Taxonomical updates in The Scorpion Files for Scorpionidae - NTNU

Taxonomical updates in The Scorpion Files for Scorpionidae (2008 )
Taxa Status Distribution Comments Reference
Aops New genus Australia The family Urodacidae was
abolished by Soleglad, Fet &
Kovarik in 2005 and the subfamily
Urodacinae was transfered to
Scorpionidae. Volschenk & Prendini
have not accepted this decision and
retained the family Urodacidae (and
placed the new genus and species in
this family because of its close
relationship to Urodacus). The
taxonomy of The Scorpion Files
follows Soleglad & Fet (2005), and I
have here listed Aops in
Scorpionidae. This is formally not
correct, but is the best choice in the
current situation until a new revision
on higher scorpion systematics is
Cazierius scaber (Pocock,
1893)
Cazierius tatae Santiago-
Blay, 2009
published.
New comb. Jamaica New name: Heteronebo scaber
(Pocock, 1893)
Synonymization
Regarded as junior synonym of C.
garridoi Armas, 2005 and
synonymised.
2013 © Jan Ove Rein, The Scorpion Files
Volschenk ES, Prendini L. Aops
oncodactylus, gen. et sp. nov, the
frist troglobitic urodacid
(Urodacidae: Scorpiones), with a
re-assessment of cavernicolus,
troglobitic and troglomorphic
scorpions. Invertebrate
Systematics. 2008;22:235.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
de Armas LF. Acerca de algunos
alacranes (Scorpiones: Buthidae,
Scorpionidae) recientemente

Cryptoiclus Teruel &
Kovarik, 2012
Cryptoiclus rodriguezi Teruel
& Kovarik, 2012
Didymocentus armasi Teruel
& Rodriguez, 2008
Didymocentus jaumei Armas,
1976
New sp.
Puerto Rico
2013 © Jan Ove Rein, The Scorpion Files
descritos o registrados de Puerto
Rico e Isla Mona, Antillas
Mayores. Boletin Sociedad
Entomológica Aragonesa.
2009(45):298.
Santiago-Blay JA. Systematics
and some aspects of the biology
of the scorpions (Arachnida) of
the greater Puerto Rico region: A
biosystematic synopsis.
Entomological News. 2009 Jan-
Feb;120 (1):109-24.
New genus Cuba Endemic to eastern Cuba. Teruel R, Kovarik F. Scorpions
of Cuba: Jakub Rolcik - Clarion
Production; 2012 (page 168-174)
New sp. Cuba Endemic to eastern Cuba. Teruel R, Kovarik F. Scorpions
of Cuba: Jakub Rolcik - Clarion
Production; 2012 (page 168-174)
New sp. Cuba Teruel R, Rodriguez TM. La
subfamilia Diplocentrinae
(Scorpiones: Scorpionidae) en
Cuba. Quinta parte: el genero
Didymocentrus Kraepelin 1905.
Boletin Sociedad Entomológica
Restored
species
Species is valid and is redescribed
and updated as to their respective
diagnosis and geographical
distribution.
Aragonesa. 2008(42):53-78
Teruel R, Rodriguez TM. La
subfamilia Diplocentrinae
(Scorpiones: Scorpionidae) en
Cuba. Quinta parte: el genero
Didymocentrus Kraepelin 1905.

Didymocentus sanfelipensis Restored
Species is valid and is redescribed
Boletin Sociedad Entomológica
Aragonesa. 2008(42):53-78
Teruel R, Rodriguez TM. La
Armas, 1976
species
and updated as to their respective subfamilia Diplocentrinae
diagnosis and geographical (Scorpiones: Scorpionidae) en
distribution.
Cuba. Quinta parte: el genero
Didymocentrus Kraepelin 1905.
Boletin Sociedad Entomológica
Aragonesa. 2008(42):53-78
Diplocentrus bicolor New sp. Mexico Contreras-Felix G, Santibanez-
Contreras-Felix & Santibanez-
Lopez CE. Diplocentrus bicolor
Lopez, 2011
sp. n. (Scorpiones:
Diplocentridae) from Jalisco,
Mexico. Zootaxa.
2011(2992):61-8.
Diplocentrus formosus Armas New status Reinstated as valid species. Santibanez-Lopez CE, Francke
& Martin-Frias, 2003
Previously synonymized with OF, Prendini L. Systematics of
Diplocentrus tehuano Francke, the keyserlingii group of
1977.
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
Diplocentrus gladiator New sp. Mexico Species described in 1999, but I Beutelspacher CR, Trujillo
didn’t learn about this species before Olivera M. Una especie nueva de
October 2008. Thanks to Oscar Diplocentrus Peters
Francke for informing me!
(Scorpionida: Diplocentridae) de
Chiapas, Mexico. Rev Nica Ent.
1999;50:1-11.
Diplocentrus kraepelini New sp. Mexico Santibanez-Lopez CE, Francke
Santibanez-Lopez, Francke &
OF, Prendini L. Systematics of
2013 © Jan Ove Rein, The Scorpion Files

Prendini, 2013 the keyserlingii group of
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
Diplocentrus lachua Armas.
Trujillo & Agreda, 2011
Diplocentrus landelinoi
Trujillo & Armas, 2012
Diplocentrus longimanus
Santibanez-Lopez, Francke &
Athanasiadis, 2011
2013 © Jan Ove Rein, The Scorpion Files
2013 (3777):1-47.
New sp. Guatemala de Armas LF, Trujillo RE,
Agreda EO. Nueva especie de
Diplocentrus Peters, 1851
(Scorpionidae: Diplocentrinae)
del noroeste de alta Verapaz,
Guatemala. Boletin de la SEA.
2011(49):113-7.
New sp. Guatemala Trujillo RE, de Armas LF. Dos
especie nuevas de Diplocentrus
Peters, 1861 (Scorpionidae:
Diplocentrinae) de Guatemala.
Revista Iberica de Arachnologia.
2012 (21):131-8.
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Athanasiadis MC. The
genus Diplocentrus Peters
(Scorpiones: Diplocentridae) in
Morelos, Mexico. Revista
Iberica de Arachnologia.
Diplocentrus majahuensis New sp. Mexico Species described in 2003, but I
didn’t learn about this species before
October 2008. Thanks to Oscar
Francke for informing me!
2011;19:3-13.
Baldazo Monsivaiz JG.
Diplocentrus majahuensis,
nueva especie de alcran del
estado de Guerrero, Mexico
(Scorpiones, Diplocentridae).

Diplocentrus motagua Armas
& Trujillo, 2009
Diplocentrus oxlajujbaktun
Trujillo & Armas, 2012
Diplocentrus poncei Francke
& Quijano-Ravell, 2009
Diplocentrus sagittipalpus
Santibanez-Lopez, Francke &
Prendini, 2013
Diplocentrus sissomi
Santibanez-Lopez, Francke &
Prendini, 2013
New sp. Guatemala
Entomologia Mexicana.
2003;2:73-7.
de Armas LF, Trujillo RE.
Nueva especie de Diplocentrus
Peters, 1861 (Scorpiones:
Scorpionidae) de Guatemala.
Boletin Sociedad Entomológica
Aragonesa. 2009(45):67-72.
New sp. Guatemala Trujillo RE, de Armas LF. Dos
especie nuevas de Diplocentrus
Peters, 1861 (Scorpionidae:
Diplocentrinae) de Guatemala.
Revista Iberica de Arachnologia.
2012 (21):131-8.
New sp. Mexico Authors using family Diplocentridae Francke OF, Quijano-Ravell AF.
Una especie nueva Diplocentrus
(Scorpiones: Diplocentridae) del
estado de Michoacan, Mexico.
Revista Mexicana de
Biodiversidad 2009;80:659-63.
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Prendini L. Systematics of
the keyserlingii group of
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Prendini L. Systematics of
the keyserlingii group of
2013 © Jan Ove Rein, The Scorpion Files

Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
Diplocentrus tenango New sp. Mexico Santibanez-Lopez CE, Francke
OF. A new species of
Diplocentrus (Arachnida:
Scorpiones) from Oaxaca,
Mexico. Zootaxa.
Heterometrus atrascorpius
Mirza, Joshi, Desouza &
Sanap, 2012
Heterometrus liangi Zhu &
Yang, 2007
Heterometrus telanganaensis
Javed, Mirza, Tampal &
Lourenco, 2010
2013 © Jan Ove Rein, The Scorpion Files
2008(1742):53-60
New sp. India Mirza ZA, Joshi D, Desouza G,
Sanap RV. Description of a new
species of scorpion of the genus
Heterometrus Ehrenberg, 1828
(Scorpiones: Scorpionidae) from
the Western Ghats, India. Indian
Journal of Arachnology.
New sp. Vietnam Haven’t seen this paper (in Chinese).
Info based on bibliographical data
only.
2012;1(2):1-8.
Zhu MS, Yang X-F. Two species
of the genus Heterometrus
Ehrenberg, 1828 (Scorpionidae)
from South Vietnam sold in pet
shops in China. Acta
Arachnologica Sinica. 2007;16
(2):92-103.
New sp. India Maqsood Javed SM, Mirza ZA,
Tampal F, Lourenco WR. A new
species of the genus
Heterometrus Ehrenberg, 1828
(Scorpiones: Scorpionidae) from

Heteronebo franckei
Stockwell, 1985
Heteronebo jamaicae
jamaicae Francke, 1978
Heteronebo jamaicae
occidentalis Francke, 1978
Heteronebo jamaicae
portlandensis Francke, 1978
Heteronebo scaber (Pocock,
1893)
Synonymization Synonymized with Heteronebo
scaber (Pocock, 1893)
Synonymization Synonymized with Heteronebo
scaber (Pocock, 1893)
Synonymization Synonymized with Heteronebo
scaber (Pocock, 1893)
Synonymization Synonymized with Heteronebo
scaber (Pocock, 1893)
New comb. Jamaica Previous name: Cazierius scaber
(Pocock, 1893) + senior synonym of
Heteronebo franckei Stockwell,
1985, H. jamaicae jamaicae
Francke, 1978, H. jamaicae
2013 © Jan Ove Rein, The Scorpion Files
India with notes on its natural
history. Boletin de la SEA.
2010(47):143-8.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica

Pandinus awashensis
Kovarik, 2012
Pandinus gregoryi (Pocock,
1896)
Pandinus lowei Kovarik, 2012 New sp. Democratic
Republic of
Congo
Pandinus mazuchi Kovarik,
2011
Pandinus militaris Pocock,
1900
occidentalis Francke, 1978 & H.
jamaicae portlandensis Francke,
1978
2013 © Jan Ove Rein, The Scorpion Files
Aragonesa. 2009; (44):103-10.
New sp. Ethiopia Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
New status Considered a synonym of Pandinus
exitialis (Pocock, 1888) by Kovarik,
2003, but restored species status in
Kovarik’s (2009) Illustrated catalog
of scorpions (page 55).
Euscorpius. 2012(141):1-22.
Kovarik F. Illustrated catalog of
scorpions. Part I. Introductory
remarks; keys to families and
genera; subfamily Scorpioninae
with illustrated keys to
Heterometrus and Pandinus
species. Prague: Clarion
Production; 2009.
Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
Euscorpius. 2012(141):1-22.
New sp. Ethiopia Kovarik F. A review of the
subgenus Pandinus Thorell, 1876
with descriptions of two new
species from Uganda and
Ethiopia (Scorpiones:
Scorpionidae). Euscorpius.
Synonymization This taxa was judged as a junior
synonym of Pandinus cavimanus
2011(129):1-18.
Kovarik F. A checklist of
scorpions (Arachnida) in the

Pandinus somalilandus
Kovarik, 2012
Pandinus sundanicus Hirst,
1911
Pandinus ugandaensis
Kovarik, 2011
Oiclus nanus Teruel &
Chazal, 2010
(Pocock, 1888) in 2002, but I didn’t
discover this paper until July 2009.
2013 © Jan Ove Rein, The Scorpion Files
collection of the
Forschungsinstitut und
Naturmuseum Senckenberg,
Frankfurt am Main, Germany.
Serket. 2002 April;8 (1):1-23.
New sp. Somaliland Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
New status Sudan Previous status:
Pandinus exitialis sudanicus Hirst,
1911 (in part)
Pandinus magretti Birula, 1928 (in
part).
Euscorpius. 2012(141):1-22.
Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
Euscorpius. 2012(141):1-22.
New sp. Uganda Kovarik F. A review of the
subgenus Pandinus Thorell, 1876
with descriptions of two new
species from Uganda and
Ethiopia (Scorpiones:
Scorpionidae). Euscorpius.
New Sp. Guadeloupe,
Lesser Antilles
2011(129):1-18.
Teruel R, Chazal L. A new
species of the genus Oiclus
Simon, 1880 (Scorpiones:
Scorpionidae: Diplocentrinae)
from Guadeloupe, Lesser
Antilles. Euscorpius.
2010(92):1-9.
Oiclus questeli Teruel, 2008 New Sp. Lesser Antilles Teruel R. A new species of

Scorpio birulai Fet, 1997 New rank Morocco Previous subspecies status: Scorpio
maurus subtypicus Birula, 1910
Scorpio ennedi Lourenco,
Duhem & Cloudsley-
Thompson, 2012
Scorpio fuliginosus Pallary,
1928
2013 © Jan Ove Rein, The Scorpion Files
Oiclus Simon 1880 (Scorpiones:
Scorpionidae: Diplocentrinae)
from Saint-Barthelemy, Lesser
Antilles. Boletin Sociedad
Entomológica Aragonesa.
2008(43):95-9.
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
New sp. Chad Lourenco WR, Duhem B,
Cloudsley Thompson JL.
Scorpions from Ennedi, Kapka
and Tibesti, the mountains of
Chad, with descriptions of nine
new species (Scorpiones:
Buthidae, Scorpionidae).
Arthropoda Selecta.
New rank Morocco Previous status: Heterometrus
fuliginosus Pallary, 1928
2012;21(4):307-38.
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio hesperus Birula, 1910 New rank Morocco Previous subspecies status: Scorpio Lourenco WR. Reanalysis of the

Scorpio mogadorensis Birula,
1910
Scorpio niger Lourenco &
Cloudsley-Thompson, 2012
Scorpio occidentalis Werner,
1936
New rank Morocco Previous subspecies status: Scorpio
maurus mogadorensis Birula, 1910
maurus hesperus Birula, 1910 genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2013 © Jan Ove Rein, The Scorpion Files
2009;15(181):99-113.
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
New sp. Niger Lourenco WR, Cloudsley
Thompson JL. About the
enigmatic presence of the genus
Scorpio Linnaeus, 1758 in
Congo with the description of a
new species from Niger
(Scorpiones, Scorpionidae).
New rank Mali, Mauritania
& Senegal
Previous subspecies status: Scorpio
maurus occidentalis Werner, 1936
Serket. 2012;13(1/2):1-7.
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio punicus Fet, 2000 New rank Algeria, Libya, Previous subspecies status: Scorpio Lourenco WR. Reanalysis of the

Scorpio savanicola Lourenco,
2009
Scorpio sudanensis Lourenco
& Cludsley-Thompson, 2009
Scorpio weidholzi Werner,
1929
Urodacus butleri Volschenk,
Harvey & Prendini, 2012
Morocco &
Tunisia
maurus tunetanus Birula, 1910 genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2013 © Jan Ove Rein, The Scorpion Files
2009;15(181):99-113.
New sp. Cameroon Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
New sp. Sudan Lourenco WR, Cloudsley
Thompson JL. A new species of
the genus Scorpio Linnaeus 1758
from Sudan (Scorpiones,
Scorpionidae). Boletin Sociedad
Entomológica Aragonesa. 2009
New rank Morocco Previous subspecies status: Scorpio
maurus weidholzi Werner, 1929
(45):123-6.
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
New sp. Australia Volschenk ES, Harvey MS,
Prendini L. A new species of

2013 © Jan Ove Rein, The Scorpion Files
Urodacus (Scorpiones:
Urodacidae) from Western
Australia. American Museum
Novitates. 2012(3748):1-18.