist. 28: 97-103.
Barthell, J. F., J. M. Hranitz, R. W. Thorp and Maria K. Shue. 2002. High temperature
responses in two exotic leafcutting bees: Megachile apicalis and M. rotundata
(Hymenoptera: Megachilidae). Pan-Pacific Entomologist. 78: 235-246.
Barthell, J. F., J. M. Randall, R. W. Thorp, and A. M. Wenner. 2001. Promotion of seed set in
yellow star-thistle by honey bees: evidence of an invasive mutualism. Ecological
Applications. 11: 1870-1883.
Barthell, J. F., D. M. Bromberger, H. V. Daly and R. W. Thorp. 1999. Nesting biology of the
solitary digger bee Habropoda depressa (Hymenoptera: Anthophoridae) in urban and island
environments. Journal of the Kansas Entomological Society. 71: 116-136.
Frankie, G. W., R. W. Thorp, L. E. Newstrom-Lloyd, M. A. Rizzardi, J. F. Barthell, T. L.

Dr. John Barthell 4
Griswold, J. - Y. Kim and S. Kappagoda. 1998. Monitoring solitary bees in modified
wildland habitats: implications for bee ecology and conservation. Environmental
Entomology. 27: 1137-1148.
Barthell, J. F., G. W. Frankie and R. W. Thorp. 1998. Invader effects in a community of cavity
nesting megachilid bees (Hymenoptera: Megachilidae). Environmental Entomology. 27:
240-247.
Barthell, J. F., T. L. Griswold, G. W. Frankie and R. W. Thorp. 1997. Osmia (Hymenoptera:
Megachilidae) diversity at a site in central coastal California. Pan-Pacific Entomologist. 73:
141-151.
Barthell, J. F. and J. H. M. Knops. 1997. Visitation of evening primrose by carpenter bees:
evidence of a “mixed” pollination syndrome. Southwestern Naturalist. 42: 86-93.
Knops, J. H. M. and J. F. Barthell. 1996. Flower abundance in a population of sky lupine
(Lupinus nanus) over three years in central coastal California. Madroñ o. 43: 85-92.
Barthell, J. F. and R. W. Thorp. 1995. Nest usurpation among females of an introduced leafcutter
bee, Megachile apicalis. Southwestern Entomologist. 20: 117-124.
Barthell, J. F. and R. Stone. 1995. Recovery of the parasite Triarthria spinipennis (Meigen)
(Diptera: Tachinidae) from an inland California population of the introduced European
earwig. Pan-Pacific Entomologist. 71: 137-141.
Barthell, J. F. and H. V. Daly. 1995. Male size variation and mating site fidelity in a population
of Habropoda depressa Fowler (Hymenoptera: Anthophoridae). Pan-Pacific Entomologist.
71: 149-156.
Gordon, D. M., J. F. Barthell, R. E. Page, M. K. Fondrk and R. W. Thorp. 1995. Colony
performance of selected honey bee (Hymenoptera: Apidae) strains used for alfalfa
pollination. Journal of Economic Entomology. 88: 51-57.
Frankie, G. W., L. Newstrom, S. B. Vinson and J. F. Barthell. 1993. Nesting-habitat preferences
of selected Centris bee species in Costa Rican Dry Forest. Biotropica. 25: 322-333.
Thorp, R. W., G. W. Frankie, J. F. Barthell, D. Gordon, L. Newstrom, T. Griswold, J. Schmidt
and S. Thoenes. 1992. Long-term studies to gauge effects of invading bees. California
Agriculture. 46: 20-23.
Frankie, G. W., S. B. Vinson, L. E. Newstrom and J. F. Barthell. 1988. Nest site and habitat
preferences of Centris bees in the Costa Rican dry forest. Biotropica. 20: 301-310.
Frankie, G. W., J. W. Brewer, W. Cranshaw and J. F. Barthell. 1987. Abundance and natural
enemies of the spindle gall midge, Pinyonia edulicola Gagne, in natural and urban stands of

Dr. John Barthell 5
Pinyon pine in Colorado (Diptera: Cecidomyiidae). Journal of the Kansas Entomological
Society. 60: 133-144.
Frankie, G. W., J. B. Fraser and J. F. Barthell. 1986. Geographic distribution of Synanthedon
sequoiae host plant suceptibility on Monterey pine in adventive and native stands in
California (Lepidoptera: Sesiidae). Pan-Pacific Entomologist. 62: 29-40.
BOOK CHAPTERS, ESSAYS, PROCEEDINGS & REVIEWS (Student Names Boldfaced)
Clement, M., J. Lea, L. Pham, B. Smith, D. Smith, N. Wagner, C. Simmons and J. Barthell.
2009. Hypatia of Alexandria: a comparative perspective on her role in history. Math
Horizons. (In Press.)
Barthell, J. F. 2007. UCO leader in residence: one dean’s perspective. New Plains Review. 8:
15-21.
Barthell, J. F., M. L. Clement, D. S. Song, A. N. Savitski, J. M. Hranitz, H. Wells, T.
Petanidou, R. W. Thorp, A. M. Wenner and T. L. Griswold. 2008. Nectarivore-plant
interactions on Santa Cruz Island (USA) and the Aegean island of Lesvos (Greece):
implications for yellow star-thistle invasion. Seventh California Islands Symposium.
Institute for Wildlife Studies, Arcata, California, USA. (In Press.)
Wenner, A. M., R. W. Thorp and J. F. Barthell. 2008. Biocontrol and eradication of feral honey
bee colonies from Santa Cruz Island, California: a summary. Seventh California Islands
Symposium. Institute for Wildlife Studies, Arcata, California, USA. (In Press.)
Barthell, J. F. and H. Wells. 2007. Honey bee globalization: why the world should not be flat.
Business Review Journal of the St. Joseph’s College of Business Administration. 2: 51-63.
Barthell, J. F., R. W. Thorp, A. M. Wenner, J. M. Randall and D. S. Mitchell. 2005. Seed set in
a non-native, self-compatible thistle on Santa Cruz Island: implications for the invasion of
an island ecosystem. In D. K. Garcelon and C. A. Schwemm, editors. Pp. 185-191. Sixth
California Islands Symposium. Institute for Wildlife Studies, Arcata, California, USA.
Barthell, J. F. 2003. The honey bee in America: An organism of contradictions. In Y. A.
Schnoeker-Shorb and T. L. Shorb, editors. Blessed “Pests” of the beloved west: an
affectionate collection on insects and their kin. Native West Press, Prescott, Arizona, USA.
Barthell, J. F., R. W. Thorp, G. W. Frankie, J.-Y. Kim and J. M. Hranitz. 2003. Impacts of
introduced solitary bees on natural and agricultural systems: the case of the leafcutting bee,
Megachile apicalis Spinola (Hymenoptera: Megachilidae). In K. Strickler and J. H. Cane
and J. Brown, editors. Pp. 151-162. For non-native crops, whence pollinators of the future?
Thomas Say Publications in Entomology, Entomological Society of America, Lanham,
Maryland.
Barthell, J. F., R. W. Thorp, A. M. Wenner and J. M. Randall. 2000. Yellow star-thistle,

Dr. John Barthell 6
gumplant and feral honey bees on Santa Cruz Island: a case of invaders assisting invaders.
Pages 269-273. In D. R. Browne, K. L. Mitchell and H. W. Chaney (eds.), Fifth California
Islands Symposium. Santa Barbara Museum of Natural History, Santa Barbara, California.
MBC Applied Environmental Sciences.
Wenner, A. M., R. W. Thorp and J. F. Barthell. 2000. Removal of European honey bees from
Santa Cruz Island ecosystem. Pages 256-260. In D. R. Browne, K. L. Mitchell and H. W.
Chaney (eds.), Fifth California Islands Symposium. Santa Barbara Museum of Natural
History, Santa Barbara, California. MBC Applied Environmental Sciences (under contract of
U.S. Dept. of the Interior).
Thorp, R. W., A. M. Wenner and J. F. Barthell. 2000. Pollen and nectar resource overlap among
bees on Santa Cruz Island. Pages 261-268. In D. R. Browne, K. L. Mitchell and H. W.
Chaney (eds.), Fifth California Islands Symposium. Santa Barbara Museum of Natural
History, Santa Barbara, California. MBC Applied Environmental Sciences (under contract of
the U.S. Dept. of the Interior).
Thorp, R. W., A. M. Wenner and J. F. Barthell. 1994. Flowers visited by honey bees on Santa
Cruz Island. Pages 351-365. In W. L. Halverson and G. J. Meander (eds.), Fourth California
Islands Symposium: Update on the status of resources. Santa Barbara Museum of Natural
History, Santa Barbara, California.
Vinson, S. B., G. W. Frankie and J. Barthell. 1993. Threats to the diversity of solitary bees in a
neotropical dry forest in Central America. Pages 53-81. In Hymenoptera and Biodiversity.
J. LaSalle and I. D. Gauld (eds). CAB International, Wallingford, UK.
Frankie, G. W., S. B. Vinson, L. E. Newstrom, J. F. Barthell, W. A. Haber and J. K. Frankie.
1990. Plant phenology, pollination ecology, pollinator behaviour, and conservation of
pollinators in a neotropical dry forest. Pages 37-47. In K. S. Bawa and M. Hadley, editors.
Man and the Biosphere Series, Volume 7, Reproductive Ecology of Tropical Forest Plants.
UNESCO, Paris, France and Parthenon Publishing Group, Park Ridge, New Jersey, USA.
RECENTLY FUNDED OR PENDING GRANTS
“REU SITE: Ecology and behavior of honey bees and solitary bees”. 2009. National Science
Foundation. PI = J. F. Barthell, Co-PIs = H. Wells and C. I. Abramson. $205,680.00.
(Pending)
“STEM Double Bridge: Connecting High Schools, Community Colleges, and Universities for
Tomorrow’s Leaders in Science, Technology, Engineering, and Mathematics”. PI = W. R.
Chen, Co-PIs: J. F. Barthell and G. M. Wilson. $1,998,750.00. (Pending)
“REU SITE: Behavior and ecology of honey Bees and leafcutting Bees”. 2006. National
Science Foundation. PI = J. F. Barthell, Co-PIs = H. Wells and C. I. Abramson.
$188,680.00. (Funded)

Dr. John Barthell 7
“Nectar Standing Crop and Plant Competition for Pollinators: Testing a Model of Plant
Invasion”. 2005. UCO Faculty Grant. PI = J. F. Barthell. $7,113.08. (Funded)
“Nonnative Bee Pollination of a Native Plant: A Perturbation Experiment”. 2004. UCO Faculty
Grant. PI = J. F. Barthell. $5,093.72. (Funded)
“Supporting Undergraduate Research and Education for the Science, Technology, Engineering
and Mathematics Talent Expansion” (SURE-STEP). 2004. National Science
Foundation. PI = J. F. Barthell (formerly S. N. Rao). $1,999,050.00. (Funded)
AWARDS
Herbert S. Dordick UCO Outstanding Mentor Award
Sigma Xi Outstanding Researcher Award - Univ. Central Oklahoma Chapter
Hauptman Fellow for College of Mathematics and Science
Neely Excellence in Teaching Award