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pQ&C.i?J <strong>pq</strong><br />

The fruit flies of the Canary Islands (Diptera: Tephritidae)<br />

BERNHARD MERZ<br />

stand. Copenhagen, Denmark. July 1992. ISSN OO13-87l1.<br />

Merz, B.: The fruit flies of the Canary Islands (Diptera: Tephritidae). Ent. scand. 23: 215-231.<br />

This study updates our knowledge of the Tephritidae of the Canary Isiands. The 30 recorded spe-<br />

cies are keyed and briefly discussed. iephritis lureipes sp. n. is described and Trupanea tubulata<br />

Munro is recorded for the first time from the Canary Islands. Three records are regarded as er-<br />

roneous: Aciura coryli (Rossi), Dectodesis augur (Frauenfeld) and Paroxyna absinthii<br />

(Fabricius). Tephritis reticulata Becker, 7: duplex Becker and Paroxyna rnulrirnuculosu Dirlbek<br />

& Dirlbek are transferred to Cumpiglossu (combs. n.). The following new synonymies are pro-<br />

posed (junior synonyms in parantheses): Capititesramulosa (Loew) (= Urelliaperfecta Becker),<br />

Campiglossa duplex (Becker) ( = Paroxyna aequalis Hering), Campiglossa reticulata (Becker)<br />

( = Ephritis occuíta Becker). Oedaspis quinquefasciata Becker ( = Oedaspis heringi Hendel),<br />

Ephritis praecox (Loew) (=íéphritispoecilura Loew). Lectotypes are designated for Paroxyna<br />

aequalis Hering, iephritis reticulata Becker, Tetanocera walkeri Wollaston and Pyperapraecox<br />

Loew. Seven fruit fly species were reared for the first time, and 12 new hit fly - piant reiation-<br />

ships are mentioned. Furthermore Paroxyna martii (Becker) and Urelliosomn guimari (Becker)<br />

are recorded for the first time from Southern Spain and iephritispraecox from Austria.<br />

B. Merz, Entomologisches Institut, Swiss Federal Institute of Technology, 8092 Zürich, Switzer-<br />

land.<br />

Introduction separara Becker, which is a leaf miner in several<br />

The first comprehensive study of the Diptera of the<br />

Canary Islands listed 28 fruit fly species (Becker<br />

1908a). The number of species was increased to 35<br />

by Hendel(1927), Frey (1936), Hering (1937), Frey<br />

(1958a), Dirlbek & Dirlbek (1969). Leclercq (1973)<br />

andchandler (1979). A check list ofall fruit flyspe-<br />

cies of Macaronesia was given by Dirlbek & Ohrn<br />

(1982).<br />

A careful study of the Canarian fruit flies reveals,<br />

that only 30 species and 19 genera are currently<br />

known to occur on these islands. Five of the earlier<br />

mentioned species are synonymized and three other<br />

species (Aciura coryli (Rossi) Paroxyna absinrhii<br />

(Fabricius) and Dectodesis augur (Frauenfeld)) are<br />

removed frorn the list of Dirlbek & Ohrn (1982) be-<br />

cause they were misidentified.<br />

Despite good faunistic knowledge of the fruit<br />

flies of the Canary Islands there are only scattered<br />

data on host plants and other ecological<br />

parameters. The only two species which have been<br />

reared frorn plants on the Canary Islands are<br />

Oedaspisqutnquefasciata Becker from stem galls of<br />

Schizogyne sericea (Hendel 1927) and Eirleiu<br />

0 Entomolooica rcandmi.r~ca itirp 6)<br />

Umbelliferae (= Apiaceae) (Hering 1927).<br />

The following study presents host plant data for<br />

15 species which were al1 reared on the Canary Is-<br />

lands. There are host plant data for a further 10 spe-<br />

cies frorn outside of the Canary Islands. Four spe-<br />

cies were swept in large nurnbers on pure stands of<br />

certain plants, indicating that they may be hosts.<br />

One species. Ensina decisa Wollaston, has no host<br />

data.<br />

Material and methods<br />

This study is rnostly based on sorne 1300 specimens<br />

collected by the author on two islands, Gornera and<br />

Tenerife, during two excursions (28 April to 5 May<br />

1988and3 to IIMarch 1990). Theflieswerecollect-<br />

ed by sweeping and by rearing frorn infested plant<br />

material. In the latter case the plants were stored in<br />

paper bags which were brought into the laboratory<br />

at room ternperature. A11 flies ernerged within two<br />

months. The endernic plants were identified with<br />

Brarnwell & Bramwell(1974), the other plants with<br />

the Flora Europaea (Moore et al. 1976). Addi-<br />

tional material a me frorn the collection of T. Beck-<br />

c


*<br />

er (deposited iii the Zoologischcs Museuni der<br />

Huiiiboldt Uiiivcrsitat, Berlin (ZMHB)), the<br />

Naturhistorischcs Museum, Wien (NHMW), the<br />

Natural History Museum, London (NHM), tlie Te1<br />

Aviy University (TAU), the Zoologisch Museum,<br />

Amsterdam (ZMA) and the Zoologisk Museum,<br />

Copenliagen (ZMC). Most flies examined were col-<br />

lectedoti Teiierifeand Gomera, and thereis little in-<br />

Iortiiatioii ironi thc two descrt islaiids Fucrtevcii-<br />

iiiIa aiid Laiizaiote (Frcy 195ki).<br />

To characterizc each species froiii tlie ecological<br />

poiiit of vicw, tlie plaiit coiiiniuiiity for each record<br />

was iioted. According to Oberdorfer (1965) five<br />

plaiit coiiiiiiunities wcre recognized: semi desert<br />

slii-u b--coiii ni ii iii t y (-v~op/iy//ur>t-%o//;~o~eriu<br />

coiiiiiiuiiity). succuleiit shrub-coiiiiiiuiiity (Klci-<br />

rtiu-Euphorbiori), lauropliyll forest (Lourion<br />

mucuroriesiuiri). piiie forests (/+!->ion curiuriense)<br />

aiid siibalpiiic slirub-coiniiiuiiity (Rc/wtiu co~niiiu-<br />

iiity). For tliis stiidy two additional coniiiiunities<br />

werc distiiiguishcd: uiiciiltivatcd arcas with thistlcs<br />

(Gu/uc/iles -Curc/utcs coiiiniuiiit y) aiid ciilt ivatcd<br />

arcas (oraiiges, baiiaiias, hotel gardcns).<br />

Tlie iiiorpliological teriiiiiiology used hcrc fol-<br />

IOWS McAlpiiic (1981) willi tlie modificatioiis proposed<br />

by Foote & Steyskal (1987). Thc iioiiieiiclaiiiic<br />

;iccords with Foote (1984).<br />

Key ío speiirs<br />

Thc following kcy 10 al1 species is artificial, intend-<br />

ing to facilitate identification. For ahbreviatioiis of<br />

wing vciiatioii and cclls sce Fig. l.<br />

1. Cliacioiaxy of Iicad aiid iliorax reduced. ilic followiiig<br />

sciac absciit: ocellar. posiocular,<br />

presuiural supraalar, posiproiioial, dorsocciiiral,<br />

anterior supraalar, acrosiichal. basal scuiellar<br />

and katepisternal. Three pairs of black scapular<br />

seiae prewiit. lacc on lower pari wiili oiie<br />

round black spoi on cach side of anieiinal<br />

groove. First flagellomere niore than three iinies<br />

as long as widc. Wiiig: cell bin iwice as wide as<br />

ccll cup; wiih a small hyaliiic subapical spoi,<br />

............ Dacus oleae<br />

- Chaetotaxy of head aiid thorax different. no1 al1<br />

abovc iiiciiiioned setae absent; al lcast posi-<br />

proiiotal, dorsoceiiiral. presuiural supraalar<br />

aiid acrosiichal setae preseni. Firsi flagclloiiicre<br />

Icss than three times as loiig as wide. Wing: ccll<br />

biii iisiially as wide ascell cup; wiiig pattern vari-<br />

ahlc. ...................................... 2<br />

2. OiiIy oiic orbiial scia prcsciii, wliicli i\ always<br />

tilack ...................................... 3<br />

- l'wo orbiial sctac prcseiii.. ................... 5<br />

3. Ocellar aiid anterior scuiellar setae absent; orbi-<br />

ial sciac coiivergeiii. Wing paiiern black, wiili<br />

EN'I: SC'ANI). VOL.. 232 (1992)<br />

iwo hyaline indeniaiions al anterior margiii,<br />

iliree hyaline indeniaiioiis at posterior margin,<br />

iwo hyaline spots in cell dm and one hyaline spoi<br />

in cell r,+!. Head: proboscis capitaie; eye 1.5-2 '<br />

times as high as wide. Thorax and abdomen shi-<br />

iiy black. Legs: al leasi fin basal ihree qiiariers<br />

black. ...................... Oxyaciura iibialis<br />

- Ocellar and anterior scuiellar seiae present; orbi-<br />

ial seiae reclinaie. Wing patiern less extensive,<br />

posterior of cell cual t hyaline. Head: proboscis<br />

geiiiculate; eyc lcss thaii 1.3 times as high as<br />

wide. Tliorax not shiiiy black. Abdomen:<br />

posterior part of tergitcs witli a iiarrow ycllow<br />

band. Legs: f yellow. ........................ 4<br />

4. Two frontal setae preseiii. Wing (Fig. 2): ccll<br />

cup withoui disial exiension; disial section of<br />

veins R4rS and M convcrgent; wing palterii<br />

haiidcd. Head: proboscis siroiigly geiiiculaie,<br />

tip of hausielluni exceeding anterior margiii of<br />

palpi. Thorax and scuiellum cniirely black. only<br />

notopleiiral stripe yellow<br />

- Tliree frontal setae preseni. Wiiig (Fig. 3): cell<br />

cup with shori disial extciision; veins R4, and<br />

M f pnrallcl; wing wiili a rediiccd reiiculaic pat-<br />

ierii ni ilie apical pari. Head: proboscis horier,<br />

iip of Iiaustelliim noi exceeding anterior margiii<br />

of palpi. Tliorax aiid scuiclliiiii ycllow; SIII~UI~<br />

witli a dull black pollinosity. ....... Ensino derisa<br />

5. Only basal scuiellar seiae prcsciii; if aiiicrior<br />

scuicllar setac preseiii, tlieii they are slcndcr and<br />

sliortcr ilian onc quarier the leiigih 01 basal<br />

sciitcllar sctac; posterior orbiial scia usually<br />

whitc. ........ ....................... 6 2<br />

- l'wo pairs of s setae preseni; anierior<br />

sciiiellar seiae at leasi oiic ihird as long as basal<br />

sciiiellar setae; posterior orbiial seia variahlc. .. 15<br />

6. llsually only Iwo fronial sciac preseiil; al leasi<br />

some postocular setae black. Legs: at least f,<br />

partly darkened. Head: proboscis geiiiculate or<br />

spaiulate. Wing pattern: ccll c ai leasi centrally<br />

with dark spoi; iiever witli a dark star-likc sub-<br />

apical spoi. p : oviscapc ai basc witlioui wliiic<br />

hairs. ..................................... 7<br />

- Three frontal setae present. ilie anterior one in<br />

Goriiiirellia longirauda very shori and wliite or<br />

rarely absent; al1 postocular setae white. Legs<br />

eiitirely yellow except for Trupanea stellaia,<br />

wliich has someiimes partly darkeiied f. Head:<br />

proboscis always capitaie. Wing paiiern: cell c<br />

eiitirely hyaline; sometimes wiih a dark star-like<br />

subapical spot. Q : oviscape al base with wliite<br />

hairs. ..................................... 9<br />

7. I'roboscis spaiulaie; posierior iioiopleural scia<br />

white; katepisternal seia pale. Abdomen: tcr-<br />

giies iiniformly black. ihe lasi tergite shiiiing<br />

black. Wing pattern strongly reticulate lo ihe<br />

anal lobe; wiih an apieal fork; cell rl witli two lo<br />

thrcc hyaliiic spois, the basal one cxtenditig inio<br />

ccll r2 + ,.<br />

.................... Spalhuliria siculu<br />

- I'rohoscis geniculate; posterior notopleural seta<br />

and kaiepisiernal seta black. Abdomen: iergiies<br />

browii 10 grey, wiih iwo rows of dark spois.<br />

Wing pattern different. ........<br />

8. Wiiig paitern (Pig. 4) dark browii, forming two<br />

crossbands; wiihout apical fork; the middle of<br />

-.-,<br />

7<br />

l<br />

biiiii Ilick ol C'aiiary Islaiids 217<br />

Figs 1-8. Wings of Tephriiidae of ihe Canary Islands: (1) Terellia sp.; (2) M.vopiies nigrescens; (3) Ensino decisa; (4)<br />

Paroxyna rnarlii; (5) Dioxyrra sororcula; (6) Trupafrea insulariiin; (7) Urc~llio.wina ptinrari Q ; (8) Tephriiis luieipes.<br />

Scale bars = 1 mni.<br />

ilie ihrec hyaliiic spois iii ccll rl reaclics ilie<br />

posierior margiiiof ilieccll. iheothertwospois<br />

aresmallerand touchoiilyC;cell c browii. wiili<br />

one lo two sinall Iiyalinc spots. Head (lig. I 1):<br />

postocular setac iiiosily black. Thorax: scuiiiiii<br />

greyish. but notoplcural siripc largely browii<br />

lo black. Legs: al1 cx dark. ...... Paroxyna m wii<br />

- Wing paitern reiiculaic (Fig. 5). palc grcyisli,<br />

csp. posicrior of vciii CuAl; al1 Iiyaliiie spois in<br />

cell rl of ilie saiiic sizc, reaching the posterior<br />

niargiii of ilie cell; cell c hyaline, only cenirally<br />

with a very indisiinct dark spot. Head (Fig. 12):<br />

the long poslocular seiae are white. thc shori<br />

ones black. Thorax: scuium uniformlygreyish.<br />

wiili soiiic indisiiiici Iio.ryia sororcrila<br />

4<br />

8


Y. Aiiierior lrontal scki sliort aiid wliitc. ilic<br />

posier ior Iwo íroiiial sctac Ioiigcr aiid dwker.<br />

Wiiig paiterii wiili aii apical forh; cell rl wiili<br />

IWO Iiyaliiic iiidciiiaiicws, tlic basa1 IC iriaiigiilar,<br />

ilic apical oiic narrow, stripc likc; ccll din<br />

posiciior of r-iii crossvcin with oiic ray wliich<br />

crosscs uppcr halíoí ihccell. Abdoiiieii: at Icasi<br />

posterior iiiargin oí tergitcs ycllow-brown. p :<br />

oviscapc as ioiig as tlic lasi thrcc iergitcs coiii-<br />

Iiiiicd. oiaiigc-browii. exccpt for black ;ipcx. , ,<br />

....................... Goriiurelliu iongicuicrlu<br />

- l'liree or niore coiicoloiiroiis frontal setae prcsciit.<br />

Wiiig paiiern: wiili or witlioiit iiii apical<br />

lork, ccll rl variable. p : oviscape black. ...... 10<br />

10. Wiiig hyUliiic. wiih ycllow-brown, rcduced pattcrii:<br />

posterior oí vcin ni aiid basd Iialfoí wiiig<br />

Iipnliiic. oiily in cr aloiig tliii-cu crcissvciii wiiii<br />

wi;ill riioi: wiilioiii apical lork. ............<br />

.................... A~~~rri/liio/ihilir.\ hdiori/lii<br />

/'<br />

LY 10<br />

12<br />

- Wiiig pattcrii rnorcconiplctc: at Ic;isi ccll 111 wiih<br />

two dark rays wliich rcacli hiiid iiiargiii oí wiiig. 1 I<br />

I l. Wing (Fig. 6): witli a shori aiiicai lork; ccll rl at<br />

Ii;isc with oiic iiarrow Iiyaliiiciiidcntaiioii. whicli<br />

ismuclilongcrihan wide:arcaaiiicrioroívcin M<br />

and disial ofsc is niosily ckirk. Knob of lialicrcs<br />

darkciied. ................. Trupurrea insulurrrrn<br />

- Wiiig pattern: without apical íork; ccll rl citlier<br />

ciiiirely hlack orwiihoneor iwowidcliyaliiiciiitlciilalioiis.<br />

Kiiob of Iialicrcs yellow. .......... 12<br />

12. Lasi iergite oíabdomcn shiny black. Wing paiicrii:<br />

with a large hlack arca, coveriiig inost of<br />

ccll sc. apical hall of ccll br, niosi of cell diii.<br />

r4+ aiid in; cell rl in p (1:ig. 7) without hyalinc<br />

spot. iii U one large hyalinespot prcseni; ai apex<br />

oí wing with a iiarrow cresccnt shapctl hp;iliiic<br />

iirea; wiilioiit a star sliiiprd suhapical spot. ...<br />

......................... lJrellio~sorrr~t,~uirirori<br />

Al1 Ictgiic\gi-cy. usiiallyilciisely polliiiosc. Wiiig<br />

I:N 1. SC'ANI). VOI . ;3:2 (IY02)<br />

LJ<br />

16<br />

13 14<br />

17 18<br />

15<br />

20


220 Afivz, 11.<br />

-<br />

veiii rcaclies Iiiiid iiiargin oí wing; coiiiiiiiiaiioii<br />

oí r-in crossveiii joiiis vein C on or apical oí<br />

brown ray in cell rl; brown ray in cell dm usual-<br />

ly crosses tlie whole cell. Seiae in both sexes oí<br />

........................... 14<br />

el1 r2 coinpletcly browii<br />

hyaline spoi iii ccll rI; eiid oí<br />

vcin K, rS iiormally hyaline. al mosi wiih<br />

browii doi. .................. Triipatieo oiiioeiia<br />

~ Wiiig pattcrii: disial Iiyaliiie spot oíccll rl cxtciids<br />

iiito cell r2+3, or. ií noi. in ccll r2,,<br />

postcrior oí hyaline spot oícell rl with a srnall<br />

hyaliiic spoi; al cnd oí vein R,, wiih a dark<br />

spot ........................ Trupanea iubuloío<br />

15. Wiiig (I‘ig. 1) cxccpi íor ihc yellow sc cniircly<br />

hyaliiic. Posterior orbital setae convergeni;<br />

three concolourous Cronial setae presenl.<br />

Head: palpi exceeding anterior margin of facial<br />

iiiargiii. Thorax: aiiepisteriiuni wiih brown<br />

loiigiiudiiial stripe. Abdomen: icrgites 3-5<br />

each with two pairs oí black spots.. ... Terelliu sp.<br />

- Wiiig wiili a disiinct paiterii. Uoili orbiial sciac<br />

reclinatc. Abdomen: icrgiles unicolourous,<br />

or al iiiost with oiic pair oí hlack spois cciiirally.<br />

.................................... 16<br />

16. K4,<br />

sctulosc above. sciiilae extciid bcyoiid<br />

r-ni crossvein. Legs eniirely ycllow. 130th orbi-<br />

ial sciae dark; postocular seiae slender, black.<br />

f;roiis with dark seiulae. Two pairs of siroiig<br />

scapular seiae preseni. f yellow. 9 : 6th iergite<br />

distiiictly shorter iliaii 5ih. .................. 17<br />

-- K, :ibovc al inosi with fcw hairs al iiode.<br />

lroiis hare or wiih íew pale seiulae. Posterior<br />

orhital sciae white (oiily iii S~~ltaeiii.scirs~i/ilinl


oficii Inkiiig. Wiiig paiicrii (Fig. 4): wiilioiii<br />

siihliynliiic spois; Iiyaliiic arca bciwccii sribapical<br />

aiid iipiciil cio~sh;iiid fortiicd by a row of<br />

5-6 largc Iiyalinespois. eacli coveriiig ilie widtli<br />

of a ccll; iheanierior three big spois are siiuated<br />

iii ilic coiiiiiiiiatioii of diii-cii crossveiii io C.<br />

Siii:i11 *pccics. wing Iciigili 1.7~2.5 iiiiii. .....<br />

............................. I'uru.\:ynu rnuríu<br />

- I'roboscis gciiiculaic or capiiate, iip of liausiclliiiii<br />

iieicr exceediiig aiiicrior iiiargiii of facial<br />

iiiargiii. Aiiierior scuiellar scine iisiially ai<br />

Icasi ilirecqiiariers as loiigas basa1 sciiicllar seiiic.<br />

Wiiig paiicrii: t>ficii wiili sul~liynliiies~iois;<br />

Iiy;iliiie h:iiid hciwecii crosshaiids foriiied hy a<br />

Iiy;iliiie arca or by a row of sevcral indisiiiici,<br />

siibliyalinc rpois. Specics largci, wing Iciigili<br />

over 3.0 iii ni. .............................. 27<br />

27. Sqiiaiiiae d;irkeiied. Al1 posiociiliir sciüe dark.<br />

Ahdoiiicii: al1 lergiics exccpi l.or ilic firsi iwo<br />

covcrcd witli black hairs. Sciiiuiii iii basai haif<br />

wiili 5 loiigitudiiial stripes, ilic iwo laicral<br />

siripcs tlieiargest. Wing(l:ig. 22)wiih apaiiern<br />

01 Iiyalinc aiid subliyaliiie spois wliich foriii a<br />

subapical aiid apical crossbaiid. Proboscis<br />

ciipiiaie. Q : oviscapc browii. ... Orolasu crrhrutu<br />

Sqii;iiiiac ycllow. Al Ic;isl soiiic po\iociiler sciac<br />

wliiic. hlidoiiicn: al1 tcrgiics wiili sonic<br />

white Iiaiis. scuiuiii WilliOUl ioiigiiudiiial<br />

siripcs. Wiiig: subapieal atid apical crossbaiid<br />

arescparated bya iargc Iiyaliiicarea. Proboseis<br />

spaiiilale or griiiculaic. p : oviccape biack. .... 2H<br />

28. kuiclluiii coiivcx. sliiny hlack. Wiiig paticrii<br />

hlack (lig. 23); cell cual and inlal Iobc Wiili<br />

Iaigc hl:ick aica. I'rohoscis spaiulatc. .......<br />

only rligliilv itie dark arca; Iiyaliiic spoi iii i4 ~<br />

aiiierior of diii-cu crossveiii siiiali. usii;illy iioi<br />

exccediiig ihc middlc of ilie cell; apical spot iii<br />

rrII i , olicii ahwii. Thorax: longitiidiii;il<br />

bNI. SC'ANI). \'O1 . 23:2(1092)<br />

siripcs oii scuiuiii disíiiici. . Cunipigloissu rrricicluru<br />

- Q : aculeus wiilioiii prcapical sicp aiid apical<br />

iiidentatioii (Figs 17 & 111). Wiiig patterii liglii<br />

brown 10 brown; cells r2+! proxiinal of r-m<br />

crossvein aiid br usually wiili 1-3 large spois<br />

wliich cover mosi of ilie ccll; Iiyaiiiie spoi iii ceil<br />

r4 anterior oí dni-cu crossveiii largc. usually<br />

wider ihan half oí tlic cell; apical spoi in cell<br />

r4, always presetii. Tliorax: longiiudiiial<br />

stripes oii scutuni oftcii indisiiiici or abseiii.<br />

... 32<br />

32. Q: iip of aculeus nariowly roiinded (Fig. 17).<br />

Scuielliim ai apex witli a yellow spoi, whicli siirrouiids<br />

boih aiiicrior sciiicllar seiac. Wiiig as iii<br />

Fig. 25. ............ Carnpiglossu iiiic/riiii


224 iLlcr:. II.<br />

ZMHU); 3cr, 19. Arona, 13-26.11.1977 (Wolschrijn,<br />

ZMA); I O. Giiíiiiai 2.1V.1968 (v. Aariseii. ZMA).<br />

Goiiiera: I a, 5 hiii NNE Saii Scbastidii, 500 ni. 26.1.1981<br />

(Ellis, ZMA).<br />

Additiorral iriutc~rial. - Lectoiype Q (herc designated),<br />

2tunocerd avlkeri, 'w (T.V. Wollastoii<br />

1855-7) (NHM).<br />

Host plant. - Unknown, but tlicre are 1 u and I Q<br />

iii the NHM froni Madeira, bearing the labcl<br />

'Galactites toriteiifosa' and whicli were probably<br />

rcared froiii tliis plaiit. I have swept al1 my speci-<br />

iiiens from Galactiles tornen/osa aiid Curdtrrrs /J.w-<br />

rtocepliul,s (Conipositae).<br />

Distribution. - CANARY ISLANDS: Xeiierifc,<br />

Gomera, Gran Canaria (Frey 1936). GENERAL:<br />

Caiiary Islands and Madeira.<br />

Ecology. - On iincultivatcd groiiiid, iii iliistlc coiii-<br />

iiiuiiitics, from 0-1500 ni.<br />

Note. - I'licrc are 29 syiitypcs (labellcd as<br />

'('otypes') iii tiiccollcctioii of '1.V. Wollastoiiuiidcr<br />

tlic nanie 'walkeri' al tlie NHM. Onc Q has tlie<br />

Iiandwriiten Iñbel 'Tetanoccra ? walkcri W! This<br />

speciiiicn fits tlic concept of thc specics and is herc<br />

tlcsigiiaicd as lectotype. The otlicr 0 has no iden-<br />

tificatioii labcl aiid bcloiigs 10 Tnrpartw insrrlurum<br />

(Ikc.kcr).<br />

Canipiglossa duplex (Bccker, 1908a) coinb. n.<br />

7kphritis dirplex Ikcker. IWRa: 143. Holoiypc o,<br />

'Teiierifíe. 46799 1' (ZMHB) (examined).<br />

I'uroxyno aequalis Heririg, 1937: 253. Lecioiype cr (hcrc<br />

dcsigiialed), '1'. Orolava, 51180 V' (ZMHU) (csaiii-<br />

iiied); syn. n.<br />

Moieriol exoinineú. - 'lenerife: 58a. 31 0, El Mkdaiio.<br />

Moiiiañas de Anaga. Güíniar. Los Cristianos. Macizo de<br />

ho, Graiiadilla. El Kio. Uarraiicodel Kio. 4-15.111.1990;<br />

Jcr, 39, Saii Marcos, 26.111-2.1V.1968 (van Aariseii.<br />

TAlJ, ZMA); I u. 10, Güíniar, 2.1V.1968 (van Aartsen.<br />

TAIJ); 2 9, I'ucrlodclaCruz, 25.111-5.IV.196X(van Aari-<br />

seii, TAU); lo. Arona. 13-26.11.1977(Wolschrijn. ZMA).<br />

Ifosf planf. - Not reared. but swept at scveral sites<br />

on pure slands of Argyrantliernunt jru/escens<br />

(Coiiipocitae) togciher with C'u~~tpi~lossa retitu-<br />

IUIU.<br />

Distributiort. - CANARY ISLANDS: Riicrife.<br />

GENERAL: endeiiiic on Tcncrifc.<br />

&olug.r: - I n %)~Ro~i/i~~lluiii-%olli~ojeriu ;iiitl<br />

Kkiriiu I-irl>/iorl)iu coriiiliiiiiii ici lroiii 0-650 ni.<br />

iV.'ore.r. ~<br />

(1)<br />

Koriicycv (1989) pl;icctl iiiosi Iiasi<br />

EN l. SCANI>. VOL. 23:2 (1992)<br />

Palaearctic Puroxynaspp. in Carnpiglossa based on<br />

tlie iiiorpliology oi o gcnivalia. C duplex, C. mul-<br />

timaculosu and C. reticulofa share these characteres<br />

and are transferred to Carnpiglossa. (2) The holo-<br />

type of C duplex and tlie lectotype oí' C aequulis<br />

differ in wing pattern aiid size. The study of the<br />

speciiriciis listed above showed that these cliaracters<br />

are variable. The two iianies are, thercfore, syii-<br />

onyniized. (3) For differeiices froni C. iiiul-<br />

tirriucttlosu see note (2) under that species.<br />

Canipiglossa multimacuiosa<br />

(Diribek & Diribek, 1969) eomb. n.<br />

I'uroxynu reiirttluio ntirlliinoculoso IXrlbck s( I>irlbck<br />

1969: 9. Holotypc u, Goinera. near Tamargada, iioi located<br />

(not examiiied).<br />

/'oroxynamul/imocu/o.~u(I~irlhck & Dirlbck, 1969): üirlbck<br />

& Ohm (1982).<br />

Muleriul exoriiined. - Goinera: 17a. 109, 1.3s Kosas.<br />

13.111.1990; lo, 29. Taiiiargada. II.III.I9W, Iv. Los<br />

Aceviños. 700 in. 31.1.1981 (Ellis. ZMA).<br />

Addirirrviul rtioieriu/. - 20, I 9. Madeirn (\Voll:isioii,<br />

1855-7. NHM) (iiew lo Madeira).<br />

Has/ p/anr. - Reared from flower heads of Ar-<br />

gyrurithemitni ca/lic/rry.sirrir (Coiiiposiiac) (iiew<br />

Iios~ plaiit record).<br />

fli~~rili~tií~~i. - CANA RY ISI .A NDS: C;oiiicI~i.<br />

GENERAL: Goinera, Madcira.<br />

l2ology. - Only in tlie laiiropliyll forest fioiii<br />

300-500 ni.<br />

Notes. - (1) See note ' 1 ' under C. dirple-v. (2) C: mul-<br />

limoculoso and C duplex are very similar in exter-<br />

nal iiiorphology and differ oiily iii the tip of the<br />

aculcus (Figs 22 & 23), the colour of the sciitelliim,<br />

sliglitly in wing pattern (Figs 25 & 26) and iii thcir<br />

biology: C. inttl/iinuculosu attacks tlie flower heads<br />

of Argyrunrhemunt calliclirysttrn iii the laurophyll<br />

forcst of Gomera. C. duplex lives iii scnii-desert<br />

arcas inaiiily ín the S of Tenerií'e, where it probably<br />

at 1 ac ks A rgyran fhent u m frutescens. T hese mor-<br />

phological and biological differences are good cvi-<br />

dciicc iliat tlie two arc distiiict specics.<br />

Cartipiglossa reticulata<br />

(Becker, 1908a) comb. n.<br />

l2piiriiisreiiculoio kcker. lY08a: 143. Lecloiype a (Iiere<br />

debigiiaied). 'Teiieriffe, 46796 I ' (ZMHU) (exainiiirtl).<br />

li.phriris occw, 1x44: 407. l Ioloiy[ic Q , I'oriiigal<br />

(ZMHB) (exainiiied).<br />

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