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Early angiosperms from the Lower Cretaceous of Jixi, eastern ...

Early angiosperms from the Lower Cretaceous of Jixi, eastern ...

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Discussion: The present species is characterized by<br />

irregular venation mentioned above.<br />

Material: 2 specimens, including one questionable<br />

in identi¢cation.<br />

Locality and Horizon: Ibid., in <strong>the</strong> W-B section<br />

bed No.47.<br />

Genus Zhengia Sun et Dilcher gen.nov.<br />

Type: Zhengia chinensis Sun et Dilcher gen.et sp.<br />

nov.<br />

Generic diagnosis: Leaf or lea£et.Margin entire.<br />

Venation pinnate.Secondary veins branching dichotomously<br />

one or two times after going 1/2 distance<br />

to leaf margin.Tertiary and quaternary<br />

veins present.Marginal vein present.Leaves hypostomatic.Stomata<br />

anomocytic; guard cells<br />

sunken; subsidiary cells with collar £ange.<br />

Zhengia chinensis Sun et Dilcher gen. et sp. nov.<br />

Plate IV, 1^7<br />

Synonyms:<br />

1992a Shenkuoa caloneura Sun et Guo; Sun et al., p. 547<br />

pro parte, pl.I, ¢g.14; pl.II, ¢gs.2^6; (non pl.I,<br />

¢g.13; pl.II, ¢g.1).<br />

1996 Zhengia chinensis Sun et Dilcher, p.393, pl.1, ¢g.15;<br />

pl.2, ¢gs.7^9 [nomen nudum, nomen invalidum].<br />

Description: Leaf or lea£et more or less coriaceous,<br />

about 3.5 cm long by 2.4 cm wide; elliptic.<br />

Margin entire.Midvein nearly straight, about 0.5<br />

mm in thickness.Secondary veins arising <strong>from</strong><br />

midvein at angles <strong>of</strong> 60^70‡, and branching dichotomously<br />

one or two times after going 1/2 distance<br />

to <strong>the</strong> leaf margin.Tertiary and quaternary<br />

veins forming areoles and joining at marginal<br />

vein.Leaves hypostomatic.In upper cuticle, ordinary<br />

epidermal cells irregularly polygonal in form<br />

(average size ca.15U10 Wm), anticlinal walls<br />

nearly straight or slightly sinuous.In lower cu-<br />

Plate III.<br />

G. Sun, D.L. Dilcher / Review <strong>of</strong> Palaeobotany and Palynology 121 (2002) 91^112 103<br />

ticle, stomata elliptic to ovate, anomocytic (average<br />

size ca.30U25 Wm), randomly orientated.<br />

Guard cell sunken and subsidiary cells cutinized<br />

strongly at edges as a collar £ange.Ordinary epidermal<br />

cells irregularly polygonal in form (average<br />

size ca.25 U20 Wm), outer side <strong>of</strong> periclinal<br />

walls not smooth, but with numerous grooves.<br />

Holotype: SC10023, Plate IV, 1, 3^6.<br />

Etymology: Zhengia, named in honor <strong>of</strong> Pr<strong>of</strong>essor<br />

S.L. Zheng <strong>of</strong> <strong>the</strong> Shenyang Institute <strong>of</strong> Geology<br />

and Mineral Resources for his contributions in<br />

<strong>the</strong> collecting and research work on <strong>the</strong> <strong>Jixi</strong> early<br />

<strong>angiosperms</strong>; chinensis, country in which <strong>the</strong><br />

specimen was discovered.<br />

Discussion: The present species is characterized by<br />

coriaceous leaves, and anatomical features mentioned<br />

above.The present material is similar to<br />

Shenkuoa caloneura, but di¡ers in <strong>the</strong> secondary<br />

veins branching only one or two times, lacking<br />

exmedial looping near <strong>the</strong> leaf margin, and a presence<br />

<strong>of</strong> marginal veins.<br />

Material: 6 specimens.<br />

Locality and horizon: Ibid., in <strong>the</strong> W-B section<br />

bed No.47.<br />

Genus Xingxuephyllum Sun et Dilcher gen. nov.<br />

Type: Xingxuephyllum jixiense Sun et Dilcher gen.<br />

et sp.nov.<br />

Generic diagnosis: Leaf simple.Margin entire.<br />

Midvein prominent.Venation pinnate.Secondary<br />

veins alternate, branching 1/2 to 2/3 distance to<br />

leaf margin.Tertiary and smaller veins present.<br />

Intramarginal vein present.<br />

Xingxuephyllum jixiense Sun et Dilcher gen. et sp.<br />

nov.<br />

Plate V, 1B, 2; Plate III, ?11; Fig.4G<br />

Description: Leaf simple, about 3.5 cm long by<br />

2.0 cm wide (incomplete preservation). Margin<br />

1^3. Shenkuoa caloneura Sun et Guo emend.Sun et Dilcher.3.Showing <strong>the</strong> irregular venations.PB16777.1.Bar = 0.5 cm.<br />

2.Bar = 0.3 cm.3.Bar = 1 mm.<br />

4, 5. Reproductive organ A, SC10017.4.Bar = 0.33 cm.5.Bar = 1 cm.<br />

6, 7. Reproductive organ B, SC10018.6.Bar = 0.5 cm.7.Bar = 0.33 cm.<br />

8^10. Asiatifolium elegans Sun, Guo et Zheng emend.Sun et Dilcher, SC10019-10021.Bars = 0.5 cm.<br />

11. Shenkuoa caloneura Sun et Guo emend.Sun et Dilcher, SC10022.Bar = 0.5 cm.

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