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of the survey that produced this species, about 300 individual caves were visited, and only the three mentioned here were found to support milliped populations. This does not prove the millipeds are absent from these caves, but they were not found during the time available for exploration. Etymology: Named for Kyle Voyles, Arizona BLM Cave Coordinator, who organized and led the collecting effort that resulted in the type specimens. FIGURES 11–14. Gonopods of Pratherodesmus voylesi. Fig. 11. Gonopods in gonostome, ventral view. Fig. 12. Posterior view. Fig. 13. Lateral view. Fig. 14. Solenomere, distal zone, and subtending process, posterior view. Pratherodesmus ecclesia Shear, new species Figs. 4, 15–18, 34, 36. Types: Male holotype, two male, and four female paratypes from Cathedral Cave (part of Cathedral Cave Preserve), 10 miles south-southwest of Ash Fork, Yavapai Co., Arizona, collected 11 January 2007 by J. J. Wynne, deposited in FMNH; one male and one female paratype in CPMAB. 54 · Zootaxa 2151 © 2009 Magnolia Press SHEAR ET AL.
Description: Male. Length, 8.0 mm, width 0.75 mm. Nonsexual characters as described for P. voylesi, but metazonital setae are significantly longer and more acute (Fig 4). Pregonopodal legs slightly more crassate than postgonopodal legs. Gonopods (Figs. 15–18, 34, 35) large, gonostome walls bulging, prozonite of seventh segment notably swollen when seen dorsally. Gonocoxae hemispherical, filling gonostome, tightely appressed in midline, excavated to receive telopodites. Prefemora as in P. voylesi, but with lateral articulation less narrow, subtriangular; not obviously distally narrowed. Exomere absent, endomerite long, sinuous, appearing narrow in either anterior or posterior view, in lateral view, acute-triangular, with small basal tooth. Solenomere similar to P. voylesi, tibiotarsus reduced to absent, subtending process, large, lamellate. Female. Length, 8.0 mm, width 0.83 mm. Nonsexual characters as male, cyphopods as described for P. voylesi. FIGURES 15–18. Gonopods of Pratherodesmus ecclesia. Fig. 15. Gonopods in gonostome, ventral view. Fig. 16. Lateral view. Fig. 17. Anterior view. Fig. 18. Posterior view. NEW UNITED STATES CAVE MILLIPEDS Zootaxa 2151 © 2009 Magnolia Press · 55
Page 1 and 2: Zootaxa 2151: 47-65 (2009) www.mapr
Page 3 and 4: A recent paper by Djursvoll (2008)
Page 5 and 6: FIGURES 1, 2. Heads, collums and fi
Page 7: (200 ft.) within the cave to ~121 m
Page 11 and 12: seta of row widely separated, not a
Page 13 and 14: It now seems likely, based on size,
Page 15 and 16: Wheeler’s Deep Cave is one of fou
Page 17 and 18: FIGURES 37, 38. Fig. 37. Living exa
Page 19: Golovatch, S.I. (2009) Millipedes (

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