Contribution to the knowledge of the Reptile fauna and diversity in ...

Citation as online-first paper: North-western Journal of Zoology 10: art.131519 

NORTH-WESTERN JOURNAL OF ZOOLOGY 10 (1): online-first ©NwjZ, Oradea, Romania, 2014 

Article No.: 131519 

http://biozoojournals.3x.ro/nwjz/index.html 

Contribution to the knowledge of the Reptile fauna 

and diversity in FYR of Macedonia 

Bogoljub STERIJOVSKI 1 , Ljiljana TOMOVIĆ 2,3, * and Rastko AJTIĆ 4 

1. Macedonian Ecological Society, Faculty of Natural Sciences, Blvd. "Kuzman Josifovski - Pitu" 28/3-7 1000 Skopje, FYR of Macedonia. 

2. Institute of Zoology, Faculty of Biology, University of Belgrade, Studentski trg 16, 11000 Belgrade, Serbia. 

3. Institute for Biological Research “Siniša Stanković”, University of Belgrade, Bulevar Despota Stefanaa 142, 11000 Belgrade, Serbia. 

4. Institute for Nature Conservation of Serbia, Dr Ivana Ribara 71, 11000 Belgrade, Serbia. 

*Corresponding author, L. Tomović, E-mail: lili@bio.bg.ac.rs 

Received: 22. April 2011 / Accepted: 20. September 2013 / Available online: 23. December 2013 / Printed: June 2014 

Abstract. Although reptile fauna of FYR of Macedonia (FYROM) was the matter of interest since more than a 

century, detailed distributions of most of the species are still missing. According to published data, 32 reptile 

species inhabit FYROM. In this paper, detailed distribution data of reptile species in FYROM are presented. 

Research of herpetofauna of FYROM was conducted from 1999 until 2012 and included 393 localities. For 32 

recorded reptile species, exact localities and altitudes, as well as published records are given. Analyses of the 

reptile diversity of the FYROM showed that the regions with the highest species diversity are: the Prespa and 

Ohrid Lake Region, Skopje region, Veles region and the Dojran Lake Region, with 18-21 reptile species per 

UTM 10 x 10 squares. These regions should be considered for the future designation of important 

herpetological areas at national level. Results of comparative analysis of reptile faunas from all Balkan 

countries showed that Macedonian fauna is the most similar with Albanian (92.537) and Bulgarian (90.909) 

reptile faunas. Zoogeographic analysis showed that reptile fauna of FYROM consists of eight chorotypes in 

total, the most dominant chorotype being the Eastern-Mediterranean with 13 species, followed by the 

Turano-Mediterranean with six and Southern-European with five species. Detailed distribution data of this 

study could be basis for future action plans, protection and conservation measures of reptiles in FYROM, and 

should be implemented in the national legislation and acts. In addition, this data are imperative for the Red 

Data Book of reptilians and designation of the Important Herpetological Areas in the country. 

Key words: reptile species, faunistic, biodiversity, zoogeography, FYR of Macedonia. 

Introduction 

Although more than 9,000 species of reptiles have 

been described worldwide (Uetz 2010), taxonomic, 

geographical and biological knowledge of this 

group is still incomplete, particularly in the species-rich 

countries of the developing world 

(Measey 2006). Habitat loss and degradation are 

the principal threats affecting reptile populations 

in these regions (Cox & Temple 2009), followed by 

unsustainable trade, invasive species, pollution, 

disease and climate change (Gibbons et al. 2000). 

Regional faunistic studies are the starting point for 

threatened reptile research, and reveal regions, 

taxa and anthropogenic threat processes on which 

future research and conservation efforts should be 

targeted, to effectively maintain the local reptile 

diversity as well as world’s reptile diversity. 

The Balkan Peninsula, as a part of the Mediterranean 

basin, is generally recognized as one of 

the hot-spots of European biodiversity (Médail & 

Quézel 1999, Džukić & Kalezić 2004). FYR of Macedonia 

(FYROM), situated in the central part of 

the Balkan Peninsula, is characterized by high biological 

diversity of many plant and animal groups 

(Anonymous 2010). Published data about distribution 

of reptiles in the FYROM are mainly from two 

periods of the 20 th century. The first one falls between 

the two world wars (Doflein 1921, Buresch 

& Zonkow 1932, 1934, Karaman 1928, 1931, 1937, 

1938-39, 1939) and the second one is between the 

years 1950 and 2000 (Radovanović 1951, Dimovski 

1959a,b, 1963, 1964, 1966a,b, 1971, 1981, Džukić & 

Grubač 1988, Petrušev et al. 1990, Petkovski et al. 

2000/2001). Only recently, the number of distribution 

records for Macedonian reptile fauna started 

to increase in the published literature (Sterijovski 

& Stamatoski 2002, Sterijovski et al. 2002, 2010a,b, 

Sterijovski 2005, Jelić et al., 2012). In some of the 

recent general literature of European herpetofauna 

(Gasc et al. 1997), the distribution of the reptile 

species in FYROM are given, but precise distribution 

data are still lacking. 

The aims of this paper were to: a) use both literature 

and field data, in order to complete up-todate 

knowledge about distribution of herpetofauna 

in FYROM, b) asses regions with the highest 

reptile diversity in the country, c) propose the zo- 

UNCORRECTED PROOF


B. Sterijovski et al. 

ogeographic classification of the reptile species in 

FYROM, d) analyze the similarity of Macedonian 

reptile fauna with the neighboring countries of the 

Balkan Peninsula. 

Materials and methods 

Two types of records became part of the study: (i) records 

previously published in the literature and (ii) unpublished 

(new) records. They are as such, distinguished in 

the grid maps. Faunistic records (both literature and new 

ones) were mapped in the 10 km x 10 km UTM (universal 

Transverse Mercator) geographic coordinate grid system. 

During fourteen consecutive years (from 1999 to 

2012) numerous field trips were conducted in different 

regions of FYROM. Most field trips were done in July and 

August, and some were carried out from April to September. 

Most specimens were collected, examined for 

identification, photographed and released in original 

habitats. For identification, field guides of Radovanović 

(1951), Arnold & Ovenden (2002) were used, and taxonomy 

and nomenclature was adopted from Speybroeck et 

al. (2010). Some samples (e. g. road kills) were collected 

for more detailed studies and were deposited at the Herpetological 

collection of Macedonian Ecological Society or 

the private collection of Bogoljub Sterijovski in Skopje. 

The majority of literature data are given with exact 

localities. However, in several old references given by 

Dimovski (1960, 1963, 1966b) and Karaman (1922, 1931), 

only general distribution of certain species is given (e.g. 

mountains or regions) without precise localities. Due to 

imprecision of those data, we did not include them in the 

distribution maps and further analyses. 

For analyses of reptile’s diversity and the designation 

of centres of herpetological diversity in FYROM, we used 

an application created in Visual Basic 6.1 in the program 

WinWord 2003 (Niketić, 1999), using the method by Walter 

& Straka (1970), at National Grid UTM Reference for 

FYROM 10 x 10 km. 

Similarities with all Balkan’s countries were calculated 

according to the Sorensen Similarity Index. Data on 

the presence of the taxa follow Gasc et al. (1997); for Bosnia 

& Herzegovina, Greece (excluding Crete and Eastern 

Aegean Islands) and Bulgaria with respect to recent publications 

(Jablonski et al. 2012, Valakos et al. 2008, Stojanov 

et al. 2011). 

For zoogeographic analyses, chorotypes were identified 

according to the classification of Vigna Taglianti et al. 

(1999). Chorotype structure of the reptile fauna of FY- 

ROM was compared with that of two neighbouring countries 

of the central part of the Balkan Peninsula: Serbia 

and Montenegro. 

Results 

UNCORRECTED PROOF 

According to relevant herpetological literature 

(Doflein 1921, Buresch & Zonkow 1932, 1934, 

Karaman 1928, 1931, 1937, 1938-39, 1939, Radovanović 

1951, Dimovski 1959a,b, 1963, 1964, 

1966a,b, 1971, 1981, Džukić & Grubač 1988, 

Petrušev et al. 1990, Petkovski et al. 2000/2001, 

Džukić et al. 2001, Sterijovski 2005), 32 reptile species 

are present in FYROM. Field research conducted 

during fourteen years, confirmed the presence 

of all 32 species in this territory. Approximately 

65% of the territory was covered (393 localities 

– 188 of 294 totals UTM 10 x 10 squares). 

Unfortunately, intensive searches were not completed 

in the all parts of FYROM equally, thus 

some parts are not covered well enough with the 

data. 

For every reptile species distribution in FY- 

ROM, field data (with region, locality, altitude and 

date of finding) are given in Appendix I, and literature 

data (with localities and references) are 

presented in Appendix II. Records of all species in 

FYROM are given in Figure 1(A-C) (National Grid 

UTM 10 x 10 km Reference). 

Results of this study and published records 

show that the most common representatives (recorded 

in more than 20% of the country – 60 or 

more of total 294 UTM squares) of Macedonian 

herpetofauna are: T. hermanni, A. fragilis, L. viridis, 

L. trilineata, P. erhardii, P. muralis, N. natrix, C. austriaca, 

D. caspius, Z. longissimus and V. ammodytes 

(Table 1). 

Species that were moderately distributed (recorded 

in 5 – 20% of the territory – 15-60 of total 

294 UTM squares) in our results and/or in literature, 

data are: T. graeca, A. kitaibellii, L. agilis, P. 

tauricus, T. vermicularis, N. tessellata, P. najadum, E. 

quatuorlineata, M. monspessulanus and V. berus (Table 

1). 

According to current data, extremely rare reptilian 

species (recorded in less then 5% of the 

country – up to 15 of total 294 UTM squares) in 

FYROM are: E. orbicularis, M. rivulata, C. kotschyi, 

P. apodus, A. nigropunctatus, Z. vivipara, E. jaculus, 

H. gemonensis, T. fallax, Z. situla and V. ursinii (Table 

1). 

Analyses of the reptile diversity of the FY- 

ROM showed that the regions with the highest 

species diversity are: the Prespa and Ohrid Lake 

Region (DL 83, DL 93), Skopje region (EM 25), Veles 

region (EM 61, EM 70) and the Dojran Lake 

Region (FL 46), with 18-21 reptile species per UTM 

10 x 10 squares (Figure 2). 

Sorensen’s similarity index was used for comparative 

analysis of reptile faunas from all the Balkan 

countries. Results showed that Macedonian


Reptile fauna and diversity of FYR of Macedonia 


Figure 1/A. Records of reptile species in FYROM. (National Grid UTM 10 x 10 km Reference). 

White circles - new records; grey circles - literature records; black circles - confirmed literature records.




Figure 1/B. Records of reptile species in FYROM. (National Grid UTM 10 x 10 km Reference). 

White circles - new records; grey circles - literature records; black circles - confirmed literature records.



Figure 1/C. Records of reptile species in FYROM. (National Grid UTM 10 x 10 km Reference). 

White circles - new records; grey circles - literature records; black circles - confirmed literature records. 

Table 1. Species list and number of UTM 10 x 10 km 

grids with their records (field and literature data) 

in FYR of Macedonia. 

Species 

No. of UTM records 

Emys orbicularis 11 

Mauremys rivulata 7 

Testudo graeca 71 

Testudo hermanni 126 

Anguis fragilis 128 

Pseudopus apodus 6 

Ablepharus kitaibelii 22 

Cyrtopodion kotschyi 9 

Algyroides nigropunctatus 8 

Lacerta agilis 21 

Lacerta trilineata 134 

Lacerta viridis 106 

Podarcis erhardii 136 

Podarcis muralis 143 

Podarcis tauricus 20 

Zootoca vivipara 9 

Typhlops vermicularis 21 

Coronella austriaca 109 

Natrix natrix 94 

Natrix tessellata 50 

Dolichophis caspius 118 

Hierophis gemonensis 4 

Zamenis longissimus 68 

Zamenis situla 14 

Elaphe quatuorlineata 31 

Platyceps najadum 33 

Malpolon monspessulanus 17 

Telescopus fallax 8 

Eryx jaculus 6 

Vipera ammodytes 165 

Vipera berus 26 

Vipera ursinii 13 

fauna is most similar with the Albanian (92.537) 

and Bulgarian (90.909) reptile faunas (Table 2). 

Zoogeographic analysis showed that reptile 

fauna of FYROM consists of eight chorotypes in 

total (Table 3, Figure 3). The most dominant chorotype 

is the Eastern-Mediterranean one with 13 

species, followed by the Turano-Mediterranean 

with 6 and Southern-European with 5 species. 

Two endemic Balkan species (Algyroides nigropunctatus 

and Hierophis gemonensis) occur in the country, 

but none of them is endemic for FYROM. 

Comparison with two neighbouring countries of 

the central Balkans (Serbia and Montenegro) 

showed that representation of the reptile chorotypes 

are generally very similar, with the exception 

of an extra chorotype in Montenegro (Mediterranean 

chorotype due to the presence of Hemidactylus 

turcicus). Comparatively, Macedonian reptile 

fauna is characterized by the highest number 

of Turano-Mediterranean chorotype, with the exclusive 

presence of Eryx jaculus. 

Discussion 

Since the first published data by Doflein in 1921 

and until present day, there has been no detailed 

comprehensive study about distribution of reptiles 

that covers the complete territory of the FYROM. 

Several papers published in the 20 th century pre- 

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Figure 2. Species diversity of Reptiles in FYROM at National Grid UTM 10 x 10 km Reference. 

Table 2. Sorensen similarity index of reptile fauna of the Balkans countries. 

SSI 

FYR of 

Macedonia 

Serbia Montenegro Bulgaria Albania Greece Bosnia Croatia 

FYR of Macedonia 100.000 82.143 81.250 90.909 92.537 75.610 78.689 77.612 

Serbia 82.143 100.000 64.286 79.310 74.576 62.162 64.151 64.407 

Montenegro 81.250 64.286 100.000 72.727 80.597 63.415 91.803 89.552 

Bulgaria 90.909 79.310 72.727 100.000 84.058 76.190 69.841 69.565 

Albania 92.537 74.576 80.597 84.058 100.000 77.647 78.125 77.143 

Greece 75.610 62.162 63.415 76.190 77.647 100.000 60.759 63.529 

Bosnia 78.689 64.151 91.803 69.841 78.125 60.759 100.000 90.625 

Croatia 77.612 64.407 89.552 69.565 77.143 63.529 90.625 100.000 

Table 3. Chorotype classification of reptiles in FYROM (NSp – number of species). 

Chorotype 

NSp Species 

Centralasiatic-Europeo-Mediterranean 1 Natrix natrix 

Turano-European 1 Natrix tessellata 

Turano-Europeo-Mediterranean 1 Emys orbicularis 

European 2 Anguis fragilis, Coronella austriaca 

Euro-Siberian 3 Lacerta agilis, Zootoca vivipara, Vipera berus 

Southern-European 5 Testudo hermanni, Lacerta viridis, Podarcis muralis, Zamenis longissimus, 

Vipera ursinii 

Turano-Mediterranean 6 Testudo graeca, Pseudopus apodus, Typhlops vermicularis, Eryx jaculus, 

Platyceps najadum, Telescopus fallax 

Eastern-Mediterranean 13 Mauremys rivulata, Cyrtopodion kotschyi, Algyroides nigropunctatus, 

Lacerta trilineata, Podarcis erhardii, Podarcis tauricus, Ablepharus kitaibelii, 

Malpolon monspessulanus, Dolichophis caspius, Elaphe quatuorlineata, 

Hierophis gemonensis, Zamenis situla, Vipera ammodytes 

sented the distribution of reptiles in particular (but 

limited to) parts of the country (Buresch & Zonkow 

1932, 1934, Karaman 1928, 1931, 1937, 1938- 

39, 1939, Radovanović 1951, Dimovski 1959a,b, 

1963, 1964, 1966a,b, 1971, 1981, Džukić & Grubač 

1988, Petrušev et al. 1990, Petkovski et al. 

2000/2001, Sterijovski & Stamatoski 2002, Sterijovski 

et al. 2002, Sterijovski 2005, Sterijovski et al. 

2010a,b) or for specific taxonomic groups (e. g. Vipers 

– Jelić et al. 2012). 

Recent records of some Mediterranean species 

in the southernmost part of Serbia (the Pčinja 

UNCORRECTED PROOF



Figure 3. Comparison of numbers of identified chorotypes of reptiles of FYROM and two neighbouring countries 

of the central Balkans (Serbia and Montenegro). M – Mediterranean, C-E-M – Centralasiatic-Europeo- 

Mediterranean, T-E – Turano-European, T-E-M – Turano-Europeo-Mediterranean, E – European, 

E-S – Euro-Siberian, S-E – Southern-European, T-M – Turano-Mediterranean, E-M – Eastern-Mediterranean. 

River valley and Prizren) – P. najadum (Crnobrnja- 

Isailović & Aleksić, 1999), C. kotschyi (Ajtić & Tomović, 

2001), T. graeca (Tomović et al., 2004) and E. 

quatuorlineata (Ristić et al., 2006) are further support 

to the assumption that lack of data for the 

northern and north eastern parts of FYROM could 

be the result of incomplete research. Few novel records 

of the (sub)species of international importance 

(Vipera ursinii graeca) in the southern parts of 

Albania, may enlarge the possibility that this 

taxon could be distributed in the southern parts of 

FYROM too (Korsós et al. 2008). Future studies 

should also focus on bordering areas with 

neighbouring countries (Greece, Bulgaria, Albania, 

Serbia) for the comparative studies of the species 

diversity (e. g. Asimakopoulos 1994, Ioannidis & 

Bousbouras 1997, Haxhiu 1998, Crnobrnja- 

Isailović & Aleksić 1999, Ajtić & Tomović 2001, 

Petrov & Beshkov 2001, Ristić et al. 2006, Petrov et 

al. 2006). 

More comprehensive faunistic studies may 

show that some of the species currently treated as 

“extremely rare” in FYROM (e.g. P. apodus and C. 

kotschyi) could be distributed more widely, having 

in mind their general distribution and ecological 

preferences. According to the results of faunistic 

studies, reptile species that we consider to have 

the highest conservation value and that should 

have the highest priority in assessment of the conservation 

status and legal protections at the national 

level in FYROM are the following: 

(i) species that are extremely rare – e. g. P. apodus, 

A. nigropunctatus, H. gemonensis 

(ii) species which are restricted to very small 

(isolated) areas and/or specific habitats – E. orbicularis, 

M. rivulata, Z. vivipara, E. jaculus, T. vermicularis, 

V. berus 

(iii) species considered as important at the international 

level (e.g. Habitats directive, Bern convention, 

IUCN Red List and CITES) – e. g. T. 

graeca, T. hermanni, E. quatuorlineata, V. ursinii. 

The analyses of species diversity performed in 

this study identified regions that should be considered 

in the future designation of the Important 

Herpetological Areas at the national level. Among 

them are the Prespa Lake and the Ohrid Lake regions. 

They are already included in the NP 

Galičica, which makes this National Park one of 

the most important areas for the reptile conservation 

plans in FYROM. An imperative for adequate 

management of protected areas is detailed knowledge 

of the area’s biodiversity. Herpetological 

studies on diversity, distribution, and ecology of 

the species in protected areas are amongst the pri- 

UNCORRECTED PROOF



orities at the European community level (Kati et 

al. 2007). Implicitly, protected areas should serve 

as a good “shelter” for the reptile species, otherwise 

subject to anthropogenic influences and potential 

extinction in non-protected regions. However, 

it has been recently proved that this may not 

be the true – substantial population decline of several 

snake species was recorded in protected areas 

worldwide (Reading et al. 2010). Long-term population 

studies are essential for the assessment of 

the conservation status of reptile species as well as 

environmental protection programs at the local or 

regional level (Gibbons et al. 2000). Only recently, 

the only long-term population study in FYROM 

was set at Golem Grad Island at the Prespa Lake 

(Sterijovski et al. 2011, Ajtić et al. 2013). 

Unfortunately, other regions with the highest 

reptile diversity are not included in the national 

legislation as protected areas. Further more, 

Skopje and Veles regions include some of the biggest 

cities in the country (Skopje and Veles). Reptile 

communities thus probably suffer from strong 

anthropogenic pressure as concerned to habitat 

destruction and human disturbance, which are 

designated as one of the most important factors affecting 

reptile population decline and reduction of 

species range (Gibbons et al. 2000). 

Zoogeographic analyses showed that chorotype 

reptile diversity of FYROM is very high – 32 

species belong to nine chorotypes, and the most 

dominant is the Eastern-Mediterranean, followed 

by the Turano-Mediterranean. Predominance of 

Eastern-Mediterranean chorotypes was proved for 

most of the Balkan countries (Jablonski et al. 2012). 

However, dominance of the Turano- 

Mediterranean chorotype is particularly important 

for FYROM, as it distinguishes the reptile fauna of 

this country from other countries of the western 

Balkans (this study – Table 3, and Jablonski et al. 

2012). FYROM also presents an area where two 

species of this chorotype (Testudo graeca and Eryx 

jaculus) have western and/or northern distribution 

limit in Europe (Gasc et al. 1997). 

High chorotype diversity, presence of marginal 

reptile populations and occurrence of the 

two endemic species in FYROM (Algyroides nigropunctatus 

and Hierophis gemonensis) suggest high 

biogeographic and refugial importance for reptile 

diversity in the Balkan Peninsula (Hewitt 1999, 

Džukić & Kalezić 2004). 

Detailed distribution data of this study that 

cover more than 60% of the territory of FYROM 

could be basis for future action plans, protection 

and conservation measures of reptile species and 

should be implemented in the national legislation 

and acts. In addition, this data are imperative for 

the Red Data Book of reptilians and the designation 

of the important herpetological areas in FY- 

ROM. 

Acknowledgements. We would like to thank Dr Marjan 

Niketić and Dr Gordana Tomović (Belgrade) for help with 

distribution maps and biodiversity analyses. Also we 

thank Metodija Velevski, Dr Slavčo Hristovski and Dr 

Ljupčo Melovski for field distribution data. L. Tomović 

was partly funded by the Ministry of Education, Sciences 

and Technological Development of Republic of Serbia, 

grant No. 173043. 

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+ Appendix I – Field data – 10 pages (pp.XXXXX) 

+ Appendix II – Literature data – 4 pages (pp.XXXXX) 

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Appendix I – Field data 

Emys orbicularis (Linnaeus, 1758) 

Ilinska Planina Mt.: Belčišta village, Belčiško Blato, 850, 25.07.2008; 

Kavadarci Region: Rosoman village, Mrzenska propast, 11.10.2005; 

Kočani Region: Dolni Podlog village, 05.10.2002; Lake Dojran 

Region: Dojran, 150, 27.03.2008; Lake Ohrid Region: Kališta village, 

750, 15.07.2006; St. Naum Monastery, 708, 20.06.2009; Struga, 

750,15.07.2006; Lake Prespa Region: Stenje village, 850, 23.07.2005; 

Skopje Region: Katlanovsko Blato, 340, 30.05.2009; Strumica Region: 

Gabrovo village, Dubrava, 210, 08.06.2008; Monospitovsko Blato, 

210, 23.05.2008 

Mauremys rivulata (Valenciennes, 1833) 

Gevgelija Region: River Kovanska Reka, 60 m, 25.05.2008; Skopje 

Region: Katlanovsko Blato, 340 m, 30.05.2009; Strumica Region: 

Monospitovsko Blato, 200 m, 09.06.2007. 

Testudo graeca Linnaeus, 1758 

Babuna Mt.: Trojaci village, 582, 16.08.2010; Berovo Region: 

Vladimirovo village, 870, 24.08.2009; Bigla Mt.: Smilevo village, 

1000, 14.05.2007; Zlatari village, 1047, 19.03.2010; Bilino Mt.: 

Podržikonj village, 1000, 18.03.2007; Bistra Mt.: Lazaropole village, 

1250, 18.05.2006; Straža pass, 1200, 17.05.2006; Bitola Region: 

Golemo Konjari village, 610, 28.04.2006; Lavci village, 960, 

29.04.2006; Bogdanci Region: Gjavato village, 100, 03.05.2005; Demir 

Hisar Region: Demir Hisar, 820, 23.07.2008; Vranešnica village, 

1100, 24.07.2008; Demir Kapija Region: Došnica village, 300, 

01.06.2009; Mala Javorica River, 140, 01.04.2007; Dešat Mt.: 

Žirovnica village, 1200, 28.07.2008; Galičica Mt.: Bolno village, 1000, 

12.03.2010; Gevgelija Region: Kovanska Reka River, 60, 25.05.2008; 

Smokvica village, 01.04.2007; Gradsko Region: Crna reka River, 

mouth to Vardar River, 80, 02.06.2001; Ilinska Planina Mt.: Belčišta 

village, Belčiško Blato, 850, 25.07.2008; Jablanica Mt.: Modrič 

village, 1080, 27.07.2008; Jakupica Mt.: Nežilovo village, 1000, 

17.07.1999; Karaorman Mt.: Botun village, 950, 29.08.2008; 

Kavadarci Region: Tumba hill, 597, 17.08.2010; Kičevo Region: 

Izvor village, 1000, 25.07.2008; Kitka Mt.: Gorno Količani village, 

800, 01.09.2006; Kočani Region: Obleševo village, 305, 09.10.2008; 

Kozjak Mt.: Malotino village, 16.06.2010; Kožuf Mt.: Lukar, 1047, 

17.07.2005; Kratovo Region: Kratovo, 712, 17.03.2007; Kuklica 

village, 700, 16.06.2011; Stracin village, 13.04.2007; Kumanovo 

Region: Pelince village, 526, 20.03.2007; Vaksince village, 600, 

30.05.2008; Lake Dojran Region: Crničani village, 180, 02.08.2002; 

Makedonski Brod Region: Slatina village, 720, 24.07.2008; 

Maleševski Planini Mts.: Lake Berovsko Ezero, 980, 23.08.2009; 

Ramna reka River, 1000, 06.08.2010; Mangovica Mt.: Gorno 

Djudjance village, 29.04.2012; Sudić village, 14.07.2011; Mariovo 

Region: Vitolište village, 1000, 20.09.2006; Osogovski Planini Mts.: 

Gorno Gradče vilage, 650, 22.07.2007; Ivova Glava, 1200, 30.08.2008; 

Popova Čuka hill, 597, 10.10.2008; Rusov Grob, 850, 11.10.2008; 

Sokolarci village, 550, 17.04.2009; Štalkovica village, 717, 14.08.2008; 

Zletovska reka River, 800, 05.06.2008; Ovče Pole Region: Mečkuevci 

village, 17.08.2005; Plačkovica Mt.: Zrnovci village, 590, 09.10.2008; 

Plakenska Planina Mt.: Streževo village, 787, 14.05.2007; Prilep 

Region: Brailovo village, 950, 14.03.2010; Dabnica village, 

17.05.2009; Skopje Region: Dervenska Klisura gorge, 570, 

30.03.2007; Divlje village, 800, 14.08.2007; Jabolci village, 700, 

15.04.2009; Kačanička klisura gorge, 300, 31.03.2007; Lepenec River, 

15.05.2001; Orman village, 370, 30.04.2006; Treska River gorge, 

Matka, 340, 12.09.2004; Skopska Crna Gora Mt.: Čučer village, 580, 

30.04.2006; Slaviško pole Region: Psača village, 540, 18.03.2007; 

Stogovo Mt.: Papradnik village, 700, 24.07.2006; Strumica Region: 

Novo Selo village, 250, 24.05.2008; Sveti Nikole Region: Dorfulija 

village, 12.08.2004; Šar Planina Mt.: Gajre village, 650, 20.04.2000; 

Novo Selo village, 1340, 20.07.2009; Rogačevo village, 900, 

30.07.2008; Vrutok village, 1000, 30.07.2008; Štip Region: Leskovica 

village, 400, 20.03.2006; Veles Region: Babuna River, 180, 27.05.2001; 

Babuna River, Gradište, 366, 19.03.2008; Bair hill, 380, 01.05.2001; 

Bekirlija village, 300, 10.06.2009; Bogomila village, 680, 16.09.1999; 

Crkvino village, 03.04.2005; Kišino village, 250, 02.05.2005; Kočilari 

village, 15.04.2005; Vlaina Planina Mt.: Judovi Livadi, 1000, 

06.08.2010. 

Testudo hermanni Gmelin, 1789 

Babuna Mt.: Trojaci village, 582, 25.07.2010; Berovo Region: Blatec 

village, 950, 24.08.2009; Vladimirovo village, 870, 24.08.2009; Bistra 

Mt.: Daboec, 1000, 27.08.2010; Bitola Region: Dobruševo village, 

595, 22.09.2006; Golemo Konjari village, 610, 28.04.2006; Lavci 

village, 960, 29.04.2006; Mogila village, 580, 29.04.2006; Topolčani 

village, 600, 28.04.2006; Bogdanci Region: Gjavato village, 100, 

03.05.2005; Delčevo Region: Grad village, 800, 17.09.2009; Demir 

Hisar Region: Demir Hisar, 820, 23.07.2008; Vranešnica village, 900, 

24.07.2008; Demir Kapija Region: Došnica village, 300, 01.06.2009; 

Golema Javorica River, 150, 10.03.2007; Koprišnica village, 500, 

29.06.2005; Galičica Mt.: Čokolsko, 730, 30.07.2006; Konjsko village, 

1000, 25.08.2007; Leskoec village, 980, 25.07.2005; Gevgelija Region: 

Gorničet village, 210, 26.05.2008; Kovanska Reka River, 60, 

25.05.2008; Gradsko Region: Ulanci village, 196, 14.09.2009; Ilinska 

Planina Mt.: Belčišta village, Belčiško Blato, 850, 25.07.2008; Belica 

village, 900, 01.09.2008; Gradišteno, 1018, 18.07.2008; Jablanica Mt.: 

Višni village, 14.07.2006; Jakupica Mt.: Kapina village, 800, 

08.09.2010; Nežilovo village, 1000, 16.09.1999; Karaorman Mt.: 

Botun village, 950, 29.08.2008; Pesočani village, 950, 27.08.2010; 

Kavadarci Region: Lake Tikveško Ezero, Kamenica River, 

26.04.2005; Ljubaš hill, 580, 19.08.2008; Rosoman village, 200, 

13.09.2009; Kičevo Region: Srbjani village, 870, 26.07.2008; Kočani 

Region: Obleševo village, 305, 09.10.2008; Kozjak Mt.: Bešište 

village, 900, 22.05.2007; Malotino village, 18.04.2009; Kožuf Mt.: 

Kumanova Glava, 1066, 11.07.2005; Sermenin village, 400, 

26.05.2008; Kratovo Region: Dimonce village, 530, 15.06.2011; 

Kratovo, 712, 17.03.2007; Krilatica village, 698, 11.06.2011; Kuklica 

village, 700, 16.06.2011; Stracin village, 592, 16.03.2007; Šopsko 

rudare village, 580, 16.06.2011; Talašmance village, 639, 11.06.2011; 

Tatićev Kamen hill, 1013, 10.08.2012; Kriva Palanka Region: Čuka, 

1320, 21.05.2011; Drenje village, 680, 13.06.2011; Ivankovci, 600, 

12.06.2011; Zlatkovci, 617, 11.06.2011; Kruševo Region: Crnilište 

village, 646, 26.07.2010; Kumanovo Region: Beljakovce village, 470, 

21.03.2007; Dragomance village, 501, 09.08.2012; Klečevce village, 

366, 14.06.2011; Kokošinje village, 760, 12.05.2012; Mlado 

Nagoričane village, 398, 21.03.2007; Orašac village, 500, 21.03.2007; 

Pčinja River, Bislimska Gorge, 07.04.2005; Pelince village, 526, 

20.03.2007; Vaksince village, 600, 30.05.2008; Lake Dojran Region: 

Crničani village, 180, 09.06.2009; Čaušli village, 720, 10.06.2006; 

Lake Prespa Region: Carev Dvor village, 860, 11.03.2010; Ezerani 

village, 860, 11.03.2010; Golem Grad Island, 850, 02.08.2004; 

Konjsko village, 950, 22.07.2005; Stenje village, 860, 17.04.2006; 

Makedonski Brod Region: Krapa village, 1046, 24.07.2010; 

Makedonski Brod, 700, 24.07.2008; Samokov village, 900, 24.07.2008; 

Slatina village, 720, 24.07.2008; Suvodol village, 600, 24.07.2008; 

Volče village, 1000, 04.06.2012; Maleševski Planini Mts.: Lake 

Berovsko Ezero, 980, 23.08.2009; Mangovica Mt.: Gorno Djudjance 

village, 29.04.2012; Mariovo Region: Manastir village, Rasinbegov 

most, 23.08.2005; Peštani village, 27.07.2008; Raec, 14.04.2012; 

Staravina village, 860, 02.06.2010; Vitolište village, 1000, 20.09.2006; 

Negotino Region: Eneševo village, 290, 16.08.2010; Pepelište village, 

193, 15.08.2010; Nidže Mt.: Skočivir village, 700, 18.07.2002; 

Ogražden Mt.: Bezgaštevo village, 630, 22.07.2000; Lake Čaušica 

Ezero, 575, 17.07.2000; Osogovski Planini Mts.: Buov Rid, 848, 

30.08.2008; Dobrevo village, 603, 13.08.2008; Golak hill, 400, 

10.10.2008; Kriva Jabuka, 850, 15.07.2008; Leski village, 910, 

21.07.2007; Moždivnjak village, 700, 18.03.2007; Orizari village, 406, 

11.10.2008; Popova Čuka hill, 597, 10.10.2008; Ramni Nivi, 830, 

15.05.2008; Rusov Grob, 850, 11.10.2008; Sinkovica, 1000, 22.07.2007; 

Sokolarci village, 550, 16.04.2009; Šlegovo village, 798, 16.05.2008; 

Štalkovica village, 717, 14.08.2008; Tursko Rudari village, 550, 

14.08.2008; Varovište village, 850, 24.08.2008; Zletovska reka River, 

800, 05.06.2008; Pelister Mt.: Capari village, 950, 12.04.2002; 

Plačkovica Mt.: Zrnovci village, 590, 09.10.2008; Plakenska Planina 

Mt.: Brežani village, 1080, 19.07.2008; Sloeštica village, 720, 

UNCORRECTED PROOF



20.07.2008; Streževo village, 787, 14.05.2007; Virovo village, 840, 

20.07.2008; Prilep Region: Brailovo village, 950, 14.03.2010; Čelopeci 

village, 593, 13.03.2010; Kanatlarci village, 16.05.2009; Krivogaštani 

village, 670, 14.03.2010; Lažani village, 630, 13.03.2010; Prilep, 

Markovi Kuli, 16.06.2012; Prisad village, Sivec mine, 246, 

17.06.2012; Rusjaci village, 620, 12.03.2010; Zrze village, 700, 

06.06.2012; Probištip Region: Kundino village, 750, 13.08.2008; 

Radoviš Region: Babino Platno, 547, 21.03.2006; Skopje Region: 

Blace village, 600, 22.03.2009; Brazda village, 450, 22.03.2009; 

Čaršija, 270, 14.04.2002; Dervenska Klisura gorge, 570, 30.03.2007; 

Divlje village, 800, 14.08.2007; Jabolci village, 700, 15.04.2009; 

Kačanička klisura gorge, 300, 31.03.2007; Kučkovo village, 600, 

14.09.2002; Lepenec River, 15.05.2001; Orman village, 450, 

26.04.2002; Sopište village, 710, 15.04.2009; Taorska Klisura gorge, 

300, 29.06.2007; Treska River gorge, Matka, 340, 12.09.2004; Skopska 

Crna Gora Mt.: Čučer village, 600, 22.03.2009; Slaviško pole Region: 

Ginovci village, 500, 20.04.2007; Luda Mogila hill, 673, 23.05.2006; 

Psača village, 540, 18.03.2007; Rankovce village, 490, 12.06.2011; 

Rankovce village, 480, 20.04.2007; Vetunica village, 625, 11.06.2011; 

Stogovo Mt.: Papradnik village, 700, 24.07.2006; Struga Region: 

Radožda village, 10.04.2010; Strumica Region: Novo Selo village, 

250, 24.05.2008; Sveti Nikole Region: Gorobinci village, 450, 

20.03.2006; Štip Region: Bogoslovec village, 755, 12.06.2009; 

Erdželia village, 260, 11.06.2009; Leskovica village, 400, 21.03.2006; 

Radanja village, 400, 21.03.2006; Sofilari village, 320, 21.03.2006; 

Valandovo Region: Sobri village, 170, 25.05.2008; Udovo village, 

150, 30.03.2012; Veles Region: Babuna River, 180, 27.05.2001; 

Babuna River, Gradište, 170, 20.09.2010; Bekirlija village, 200, 

02.05.2005; Bogomila village, 680, 16.09.1999; Dolno Vranovci 

village, 260, 16.09.1999; Kišino village, 400, 10.06.2009; Slp village, 

410, 20.04.2009; Vardar River, Asarlak, 180, 15.08.2001; Veles, 

Kameno Brdo, 420, 20.09.2010; Vinica Region: Vinica, Kale, 470, 

19.09.2010; Vlaina Planina Mt.: Judovi Livadi, 1100, 06.08.2010. 

Anguis fragilis Linnaeus, 1758 

Berovo Region: Blatec village, 950, 24.08.2009; Vladimirovo village, 

870, 24.08.2009; Bigla Mt.: Smilevo village, 1000, 14.05.2007; Zlatari 

village, 1047, 19.03.2010; Bilino Mt.: Podržikonj village, 1000, 

18.03.2007; Bistra Mt.: Galičnik village, 1200, 17.05.2006; Galičnik 

village, Sv. Petka church, 30.06.2012; Lazaropole village, 1400, 

18.05.2006; Mavrovi Anovi, 1600, 17.05.2006; Bitola Region: 

Dobruševo village, 595, 22.09.2006; Golemo Konjari village, 610, 

28.04.2006; Lavci village, 960, 29.04.2006; Mogila village, 580, 

29.04.2006; Topolčani village, 600, 28.04.2006; Čučer-Sandevo 

Region: Brest village, 1146, 09.08.2012; Delčevo Region: Crnik 

village, 983, 17.09.2009; Grad village, 800, 17.09.2009; Vetroviti Rid, 

753, 16.09.2009; Demir Hisar Region: Demir Hisar, 820, 23.07.2008; 

Dešat Mt.: Žirovnica village, 1200, 28.07.2008; Galičica Mt.: Bolno 

village, 1000, 12.03.2010; Dolno Studino, 1450, 25.08.2007; Leskoec 

village, 980, 26.07.2005; Lipona Livada, 1540, 28.07.2005; Pljuska, 

940, 25.07.2005; Preseka, 1550, 23.07.2006; Tomoros peak, 1620, 

25.07.2006; Volkolegalo, 1100, 27.07.2005; German Mt.: Vojnova 

češma, 900, 01.04.2007; Gradsko Region: Ulanci village, 180, 

14.09.2009; Ilinska Planina Mt.: Belica village, 940, 03.08.2010; 

Gradišteno, 1018, 18.07.2008; Sveti Ilija peak, 1544, 22.07.2008; 

Šoburki, 1400, 17.07.2008; Jablanica Mt.: Modrič village, 1080, 

27.07.2008; Jakupica Mt.: Garvan, 1100, 17.07.1999; Kapina village, 

800, 08.09.2010; Nežilovo village, 1000, 16.09.1999; Selište village, 

1200, 08.09.2010; Karaorman Mt.: Botun village, 950, 29.08.2008; 

Čardak, 1150, 27.08.2010; Popova Livada, 1714, 27.08.2010; 

Kavadarci Region: Rosoman village, 200, 13.09.2009; Kičevo Region: 

Izvor village, 1000, 25.07.2008; Srbjani village, 870, 26.07.2008; Kitka 

Mt.: Kitka hut, 1100, 01.09.2006; Kočani Region: Obleševo village, 

305, 09.10.2008; Korab Mt.: Strezimir, 1200, 29.07.2008; Kozjak Mt.: 

Bešište village, 900, 22.05.2007; Kratovo Region: Kovačevci, 650, 

16.06.2011; Kratovo, 712, 16.03.2007; Kuklica village, 700, 16.06.2011; 

Stracin village, 592, 16.03.2007; Talašmance village, 639, 11.06.2011; 


1320, 21.05.2011; Golema Crcorija village, 1120, 20.05.2011; Ogut 

village, 900, 20.05.2011; Tlminci village, 618, 12.06.2011; Trnovo 

village, 1100, 20.05.2011; Uzem village, 950, 13.06.2011; Zlatkovci, 

617, 11.06.2011; Kruševo Region: Crnilište village, 646, 26.07.2010; 

Kumanovo Region: Beljakovce village, 470, 21.03.2007; Dragomance 

village, 501, 09.08.2012; Orašac village, 500, 21.03.2007; Pelince 

village, 526, 20.03.2007; Lake Ohrid Region: Elšani village, 820, 

26.08.2007; Lake Prespa Region: Carev Dvor village, 860, 11.03.2010; 

Ezerani village, 860, 11.03.2010; Konjsko village, 950, 22.07.2005; 

Stenje village, 870, 24.07.2005; Makedonska Kamenica Region: 

Dramče village, 800, 31.03.2009; Makedonski Brod Region: Belica 

village, 900, 24.07.2008; Dolno Botušje village, 970, 03.06.2012; 

Kosovo village, 680, 04.06.2012; Makedonski Brod, 700, 24.07.2008; 

Samokov village, 900, 24.07.2008; Slatina village, 720, 24.07.2008; 

Suvodol village, 600, 24.07.2008; Volče village, 1000, 04.06.2012; 

Maleševski Planini Mts.: Klepalo hill, 1400, 22.08.2009; Lake 

Berovsko Ezero, 980, 23.08.2009; Ramna reka River, 1000, 

06.08.2010; Mariovo Region: Vitolište village, 1000, 20.09.2006; 

Negotino Region: Eneševo village, 290, 16.08.2010; Pepelište village, 

193, 15.08.2010; Nidže Mt.: Redir, 1580, 15.07.2002; Osogovski 

Planini Mts.: Bukovec, 1423, 15.05.2008; Buov Rid, 848, 30.08.2008; 

Džikovska Maala, 850, 01.08.2008; Golak hill, 400, 10.10.2008; Gorno 

Gradče vilage, 610, 21.04.2007; Jamište village, 906, 18.07.2008; 

Kapetanska Maala, 1141, 31.03.2009; Katanska Maala, 850, 


11.10.2008; Panagjur, 1669, 15.07.2007; Sokolarci village, 550, 

16.04.2009; Štalkovica village, 717, 14.08.2008; Toranica mine, 1068, 

19.03.2007; Tošino Boište, 1281, 24.08.2008; Vlaški Kolibi, 1633, 

24.08.2008; Pelister Mt.: Kunina Stena, 1600, 12.04.2002; Molika 

hotel, 1445, 12.07.2001; Plačkovica Mt.: Zrnovci village, 590, 

09.10.2008; Plakenska Planina Mt.: Babino village, 680, 19.07.2008; 

Streževo village, 787, 14.05.2007; Prilep Region: Brailovo village, 

950, 14.03.2010; Čelopeci village, 593, 13.03.2010; Krivogaštani 

village, 670, 14.03.2010; Lažani village, 630, 13.03.2010; Rusjaci 

village, 620, 12.03.2010; Zrze village, 700, 06.06.2012; Probištip 

Region: Kundino village, 750, 13.08.2008; Radoviš Region: Babino 

Platno, 547, 21.03.2006; Selečka Planina Mt.: Čanište village, 520, 

23.05.2007; Skopje Region: Blace village, 600, 22.03.2009; Brazda 

village, 450, 22.03.2009; Dervenska Klisura gorge, 570, 30.03.2007; 

Divlje village, 800, 14.08.2007; Jabolci village, 700, 15.04.2009; 

Orman village, 250, 09.05.2004; Sopište village, 710, 15.04.2009; 

Skopska Crna Gora Mt.: Biljak peak, 1250, 22.03.2009; Čučer village, 

600, 22.03.2009; Lipkovo village, 580, 22.03.2007; Pržar peak, 1200, 

22.03.2009; Slaviško pole Region: Ginovci village, 500, 20.04.2007; 

Psača village, 540, 18.03.2007; Rankovce village, 490, 12.06.2011; 

Rankovce village, 480, 20.04.2007; Vetunica village, 625, 11.06.2011; 

Stogovo Mt.: Bijak Dorok peak, 1900, 15.05.2007; Kodžadžik village, 

1200, 15.05.2007; Papradnik village, 700, 24.07.2006; Strumica 

Region: Gabrovo village, Šuntevica, 210, 27.05.2008; Suva Gora Mt.: 

Konjski Grob, 1030, 31.07.2010; Sveti Nikole Region: Gorobinci 

village, 450, 20.03.2006; Šar Planina Mt.: Gorno Jelovce village, 1216, 

17.05.2007; Lešnica, 1450, 16.07.2009; Novo Selo village, 1340, 

20.07.2009; Popova Šapka, 1550, 16.07.2009; Rogačevo village, 900, 

30.07.2008; Selce Keč, 1200, 20.07.2009; Torbeški Most, 1350, 

01.07.2006; Vrutok village, 1000, 30.07.2008; Štip Region: Bogoslovec 

village, 755, 12.06.2009; Erdželia village, 260, 11.06.2009; Leskovica 

village, 400, 21.03.2006; Radanja village, 400, 21.03.2006; Sofilari 

village, 320, 21.03.2006; Valandovo Region: Udovo village, 150, 

30.03.2012; Veles Region: Babuna River, Gradište, 170, 20.09.2010; 

Bogomila village, 680, 16.09.1999; Dolno Vranovci village, 260, 

16.09.1999; Kišino village, 400, 10.06.2009; Slp village, 410, 

20.04.2009; Veles, Kameno Brdo, 420, 20.09.2010; Vinica Region: 

Vinica, Kale, 470, 19.09.2010; Vlaina Planina Mt.: Judovi Livadi, 

1200, 06.08.2010. 

Pseudopus apodus (Pallas, 1775) 

Bogdanci Region: Gjavato village, 100, 03.05.2005; Demir Kapija 

Region: Klisura village, 18.04.2011; Gevgelija Region: Miravci 

village, Petruška Reka, češma Ilinden 1946, 01.04.2007; Lake Dojran 

Region: Crničani village, 180, 09.06.2009; Valandovo Region: Udovo 

village, 150, 30.03.2012. 


Ablepharus kitaibelii Bibron & Bory, 1833 

Bogdanci Region: Gjavato village, 100, 29.06.2008; Demir Kapija



Region: Došnica village, 300, 29.06.2005; Koprišnica village, 500, 

29.06.2005; Galičica Mt.: Lipona Livada, 1350, 28.07.2005; Prisoj, 950, 

26.07.2006; Suvo Pole, 1200, 28.07.2003; Tomoros peak, 1350, 

25.07.2006; Gevgelija Region: Gorničet village, 210, 26.05.2008; 

Kovanska Reka River, 60, 25.05.2008; Ilinska Planina Mt.: Lokvišta, 

950, 18.07.2008; Jakupica Mt.: Nežilovo village, 980, 17.07.1999; 

Kožuf Mt.: Sermenin village, 400, 26.05.2008; Lake Dojran Region: 

Crničani village, 180, 01.07.2000; Lake Prespa Region: Konjsko 

village, 950, 12.08.2010; Ogražden Mt.: Bezgaštevo village, 630, 

22.07.2000; Osogovski Planini Mts.: Tursko Rudari village, 550, 

28.08,2009; Zletovska reka River, 800, 01.07.2008; Pelister Mt.: Niže 

Pole village, Babin Zab, 15.08.2002; Plakenska Planina Mt.: Brežani 

village, 915, 18.07.2008; Skopje Region: Taorska Klisura gorge, 300, 

18.06.2006; Treska River gorge, Matka, 340, 12.09.2004. 

Cyrtopodion kotschyi (Steindachner, 1870) 

Gevgelija Region: Gjavato village, 100 m, 29.06.2008; Skopje Region: 

Volkovo village, 350 m, 29.09.2010. 

Algyroides nigropunctatus (Duméril & Bibron, 1839) 

Jablanica Mt.: Modrič village, 21.05.2006; Lake Ohrid Region: 

Trpejca village, 750, 29.08.2007; Skopje Region: Treska River gorge, 

Matka, 340, 12.09.2004; Treska River gorge, Sv. Nikola Šiševski 

Monastery, 07.08.2004. 

Lacerta agilis Linnaeus, 1758 

Bilino Mt.: Luke, 1400, 18.03.2007; Dešat Mt.: Golema Stena, 2000, 

05.07.2004; Lake Gorno Gaševo Ezero, 1850, 05.07.2004; Jablanica 

Mt.: Lake Vevčansko Ezero, 12.08.2005; Korab Mt.: Ciganski Prolaz, 

2330, 04.07.2004; Kabaš, 1955, 04.07.2004; Osogovski Planini Mts.: 

Juručki Grobišta, 1643, 30.08.2008; Markova Stupka, 1856, 

19.07.2007; Ruen peak, 2245, 17.07.2007; Sultan Tepe peak, 1900, 

17.07.2007; Vlaški Kolibi, 1633, 24.08.2008; Pelister Mt.: Bojadžiev 

Vrv peak, 1744, 18.07.2001; Čuka peak, 1650, 18.08.2002; Jorgov 

Kamen, 1744, 13.07.2001; Kopanki, 1650, 11.07.2001; Kunina Stena, 

1850, 12.04.2002; Lake Golemo Ezero, 2350, 19.07.2001; Markovo 

Bačilo, 1850, 17.08.2002; Mrazarnik, 1744, 19.07.2001; Muza, 2350, 

19.07.2001; Niže Pole village, 10.08.2002; Niže Pole village, Stjaj, 

12.08.2002; Niže Pole village, Babin Zab, 15.08.2002; Pelister peak, 

2400, 19.10.2002; Široko Stapalo peak, 2415, 19.10.2002; Šar Planina 

Mt.: Dedel Beg, 1900, 01.07.2006; Idrizova Dupka, 1950, 01.07.2006; 

Lera, 2100, 01.07.2006; Mountain hut "Ljuboten", 1700, 25.08.2007; 

Popova Šapka, 1700, 16.07.2009; Venec, 2000, 01.07.2006. 

Lacerta trilineata Bedriaga, 1886 

Berovo Region: Blatec village, 950, 24.08.2009; Berovo Region: 

Vladimirovo village, 870, 24.08.2009; Bigla Mt.: Smilevo village, 

1000, 14.05.2007; Bigla Mt.: Zlatari village, 1047, 19.03.2010; Bilino 

Mt.: Podržikonj village, 1000, 18.03.2007; Bistra Mt.: Galičnik 

village, 1200, 17.05.2006; Bitola Region: Dobruševo village, 595, 

22.09.2006; Bitola Region: Golemo Konjari village, 610, 28.04.2006; 

Bitola Region: Lavci village, 960, 29.04.2006; Bitola Region: Mogila 

village, 580, 29.04.2006; Bitola Region: Topolčani village, 600, 

28.04.2006; Bogdanci Region: Gjavato village, 100, 03.05.2005; 

Čučer-Sandevo Region: Brest village, 1146, 09.08.2012; Delčevo 

Region: Crnik village, 983, 17.09.2009; Delčevo Region: Grad 

village, 800, 17.09.2009; Delčevo Region: Vetroviti Rid, 753, 

16.09.2009; Demir Hisar Region: Demir Hisar, 820, 23.07.2008; 

Demir Kapija Region: Dren village, 21.05.2005; Demir Kapija 

Region: Mala Javorica River, 140, 01.04.2007; Dešat Mt.: Žirovnica 

village, 1200, 28.07.2008; Galičica Mt.: Bolno village, 1000, 

12.03.2010; Galičica Mt.: Konjsko village, 1100, 24.08.2007; Galičica 

Mt.: Suvo Pole, 1300, 28.07.2003; Galičica Mt.: Volkolegalo, 1100, 

27.07.2005; Galičica Mt.: Zli Dol, 965, 29.07.2006; German Mt.: 

Vojnova češma, 900, 17.03.2007; Gradsko Region: Nogaevci village, 


Planina Mt.: Belica village, 900, 03.08.2010; Ilinska Planina Mt.: 

Gradišteno, 1018, 18.07.2008; Jablanica Mt.: Modrič village, 1080, 

27.07.2008; Jakupica Mt.: Kapina village, 800, 08.09.2010; Jakupica 

Mt.: Selište village, 1200, 08.09.2010; Karaorman Mt.: Botun village, 

950, 29.08.2008; Kavadarci Region: Rosoman village, 200, 

13.09.2009; Kičevo Region: Izvor village, 1000, 25.07.2008; Kičevo 

Region: Srbjani village, 870, 26.07.2008; Kitka Mt.: Čestak, 846, 

01.09.2006; Kočani Region: Obleševo village, 305, 09.10.2008; Kozjak 

Mt.: Bešište village, 900, 22.05.2007; Kratovo Region: Dimonce 

village, 530, 15.06.2011; Kratovo Region: Kratovo, 712, 17.03.2007; 

Kratovo Region: Krilatica village, 698, 11.06.2011; Kratovo Region: 

Kuklica village, 700, 16.06.2011; Kratovo Region: Stracin village, 

592, 16.03.2007; Kratovo Region: Šopsko rudare village, 580, 

16.06.2011; Kratovo Region: Talašmance village, 639, 11.06.2011; 

Kratovo Region: Tatićev Kamen hill, 1013, 10.08.2012; Kriva 

Palanka Region: Čuka, 1320, 21.05.2011; Kriva Palanka Region: 

Drenje village, 680, 13.06.2011; Kriva Palanka Region: Golema 

Crcorija village, 1120, 20.05.2011; Kriva Palanka Region: Ivankovci, 

600, 12.06.2011; Kriva Palanka Region: Tlminci village, 618, 

12.06.2011; Kriva Palanka Region: Zlatkovci, 617, 11.06.2011; 

Kruševo Region: Crnilište village, 646, 26.07.2010; Kumanovo 

Region: Beljakovce village, 470, 21.03.2007; Kumanovo Region: 

Dragomance village, 501, 09.08.2012; Kumanovo Region: Klečevce 

village, 366, 14.06.2011; Kumanovo Region: Kokošinje village, 760, 

12.05.2012; Kumanovo Region: Orašac village, 500, 21.03.2007; 

Kumanovo Region: Pelince village, 526, 20.03.2007; Lake Dojran 

Region: Čaušli village, 720, 10.06.2006; Lake Ohrid Region: 

Ljubaništa village, 750, 25.08.2007; Lake Prespa Region: Carev Dvor 

village, 860, 11.03.2010; Lake Prespa Region: Ezerani village, 860, 

11.03.2010; Lake Prespa Region: Konjsko village, 950, 22.07.2005; 

Lake Prespa Region: Oteševo village, 910, 28.07.2006; Lake Prespa 

Region: Stenje village, 850, 10.08.2004; Makedonska Kamenica 

Region: Dramče village, 800, 31.03.2009; Makedonski Brod Region: 

Belica village, 940, 24.07.2008; Makedonski Brod Region: Dolno 

Botušje village, 970, 03.06.2012; Makedonski Brod Region: Kosovo 

village, 680, 04.06.2012; Makedonski Brod Region: Makedonski 

Brod, 700, 24.07.2008; Makedonski Brod Region: Samokov village, 

900, 24.07.2008; Makedonski Brod Region: Slatina village, 720, 

24.07.2008; Makedonski Brod Region: Suvodol village, 600, 

24.07.2008; Makedonski Brod Region: Volče village, 1000, 

04.06.2012; Maleševski Planini Mts.: Lake Berovsko Ezero, 980, 

23.08.2009; Mariovo Region: Brest, Agro "Mariovo", 16.04.2005; 

Mariovo Region: Manastir village, road to Zović village, 15.04.2012; 

Mariovo Region: Raec, 14.04.2012; Mariovo Region: Rapeš village, 

Crna River, Rapeški bridge, 29.08.2004; Mariovo Region: Vitolište 

village, 1000, 20.09.2006; Negotino Region: Eneševo village, 290, 

16.08.2010; Negotino Region: Pepelište village, 193, 15.08.2010; 

Osogovski Planini Mts.: Buov Rid, 848, 30.08.2008; Osogovski 

Planini Mts.: Dobrevo village, 603, 13.08.2008; Osogovski Planini 

Mts.: Golak hill, 330, 10.10.2008; Osogovski Planini Mts.: Gorno 

Gradče vilage, 610, 21.04.2007; Osogovski Planini Mts.: Jamište 

village, 906, 18.07.2008; Osogovski Planini Mts.: Kapetanska Maala, 

1141, 31.03.2009; Osogovski Planini Mts.: Katanska Maala, 850, 

31.03.2009; Osogovski Planini Mts.: Kunovo village, 1266, 

15.07.2008; Osogovski Planini Mts.: Leski village, 910, 21.07.2007; 

Osogovski Planini Mts.: Luti Rid, 1062, 01.08.2008; Osogovski 

Planini Mts.: Luto Brdo, 1332, 01.08.2008; Osogovski Planini Mts.: 

Moždivnjak village, 700, 18.03.2007; Osogovski Planini Mts.: Orizari 

village, 406, 11.10.2008; Osogovski Planini Mts.: Plavica, 1297, 

16.05.2008; Osogovski Planini Mts.: Popova Čuka hill, 597, 

10.10.2008; Osogovski Planini Mts.: Ramni Nivi, 830, 15.05.2008; 

Osogovski Planini Mts.: Sinkovica, 1300, 22.07.2007; Osogovski 

Planini Mts.: Sokolarci village, 550, 16.04.2009; Osogovski Planini 

Mts.: Straovica village, 1190, 01.08.2008; Osogovski Planini Mts.: 

Šlegovo village, 798, 16.05.2008; Osogovski Planini Mts.: Toranica 

mine, 1068, 19.03.2007; Osogovski Planini Mts.: Tursko Rudari 

village, 550, 14.08.2008; Osogovski Planini Mts.: Varovište village, 


Pelister Mt.: Niže Pole village, 10.08.2002; Pelister Mt.: Niže Pole 

village, Babin Zab, 15.08.2002; Pelister Mt.: Tutunica, 1150, 

10.08.2002; Plačkovica Mt.: Zrnovci village, 590, 09.10.2008; 

Plakenska Planina Mt.: Brežani village, 915, 18.07.2008; Plakenska 

Planina Mt.: Streževo village, 787, 14.05.2007; Plakenska Planina 

Mt.: Virovo village, 840, 20.07.2008; Prilep Region: Brailovo village, 

950, 14.03.2010; Prilep Region: Čelopeci village, 593, 13.03.2010; 

Prilep Region: Krivogaštani village, 670, 14.03.2010; Prilep Region: 

UNCORRECTED PROOF



Lažani village, 630, 13.03.2010; Prilep Region: Prilep, Markovi Kuli, 

16.06.2012; Prilep Region: Rusjaci village, 620, 12.03.2010; Prilep 

Region: Zrze village, 700, 06.06.2012; Probištip Region: Kundino 

village, 750, 13.08.2008; Radoviš Region: Babino Platno, 547, 

21.03.2006; Selečka Planina Mt.: Čanište village, 520, 23.05.2007; 

Skopje Region: Blace village, 600, 22.03.2009; Skopje Region: Brazda 

village, 450, 22.03.2009; Skopje Region: Čaršija, 260, 14.04.2002; 

Skopje Region: Dervenska Klisura gorge, 570, 30.03.2007; Skopje 

Region: Divlje village, 800, 14.08.2007; Skopje Region: Jabolci 

village, 700, 15.04.2009; Skopje Region: Kožle village, Prnar, 

10.06.2007; Skopje Region: Orman village, 210, 29.05.2008; Skopje 

Region: Sopište village, 710, 15.04.2009; Skopje Region: Treska River 

gorge, Matka, 650, 12.09.2004; Skopje Region: Volkovo village, 260, 

14.09.2002; Skopska Crna Gora Mt.: Čučer village, 600, 22.03.2009; 

Skopska Crna Gora Mt.: Lipkovo village, 580, 22.03.2007; Skopska 

Crna Gora Mt.: Pržar peak, 1200, 22.03.2009; Slaviško pole Region: 

Ginovci village, 500, 20.04.2007; Slaviško pole Region: Luda Mogila 

hill, 673, 23.05.2006; Slaviško pole Region: Psača village, 540, 

18.03.2007; Slaviško pole Region: Rankovce village, 490, 12.06.2011; 

Slaviško pole Region: Rankovce village, 480, 20.04.2007; Slaviško 

pole Region: Vetunica village, 625, 11.06.2011; Stogovo Mt.: 

Kodžadžik village, 1200, 15.05.2007; Stogovo Mt.: Papradnik 

village, 700, 24.07.2006; Struga Region: Radožda village, 10.04.2010; 

Strumica Region: Gabrovo village, Dubrava, 210, 11.06.2007; 

Strumica Region: Gabrovo village, Šuntevica, 210, 27.05.2008; 

Strumica Region: Monospitovsko Blato, 210, 19.06.2007; Strumica 

Region: Novo Selo village, Rakovec, 210, 14.06.2007; Suva Gora Mt.: 

Konjski Grob, 1030, 31.07.2010; Sveti Nikole Region: Gorobinci 

village, 450, 20.03.2006; Šar Planina Mt.: Rogačevo village, 900, 

30.07.2008; Šar Planina Mt.: Selce Keč, 1200, 20.07.2009; Šar Planina 

Mt.: Vrutok village, 1000, 30.07.2008; Štip Region: Bogoslovec 

village, 755, 12.06.2009; Štip Region: Erdželia village, 260, 

11.06.2009; Štip Region: Leskovica village, 400, 21.03.2006; Štip 

Region: Radanja village, 400, 21.03.2006; Štip Region: Sofilari 



Veles Region: Bogomila village, 680, 16.09.1999; Veles Region: 

Dolno Vranovci village, 260, 16.09.1999; Veles Region: Kišino 

village, 250, 02.05.2005; Veles Region: Kišino village, 400, 

10.06.2009; Veles Region: Kočilari village, 15.04.2005; Veles Region: 

Slp village, 410, 20.04.2009; Veles Region: Veles, Kameno Brdo, 420, 

20.09.2010; Vinica Region: Vinica, Kale, 470, 19.09.2010; Vlaina 

Planina Mt.: Judovi Livadi, 1200, 06.08.2010. 

Lacerta viridis (Laurenti, 1768) 

Azot Region: Babuna River, 400, 01.05.2001; Berovo Region: Blatec 

village, 950, 24.08.2009; Vladimirovo village, 870, 24.08.2009; Bistra 

Mt.: Galičnik village, 1200, 17.05.2006; Lazaropole village, 1400, 

18.05.2006; Bitola Region: Lavci village, 960, 29.04.2006; Čučer- 

Sandevo Region: Brest village, 1146, 09.08.2012; Delčevo Region: 

Crnik village, 983, 17.09.2009; Vetroviti Rid, 753, 16.09.2009; Demir 

Hisar Region: Demir Hisar, 820, 23.07.2008; Železnec village, 830, 

20.07.2008; Dešat Mt.: Trebiško Igralište, 1500, 05.07.2004; Žirovnica 

village, 1200, 28.07.2008; Galičica Mt.: Asan Gjura, 1500, 26.08.2007; 

Leskoec village, 980, 25.07.2005; Lipona Livada, 1540, 28.07.2005; 

Pljuska, 940, 25.07.2005; Volkolegalo, 1100, 27.07.2005; Ilinska 

Planina Mt.: Dolgi Rid, 1242, 18.07.2008; Šoburki, 1400, 17.07.2008; 

Jablanica Mt.: Drenok village, 21.05.2006; Modrič village, 1080, 

27.07.2008; Modrič village, 21.05.2006; Jakupica Mt.: Selište village, 


Kičevo Region: Srbjani village, 870, 26.07.2008; Kitka Mt.: Krasta, 

1300, 02.06.2002; Korab Mt.: Strezimir, 1200, 29.07.2008; Kozjak Mt.: 

Malotino village, 18.04.2009; Kožuf Mt.: Buka, 1400, 16.07.2004; 

Kratovo Region: Dimonce village, 530, 15.06.2011; Kovačevci, 650, 

16.06.2011; Kratovo, 712, 17.03.2007; Krilatica village, 698, 

11.06.2011; Kuklica village, 700, 16.06.2011; Stracin village, 592, 

16.03.2007; Šopsko rudare village, 580, 16.06.2011; Talašmance 

village, 639, 11.06.2011; Tatićev Kamen hill, 1013, 10.08.2012; Kriva 

Palanka Region: Čuka, 1320, 21.05.2011; Drenje village, 680, 

13.06.2011; Golema Crcorija village, 1120, 20.05.2011; Ivankovci, 

600, 12.06.2011; Ogut village, 900, 20.05.2011; Tlminci village, 618, 

12.06.2011; Trnovo village, 1100, 20.05.2011; Uzem village, 950, 

13.06.2011; Zlatkovci, 617, 11.06.2011; Kumanovo Region: 

Dragomance village, 501, 09.08.2012; Klečevce village, 366, 

14.06.2011; Kokošinje village, 760, 12.05.2012; Pčinja River, 

Bislimska Gorge, 08.08.2004; Lake Ohrid Region: Elšani village, 820, 

26.08.2007; Lake Prespa Region: Oteševo village, 910, 28.07.2006; 

Stenje village, 870, 23.07.2005; Makedonski Brod Region: Dolno 

Botušje village, 970, 03.06.2012; Kosovo village, 680, 04.06.2012; 

Makedonski Brod, 700, 24.07.2008; Volče village, 1000, 04.06.2012; 

Maleševski Planini Mts.: Klepalo hill, 1400, 22.08.2009; Lake 

Berovsko Ezero, 980, 23.08.2009; Ramna reka River, 1000, 

06.08.2010; Mariovo Region: Raec, 14.04.2012; Vitolište village, 1000, 

20.09.2006; Nidže Mt.: Kopanki, 1500, 19.07.2002; Mala Reka, 1000, 

21.07.2002; Ogražden Mt.: Bezgaštevo village, 844, 17.07.2000; 

Osogovski Planini Mts.: Bukovec, 1423, 15.05.2008; Čurulinska 

čuka, 1410, 14.07.2007; Džikovska Maala, 850, 01.08.2008; Golak hill, 

330, 10.10.2008; Golem Govedarnik, 1047, 30.08.2008; Goli Rid, 1216, 

16.05.2008; Gorno Gradče vilage, 610, 20.07.2007; Gorno Gradče 

vilage, 610, 21.04.2007; Istibanja village, Sokolovci, 750, 11.10.2008; 

Ivova Glava, 1318, 30.08.2008; Jamište village, 1062, 18.07.2008; 

Jastrebnik village, 950, 22.04.2007; Kapetanska Maala, 1141, 

31.03.2009; Katanska Maala, 850, 31.03.2009; Kunovo village, 1266, 

15.07.2008; Leski village, 910, 21.07.2007; Luto Brdo, 1332, 


11.10.2008; Panagjur, 1669, 15.07.2007; Pavlevi Nivi, 1366, 

15.07.2008; Plavica, 1297, 16.05.2008; Polaki village, 806, 06.03.2007; 

Ponikva, 1614, 12.07.2007; Ramni Nivi, 830, 15.05.2008; Sokolarci 

village, 550, 13.04.2008; Štalkovica village, 717, 14.08.2008; Toranica 

mine, 1068, 19.03.2007; Tošino Boište, 1281, 24.08.2008; Varovište 

village, 850, 24.08.2008; Vlaški Kolibi, 1633, 24.08.2008; Pelister Mt.: 

Brajčino village, 1100, 18.05.2002; Capari village, 950, 12.04.2002; 

Jorgov Kamen, 1744, 13.07.2001; Kopanki, 1600, 11.08.2001; 

Magarevo village, 1150, 14.07.2001; Niže Pole village, 1165, 

13.07.2001; Rotino village, 1100, 11.07.2001; Vardišta, 1060, 

11.07.2001; Plakenska Planina Mt.: Babino village, 680, 19.07.2008; 

Brežani village, 1080, 19.07.2008; Orlov Kamen, 1684, 17.07.2008; 

Streževo village, 787, 14.05.2007; Virovo village, 840, 20.07.2008; 

Prilep Region: Brailovo village, 950, 14.03.2010; Čelopeci village, 

593, 13.03.2010; Lažani village, 630, 13.03.2010; Rusjaci village, 620, 

12.03.2010; Zrze village, 700, 06.06.2012; Radoviš Region: Babino 

Platno, 547, 21.03.2006; Selečka Planina Mt.: Čanište village, 520, 


30.03.2007; Jabolci village, 700, 15.04.2009; Kučkovo village, 450, 

14.09.2002; Lepenec River, 15.05.2001; Sopište village, 710, 

15.04.2009; Treska River gorge, Matka, 650, 12.09.2004; Treska River 

gorge, near the lake, 15.05.2005; Skopska Crna Gora Mt.: Biljak 

peak, 1250, 22.03.2009; Pržar peak, 1200, 22.03.2009; Slaviško pole 

Region: Psača village, 540, 18.03.2007; Rankovce village, 490, 

12.06.2011; Vetunica village, 625, 11.06.2011; Stogovo Mt.: 

Kodžadžik village, 1200, 15.05.2007; Papradnik village, 700, 

24.07.2006; Strumica Region: Gabrovo village, Dubrava, 210, 

11.06.2007; Monospitovsko Blato, 210, 19.06.2007; Šar Planina Mt.: 

Dedel Beg, 1460, 01.07.2006; Gajre village, 650, 10.04.2003; Gorno 

Jelovce village, 1216, 17.05.2007; Lisec village, 1150, 10.04.2003; 

Novo Selo village, 1340, 20.07.2009; Selce Keč, 1200, 20.07.2009; 

Torbeški Most, 1350, 01.07.2006; Štip Region: Leskovica village, 400, 

21.03.2006; Radanja village, 400, 21.03.2006; Sofilari village, 320, 

21.03.2006; Valandovo Region: Udovo village, 150, 30.03.2012; Veles 

Region: Bekirlija village, 200, 02.05.2005; Kišino village, 400, 

10.06.2009; Veles, Kameno Brdo, 420, 20.09.2010; Vlaina Planina 

Mt.: Judovi Livadi, 1200, 06.08.2010. 

Podarcis erhardii (Bedriaga, 1882) 

Azot Region: Bistrica village, 06.04.2007; Berovo Region: Blatec 

village, 950, 24.08.2009; Vladimirovo village, 870, 24.08.2009; Bigla 

Mt.: Smilevo village, 1000, 14.05.2007; Zlatari village, 1047, 

19.03.2010; Bilino Mt.: Podržikonj village, 1000, 18.03.2007; Bistra 

Mt.: Galičnik village, 1200, 17.05.2006; Lazaropole village, 1400, 

18.05.2006; Bitola Region: Dobruševo village, 595, 22.09.2006; 

Golemo Konjari village, 610, 28.04.2006; Lavci village, 960, 

29.04.2006; Mogila village, 580, 29.04.2006; Topolčani village, 600, 

UNCORRECTED PROOF



28.04.2006; Čučer-Sandevo Region: Brest village, 1146, 09.08.2012; 

Delčevo Region: Crnik village, 983, 17.09.2009; Grad village, 800, 

17.09.2009; Vetroviti Rid, 753, 16.09.2009; Demir Hisar Region: 

Demir Hisar, 820, 23.07.2008; Dešat Mt.: Žirovnica village, 1200, 

28.07.2008; Galičica Mt.: Bolno village, 1000, 12.03.2010; Čokolsko, 

730, 30.07.2006; Konjsko village, 1100, 25.08.2007; Leskoec village, 

850, 10.08.2004; Pljuska, 940, 25.07.2005; Prisoj, 1000, 26.07.2006; 

Suvo Pole, 1200, 28.07.2003; Zli Dol, 965, 29.07.2006; Gradsko 

Region: Ulanci village, 180, 14.09.2009; Ilinska Planina Mt.: Belica 

village, 900, 03.08.2010; Jablanica Mt.: Modrič village, 1080, 

27.07.2008; Modrič village, 21.05.2006; Jakupica Mt.: Kapina village, 

800, 08.09.2010; Nežilovo village, 1000, 16.09.1999; Selište village, 


Kavadarci Region: Lake Tikveško Ezero, Kotursko, 280, 25.08.2004; 

Rosoman village, 200, 13.09.2009; Kičevo Region: Izvor village, 

1000, 25.07.2008; Srbjani village, 870, 26.07.2008; Kitka Mt.: Čestak, 

846, 01.09.2006; Kočani Region: Obleševo village, 305, 09.10.2008; 

Korab Mt.: Strezimir, 1200, 29.07.2008; Kozjak Mt.: Bešište village, 

900, 22.05.2007; Kožuf Mt.: Alčak, 1050, 10.07.2004; Kratovo Region: 

Dimonce village, 530, 15.06.2011; Kratovo Region: Kovačevci, 650, 

16.06.2011; Kratovo, 712, 17.03.2007; Krilatica village, 698, 

11.06.2011; Kuklica village, 700, 16.06.2011; Kuklica village, 700, 

16.06.2011; Stracin village, 592, 16.03.2007; Šopsko rudare village, 

580, 16.06.2011; Talašmance village, 639, 11.06.2011; Tatićev Kamen 

hill, 1013, 10.08.2012; Kriva Palanka Region: Čuka, 1320, 21.05.2011; 

Drenje village, 680, 13.06.2011; Golema Crcorija village, 1120, 

20.05.2011; Ivankovci, 600, 12.06.2011; Ogut village, 900, 20.05.2011; 

Tlminci village, 618, 12.06.2011; Trnovo village, 1100, 20.05.2011; 

Uzem village, 950, 13.06.2011; Zlatkovci, 617, 11.06.2011; Kruševo 

Region: Crnilište village, 646, 26.07.2010; Kumanovo Region: 

Beljakovce village, 470, 21.03.2007; Dragomance village, 501, 

09.08.2012; Klečevce village, 366, 14.06.2011; Kokošinje village, 760, 

12.05.2012; Orašac village, 500, 21.03.2007; Pelince village, 526, 

20.03.2007; Lake Ohrid Region: Kališta village, 750, 15.07.2006; 

Trpejca village, 730, 30.07.2006; Lake Prespa Region: Carev Dvor 

village, 860, 11.03.2010; Ezerani village, 860, 11.03.2010; Konjsko 

village, 950, 22.07.2005; Oteševo village, 910, 27.07.2006; Stenje 

village, 870, 23.07.2005; Makedonska Kamenica Region: Dramče 

village, 800, 31.03.2009; Makedonski Brod Region: Belica village, 

940, 24.07.2008; Makedonski Brod, 700, 24.07.2008; Samokov village, 

900, 24.07.2008; Slatina village, 720, 03.08.2010; Suvodol village, 600, 


23.08.2009; Mariovo Region: Manastir village, road to Zović village, 

15.04.2012; Raec, 14.04.2012; Rapeš village, Crna River, Rapeški 

bridge, 29.08.2004; Vitolište village, 1000, 20.09.2006; Negotino 

Region: Eneševo village, 290, 16.08.2010; Pepelište village, 193, 

15.08.2010; Nidže Mt.: Mala Reka, 900, 21.07.2002; Skočivir village, 

700, 18.07.2002; Osogovski Planini Mts.: Bobovski Rid, 750, 

24.08.2008; Buov Rid, 848, 30.08.2008; Dobrevo village, 603, 

13.08.2008; Golak hill, 330, 10.10.2008; Gorno Gradče vilage, 610, 

21.04.2007; Istibanja village, Sokolovci, 750, 11.10.2008; Jamište 

village, 906, 18.07.2008; Jastrebnik village, 950, 22.04.2007; Katanska 

Maala, 850, 31.03.2009; Kriva Jabuka, 850, 15.07.2008; Kunovo 

village, 1266, 15.07.2008; Leski village, 900, 21.07.2007; Moždivnjak 

village, 700, 18.03.2007; Orizari village, 406, 11.10.2008; Plavica, 

1297, 16.05.2008; Popova Čuka hill, 597, 10.10.2008; Rajčani village, 

770, 12.04.2008; Ramni Nivi, 830, 15.05.2008; Ratkova Skala, 

06.04.2005; Rusov Grob, 850, 11.10.2008; Sokolarci village, 550, 

16.04.2009; Šlegovo village, 798, 16.05.2008; Tursko Rudari village, 

550, 14.08.2008; Varovište village, 850, 24.08.2008; Plačkovica Mt.: 

Zrnovci village, 590, 09.10.2008; Plakenska Planina Mt.: Babino 

village, 680, 19.07.2008; Brežani village, 915, 18.07.2008; Sloeštica 

village, 720, 20.07.2008; Streževo village, 787, 14.05.2007; Virovo 

village, 840, 20.07.2008; Prilep Region: Brailovo village, 950, 

14.03.2010; Čelopeci village, 593, 13.03.2010; Krivogaštani village, 

670, 14.03.2010; Lažani village, 630, 13.03.2010; Mažučište village, 

Crveni Steni, under, 07.04.2007; Prilep, Markovi Kuli, 16.06.2012; 

Prisad village, Sivec mine, 246, 17.06.2012; Rusjaci village, 620, 

12.03.2010; Zrze village, 700, 06.06.2012; Probištip Region: Kundino 



Skopje Region: Blace village, 600, 22.03.2009; Brazda village, 450, 

22.03.2009; Čaršija, 260, 14.04.2002; Dervenska Klisura gorge, 570, 


15.04.2009; Lepenec River, 15.05.2001; Orman village, 210, 

29.05.2008; Sopište village, 710, 15.04.2009; Treska River gorge, 650, 

12.09.2004; Treska River gorge, Sv. Nikola Šiševski Monastery, 

02.12.2007; Volkovo village, 400, 23.05.2004; Skopska Crna Gora 

Mt.: Biljak peak, 1250, 22.03.2009; Čučer village, 600, 22.03.2009; 

Lipkovo village, 580, 22.03.2007; Pržar peak, 1200, 22.03.2009; 

Slaviško pole Region: Ginovci village, 500, 20.04.2007; Luda Mogila 

hill, 673, 23.05.2006; Psača village, 540, 18.03.2007; Rankovce village, 

490, 12.06.2011; Rankovce village, 480, 20.04.2007; Vetunica village, 

625, 11.06.2011; Stogovo Mt.: Kodžadžik village, 1200, 15.05.2007; 

Papradnik village, 700, 24.07.2006; Struga Region: Modrič village, 

Drimkovska gorge, Modrički brigde, 27.09.2003; Strumica Region: 

Monospitovo village, 210, 28.05.2008; Monospitovsko Blato, 210, 

19.06.2007; Suva Gora Mt.: Konjski Grob, 1030, 31.07.2010; Sveti 

Nikole Region: Gorobinci village, 450, 20.03.2006; Šar Planina Mt.: 

Gorno Jelovce village, 1216, 17.05.2007; Novo Selo village, 1340, 

20.07.2009; Rogačevo village, 900, 30.07.2008; Selce Keč, 1200, 

20.07.2009; Vrutok village, 1000, 30.07.2008; Štip Region: Bogoslovec 

village, 755, 12.06.2009; Erdželia village, 260, 11.06.2009; Leskovica 

village, 400, 21.03.2006; Radanja village, 400, 21.03.2006; Sofilari 






Vinica, Kale, 470, 19.09.2010. 

Podarcis muralis (Laurenti, 1768) 

Azot Region: Brezica River, near the pond, 410, 01.05.2001; Berovo 

Region: Blatec village, 950, 24.08.2009; Berovo Region: Vladimirovo 

village, 870, 24.08.2009; Bigla Mt.: Smilevo village, 1000, 14.05.2007; 

Bigla Mt.: Zlatari village, 1047, 19.03.2010; Bilino Mt.: Podržikonj 

village, 1000, 18.03.2007; Bistra Mt.: Galičnik village, 1200, 

17.05.2006; Bistra Mt.: Lazaropole village, 1400, 18.05.2006; Bitola 

Region: Dobruševo village, 595, 22.09.2006; Bitola Region: Golemo 

Konjari village, 610, 28.04.2006; Bitola Region: Lavci village, 960, 

29.04.2006; Bitola Region: Mogila village, 580, 29.04.2006; Bitola 

Region: Topolčani village, 600, 28.04.2006; Čučer-Sandevo Region: 

Brest village, 1146, 09.08.2012; Debar Region: Crn Drim River, 

mouth, 580, 27.09.2003; Delčevo Region: Crnik village, 983, 

17.09.2009; Delčevo Region: Grad village, 800, 17.09.2009; Delčevo 

Region: Vetroviti Rid, 753, 16.09.2009; Demir Hisar Region: Demir 

Hisar, 820, 23.07.2008; Demir Hisar Region: Železnec village, 830, 

20.07.2008; Dešat Mt.: Žirovnica village, 1200, 28.07.2008; Galičica 

Mt.: Asan Gjura, 1500, 26.08.2007; Galičica Mt.: Bolno village, 1000, 

12.03.2010; Galičica Mt.: Dolno Studino, 1450, 25.08.2007; Galičica 

Mt.: Džafa, 1500, 27.08.2007; Galičica Mt.: Konjsko village, 1100, 

24.08.2007; Galičica Mt.: Leskoec village, 980, 25.07.2005; Galičica 

Mt.: Lipona Livada, 1540, 28.07.2005; Galičica Mt.: Pljuska, 940, 

25.07.2005; Galičica Mt.: Prisoj, 1000, 26.07.2006; Galičica Mt.: Suvo 

Pole, 1450, 28.07.2003; Galičica Mt.: Tomoros peak, 1620, 25.07.2006; 

Galičica Mt.: Volkolegalo, 1100, 27.07.2005; German Mt.: Vojnova 

češma, 900, 17.03.2007; Gradsko Region: Ulanci village, 180, 

14.09.2009; Ilinska Planina Mt.: Belica village, 900, 03.08.2010; 

Ilinska Planina Mt.: Dolgi Rid, 1242, 18.07.2008; Ilinska Planina Mt.: 

Golemo Ilino village, 979, 13.07.2008; Ilinska Planina Mt.: Liska 

peak, 1750, 17.07.2008; Ilinska Planina Mt.: Sveti Ilija peak, 1544, 

17.07.2008; Jablanica Mt.: Modrič village, 1080, 27.07.2008; Jakupica 

Mt.: Kapina village, 800, 08.09.2010; Jakupica Mt.: Selište village, 


Karaorman Mt.: Čardak, 1150, 27.08.2010; Kavadarci Region: 

Rosoman village, 200, 13.09.2009; Kičevo Region: Izvor village, 

1000, 25.07.2008; Kičevo Region: Srbjani village, 870, 26.07.2008; 

Kitka Mt.: Kitka hut, 1100, 01.09.2006; Kočani Region: Obleševo 

village, 305, 09.10.2008; Korab Mt.: Strezimir, 1200, 29.07.2008; 

Kozjak Mt.: Bešište village, 900, 22.05.2007; Kožuf Mt.: Buka, 1400, 

16.07.2004; Kratovo Region: Dimonce village, 530, 15.06.2011; 

Kratovo Region: Kovačevci, 650, 16.06.2011; Kratovo Region: 

UNCORRECTED PROOF



Kratovo, 712, 17.03.2007; Kratovo Region: Krilatica village, 698, 

11.06.2011; Kratovo Region: Stracin village, 592, 16.03.2007; Kratovo 

Region: Šopsko rudare village, 580, 16.06.2011; Kratovo Region: 

Šopsko rudare village, 580, 16.06.2011; Kratovo Region: Talašmance 

village, 639, 11.06.2011; Kratovo Region: Tatićev Kamen hill, 1013, 

10.08.2012; Kriva Palanka Region: Čuka, 1320, 21.05.2011; Kriva 

Palanka Region: Drenje village, 680, 13.06.2011; Kriva Palanka 

Region: Golema Crcorija village, 1120, 20.05.2011; Kriva Palanka 

Region: Ivankovci, 600, 12.06.2011; Kriva Palanka Region: Ogut 

village, 900, 20.05.2011; Kriva Palanka Region: Tlminci village, 618, 

12.06.2011; Kriva Palanka Region: Trnovo village, 1100, 20.05.2011; 

Kriva Palanka Region: Uzem village, 950, 13.06.2011; Kriva Palanka 

Region: Zlatkovci, 617, 11.06.2011; Kruševo Region: Crnilište 


21.03.2007; Kumanovo Region: Dragomance village, 501, 09.08.2012; 

Kumanovo Region: Kokošinje village, 760, 12.05.2012; Kumanovo 

Region: Orašac village, 500, 21.03.2007; Kumanovo Region: Pelince 

village, 526, 20.03.2007; Lake Ohrid Region: Elšani village, 820, 

25.08.2007; Lake Ohrid Region: St. Naum Monastery, 760, 

25.08.2007; Lake Ohrid Region: Trpejca village, 730, 30.07.2006; 

Lake Prespa Region: Carev Dvor village, 860, 11.03.2010; Lake 

Prespa Region: Ezerani village, 860, 11.03.2010; Lake Prespa Region: 

Golem Grad Island, 850, 02.08.2004; Lake Prespa Region: Konjsko 

village, 950, 22.07.2005; Lake Prespa Region: Oteševo village, 910, 

27.07.2006; Lake Prespa Region: Stenje village, 850, 10.08.2004; 

Makedonska Kamenica Region: Dramče village, 800, 31.03.2009; 

Makedonski Brod Region: Belica village, 940, 24.07.2008; 

Makedonski Brod Region: Dolno Botušje village, 970, 03.06.2012; 

Makedonski Brod Region: Kosovo village, 680, 04.06.2012; 

Makedonski Brod Region: Makedonski Brod, 700, 24.07.2008; 

Makedonski Brod Region: Samokov village, 900, 24.07.2008; 

Makedonski Brod Region: Slatina village, 720, 24.07.2008; 

Makedonski Brod Region: Suvodol village, 600, 24.07.2008; 

Makedonski Brod Region: Volče village, 1000, 04.06.2012; 

Maleševski Planini Mts.: Klepalo hill, 1400, 22.08.2009; Maleševski 

Planini Mts.: Lake Berovsko Ezero, 980, 23.08.2009; Maleševski 

Planini Mts.: Ramna reka River, 1000, 06.08.2010; Mariovo Region: 

Vitolište village, 1000, 20.09.2006; Negotino Region: Eneševo 

village, 290, 16.08.2010; Negotino Region: Pepelište village, 193, 

15.08.2010; Nidže Mt.: Prava Reka, 1400, 11.07.2002; Nidže Mt.: 

Redir, 1600, 09.07.2002; Nidže Mt.: Vlaški Dol, 1400, 11.07.2002; 

Osogovski Planini Mts.: Bobovski Rid, 750, 24.08.2008; Osogovski 

Planini Mts.: Dlgidelsko Brdo, 1200, 06.03.2007; Osogovski Planini 

Mts.: Dobrevo village, 603, 13.08.2008; Osogovski Planini Mts.: 

Duračka Reka village, 13.05.2007; Osogovski Planini Mts.: 

Džikovska Maala, 850, 01.08.2008; Osogovski Planini Mts.: Golem 

Govedarnik, 1047, 30.08.2008; Osogovski Planini Mts.: Gorno 

Gradče vilage, 620, 20.07.2007; Osogovski Planini Mts.: Ivova 

Glava, 1318, 30.08.2008; Osogovski Planini Mts.: Jamište village, 

906, 18.07.2008; Osogovski Planini Mts.: Kapetanska Maala, 1141, 

31.03.2009; Osogovski Planini Mts.: Katanska Maala, 850, 

31.03.2009; Osogovski Planini Mts.: Kriva Jabuka, 850, 15.07.2008; 

Osogovski Planini Mts.: Kunovo village, 1266, 15.07.2008; 

Osogovski Planini Mts.: Lisje Brdo, 1569, 14.07.2007; Osogovski 

Planini Mts.: Luti Rid, 1062, 01.08.2008; Osogovski Planini Mts.: 

Luto Brdo, 1332, 01.08.2008; Osogovski Planini Mts.: Moždivnjak 

village, 700, 18.03.2007; Osogovski Planini Mts.: Orizari village, 406, 

11.10.2008; Osogovski Planini Mts.: Orlovo Krilo, 05.04.2005; 

Osogovski Planini Mts.: Pavlevi Nivi, 1366, 15.07.2008; Osogovski 

Planini Mts.: Plavica, 1297, 16.05.2008; Osogovski Planini Mts.: 

Ponikva, 1614, 13.07.2007; Osogovski Planini Mts.: Popova Čuka 

hill, 597, 10.10.2008; Osogovski Planini Mts.: Rajčani village, 770, 

12.04.2008; Osogovski Planini Mts.: Ramni Nivi, 830, 15.05.2008; 

Osogovski Planini Mts.: Rusov Grob, 850, 11.10.2008; Osogovski 

Planini Mts.: Sinkovica, 1300, 22.07.2007; Osogovski Planini Mts.: 

Sokolarci village, 550, 17.04.2009; Osogovski Planini Mts.: Straovica 

village, 1190, 01.08.2008; Osogovski Planini Mts.: Štalkovica village, 

717, 14.08.2008; Osogovski Planini Mts.: Toranica mine, 1068, 

19.03.2007; Osogovski Planini Mts.: Tošino Boište, 1281, 24.08.2008; 

Osogovski Planini Mts.: Varovište village, 850, 24.08.2008; 

Osogovski Planini Mts.: Vlaški Kolibi, 1633, 24.08.2008; Pelister Mt.: 

Babin Zab, 1600, 15.08.2002; Pelister Mt.: Capari village, 1000, 

12.04.2002; Pelister Mt.: Jorgov Kamen, 1744, 13.07.2001; Pelister 

Mt.: Kopanki, 1600, 11.07.2001; Pelister Mt.: Magarevo village, 1250, 

21.07.2001; Pelister Mt.: Molika hotel, 1450, 12.07.2001; Pelister Mt.: 

Niže Pole village, 1100, 10.08.2002; Pelister Mt.: Niže Pole village, 

Babin Zab, 15.08.2002; Pelister Mt.: Rotino village, 1150, 11.07.2001; 

Pelister Mt.: Rudina, 1545, 18.05.2002; Pelister Mt.: Sapundžica 

River, 1500, 18.08.2002; Pelister Mt.: St. Bogorojca church, 1545, 

18.05.2002; Pelister Mt.: Tutunica, 1150, 10.08.2002; Plačkovica Mt.: 

Zrnovci village, 590, 09.10.2008; Plakenska Planina Mt.: Babino 

village, 680, 19.07.2008; Plakenska Planina Mt.: Brežani village, 

1080, 19.07.2008; Plakenska Planina Mt.: Sloeštica village, 720, 

20.07.2008; Plakenska Planina Mt.: Streževo village, 787, 14.05.2007; 

Plakenska Planina Mt.: Virovo village, 840, 20.07.2008; Prilep 

Region: Brailovo village, 950, 14.03.2010; Prilep Region: Čelopeci 

village, 593, 13.03.2010; Prilep Region: Krivogaštani village, 670, 

14.03.2010; Prilep Region: Lažani village, 630, 13.03.2010; Prilep 

Region: Rusjaci village, 620, 12.03.2010; Prilep Region: Zrze village, 

700, 06.06.2012; Probištip Region: Kundino village, 750, 13.08.2008; 

Radoviš Region: Babino Platno, 547, 21.03.2006; Selečka Planina 

Mt.: Čanište village, 520, 23.05.2007; Skopje Region: Blace village, 

600, 22.03.2009; Skopje Region: Brazda village, 450, 22.03.2009; 

Skopje Region: Čaršija, 260, 07.09.2003; Skopje Region: Dervenska 

Klisura gorge, 570, 30.03.2007; Skopje Region: Divlje village, 800, 

14.08.2007; Skopje Region: Jabolci village, 700, 15.04.2009; Skopje 

Region: Orman village, 450, 26.04.2002; Skopje Region: Sopište 

village, 710, 15.04.2009; Skopje Region: Treska River gorge, Matka, 

650, 12.09.2004; Skopje Region: Volkovo village, 260, 14.09.2002; 

Skopska Crna Gora Mt.: Biljak peak, 1250, 22.03.2009; Skopska Crna 

Gora Mt.: Čučer village, 600, 22.03.2009; Skopska Crna Gora Mt.: 

Lipkovo village, 580, 22.03.2007; Skopska Crna Gora Mt.: Pržar 

peak, 1200, 22.03.2009; Slaviško pole Region: Ginovci village, 500, 

20.04.2007; Slaviško pole Region: Luda Mogila hill, 673, 23.05.2006; 

Slaviško pole Region: Psača village, 540, 18.03.2007; Slaviško pole 

Region: Rankovce village, 490, 12.06.2011; Slaviško pole Region: 

Rankovce village, 480, 20.04.2007; Slaviško pole Region: Vetunica 

village, 625, 11.06.2011; Stogovo Mt.: Kodžadžik village, 1200, 

15.05.2007; Stogovo Mt.: Papradnik village, 700, 24.07.2006; Struga 

Region: Radožda village, 10.04.2010; Suva Gora Mt.: Konjski Grob, 

1030, 31.07.2010; Sveti Nikole Region: Gorobinci village, 450, 

20.03.2006; Šar Planina Mt.: Dedel Beg, 1460, 01.07.2006; Šar Planina 

Mt.: Gorno Jelovce village, 1216, 17.05.2007; Šar Planina Mt.: 

Lešnica, 1450, 16.07.2009; Šar Planina Mt.: Lisec village, 1100, 

07.05.2000; Šar Planina Mt.: Novo Selo village, 1340, 20.07.2009; Šar 

Planina Mt.: Rogačevo village, 900, 30.07.2008; Šar Planina Mt.: 

Selce Keč, 1200, 20.07.2009; Šar Planina Mt.: Torbeški Most, 1350, 

01.07.2006; Šar Planina Mt.: Vrutok village, 1000, 30.07.2008; Štip 

Region: Bogoslovec village, 755, 12.06.2009; Štip Region: Erdželia 

village, 260, 11.06.2009; Štip Region: Leskovica village, 400, 

21.03.2006; Štip Region: Radanja village, 400, 21.03.2006; Štip 

Region: Sofilari village, 320, 21.03.2006; Veles Region: Babuna 

River, Gradište, 170, 20.09.2010; Veles Region: Bair hill, 380, 

01.05.2001; Veles Region: Bogomila village, 680, 16.09.1999; Veles 

Region: Dolno Vranovci village, 260, 16.09.1999; Veles Region: 

Kišino village, 400, 10.06.2009; Veles Region: Slp village, 410, 

20.04.2009; Veles Region: Veles, Kameno Brdo, 420, 20.09.2010; 

Vinica Region: Vinica, Kale, 470, 19.09.2010; Vlaina Planina Mt.: 

Judovi Livadi, 1200, 06.08.2010. 

Podarcis tauricus (Pallas, 1814) 

Galičica Mt.: Pljuska, 940, 25.07.2005; Prisoj, 1000, 26.07.2006; 

Kavadarci Region: Vitačevo, 790, 08.07.2005; Kitka Mt.: Batinci 


15.06.2011; Lake Prespa Region: Konjsko village, 950, 22.07.2005; 

Oteševo village, 910, 27.07.2006; Negotino Region: Gorni Disan 

village, 08.06.2007; Prilep Region: Nebregovo village, Sv. Jovan 

Monatery, 07.04.2007; Veles Region: Bair hill, 380, 01.05.2001. 


Zootoca vivipara von Jacquin, 1787 

Korab Mt.: Kobilino Pole, 2215, 04.07.2004; Osogovski Planini Mts.: 

Juručki Grobišta, 1643, 30.08.2008; Ruen peak, 2245, 17.07.2007;



Sultan Tepe peak, 1900, 18.07.2007; Vlaški Kolibi, 1634, 24.08.2008; 

Šar Planina Mt.: Idrizova Dupka, 2000, 01.07.2006; Lake 

Karanikolićko Ezero, 2000, 28.07.2008; Popova Šapka, 1900, 

05.09.2007; Vakaf, 1900, 18.07.2009. 

Typhlops vermicularis (Daudin, 1803) 

Bogdanci Region: Gjavato village, 100, 03.05.2005; Demir Kapija 

Region: Dren village, 300, 03.05.2005; Koprišnica village, 500, 

29.06.2005; Gradsko Region: Ulanci village, 196, 14.09.2009; 

Kavadarci Region: Ljubaš hill, 580, 25.07.2010; Lake Dojran Region: 

Crničani village, 180, 02.08.2002; Negotino Region: Eneševo village, 

298, 27.05.2008; Osogovski Planini Mts.: Orizari village, 500, 

07.07.2009; Sokolarci village, 550, 07.07.2009; Tursko Rudari village, 

550, 07.07.2009; Skopje Region: Kačanička klisura gorge, 300, 

17.03.2008; Treska River gorge, Matka, 340, 12.09.2004; Valandovo 

Region: Sobri village, 170, 25.05.2008; Veles Region: Babuna River, 

Gradište, 366, 19.03.2008; Bekirlija village, 300, 10.06.2009. 

Coronella austriaca Laurenti, 1768 

Berovo Region: Blatec village, 950, 24.08.2009; Vladimirovo village, 

870, 24.08.2009; Bigla Mt.: Smilevo village, 1000, 14.05.2007; Zlatari 


18.03.2007; Bistra Mt.: Galičnik village, 1200, 17.05.2006; Mavrovi 

Anovi, 1600, 17.05.2006; Bitola Region: Dobruševo village, 595, 

22.09.2006; Golemo Konjari village, 610, 28.04.2006; Lavci village, 

960, 29.04.2006; Mogila village, 580, 29.04.2006; Topolčani village, 

600, 28.04.2006; Delčevo Region: Grad village, 800, 17.09.2009; 

Vetroviti Rid, 753, 16.09.2009; Demir Hisar Region: Demir Hisar, 

820, 23.07.2008; Dešat Mt.: Žirovnica village, 1200, 28.07.2008; 

Galičica Mt.: Čokolsko, 730, 30.07.2006; Dolno Studino, 1450, 

25.08.2007; Džafa, 1550, 27.08.2007; Leskoec village, 980, 25.07.2005; 

Pljuska, 940, 25.07.2005; Prisoj, 950, 26.07.2006; Suvo Pole, 1300, 

28.07.2003; Tepeno, 2000, 23.07.2006; Tomoros peak, 1620, 

25.07.2006; Zli Dol, 965, 29.07.2006; Vojnova češma, 900, 17.03.2007; 

Gradsko Region: Ulanci village, 180, 14.09.2009; Ilinska Planina Mt.: 

Sveti Ilija peak, 1544, 12.07.2008; Jablanica Mt.: Modrič village, 1080, 

27.07.2008; Jakupica Mt.: Nežilovo village, 1000, 16.09.1999; Selište 

village, 1200, 08.09.2010; Karaorman Mt.: Botun village, 950, 

29.08.2008; Čardak, 1150, 27.08.2010; Popova Livada, 1714, 

27.08.2010; Kavadarci Region: Rosoman village, 200, 13.09.2009; 

Kičevo Region: Izvor village, 1000, 25.07.2008; Srbjani village, 870, 

26.07.2008; Kitka Mt.: Kitka hut, 1100, 01.09.2006; Kočani Region: 

Obleševo village, 305, 09.10.2008; Kožuf Mt.: Gradište, 1100, 

18.07.2005; Kratovo Region: Kratovo, 712, 17.03.2007; Krilatica 

village, 698, 11.06.2011; Kuklica village, 700, 16.06.2011; Stracin 


Palanka Region: Čuka, 1320, 21.05.2011; Tlminci village, 618, 

12.06.2011; Trnovo village, 1100, 20.05.2011; Uzem village, 950, 

13.06.2011; Zlatkovci, 617, 11.06.2011; Kruševo Region: Crnilište 

village, 646, 26.07.2010; Kumanovo Region: Dragomance village, 

501, 09.08.2012; Klečevce village, 366, 14.06.2011; Orašac village, 

500, 21.03.2007; Pelince village, 526, 20.03.2007; Lake Ohrid Region: 

Gradište auto camp, 780, 26.08.2007; Lake Prespa Region: Ezerani 

village, 860, 11.03.2010; Konjsko village, 950, 22.07.2005; Oteševo 

village, 910, 28.07.2006; Stenje village, 870, 24.07.2005; Makedonski 

Brod Region: Belica village, 900, 24.07.2008; Dolno Botušje village, 

970, 03.06.2012; Kosovo village, 680, 04.06.2012; Makedonski Brod, 

700, 24.07.2008; Samokov village, 900, 24.07.2008; Slatina village, 

720, 24.07.2008; Suvodol village, 600, 24.07.2008; Volče village, 1000, 


23.08.2009; Ramna reka River, 1000, 06.08.2010; Mariovo Region: 

Vitolište village, 1000, 20.09.2006; Nidže Mt.: Budimirci village, 

1360, 21.07.2002; Mala Reka, 1000, 21.07.2002; Redir, 1600, 

09.07.2002; Osogovski Planini Mts.: Džikovska Maala, 850, 

01.08.2008; Golak hill, 400, 10.10.2008; Goli Rid, 1216, 16.05.2008; 

Juručki Grobišta, 1643, 30.08.2008; Kapetanska Maala, 1141, 

31.03.2009; Katanska Maala, 850, 31.03.2009; Kriva Jabuka, 850, 

15.07.2008; Moždivnjak village, 700, 18.03.2007; Ponikva, 1614, 

13.07.2007; Ruen peak, 2245, 17.07.2007; Sokolarci village, 550, 

17.04.2009; Sultan Tepe peak, 1900, 17.07.2007; Šlegovo village, 798, 

16.05.2008; Štalkovica village, 717, 14.08.2008; Toranica mine, 1068, 

19.03.2007; Tošino Boište, 1281, 24.08.2008; Pelister Mt.: Dihovo 

village, Dihovski Rid, 1550, 16.07.2001; Niže Pole village, 1064, 

13.07.2001; Niže Pole village, Stjaj, 12.08.2002; Palisnopje, 1650, 


Plakenska Planina Mt.: Konjarnik, 1275, 19.07.2008; Streževo 

village, 787, 14.05.2007; Prilep Region: Brailovo village, 950, 

14.03.2010; Čelopeci village, 593, 13.03.2010; Krivogaštani village, 

670, 14.03.2010; Zrze village, 700, 06.06.2012; Selečka Planina Mt.: 

Čanište village, 520, 23.05.2007; Skopje Region: Blace village, 600, 

22.03.2009; Brazda village, 450, 22.03.2009; Divlje village, 800, 

14.08.2007; Jabolci village, 700, 15.04.2009; Kučkovo village, 500, 

14.09.2002; Orman village, 350, 26.04.2002; Sopište village, 710, 

15.04.2009; Skopska Crna Gora Mt.: Biljak peak, 1250, 22.03.2009; 

Čučer village, 600, 22.03.2009; Lipkovo village, 580, 22.03.2007; 

Pržar peak, 1300, 22.03.2009; Slaviško pole Region: Ginovci village, 

500, 20.04.2007; Luda Mogila hill, 673, 23.05.2006; Rankovce village, 


625, 11.06.2011; Stogovo Mt.: Papradnik village, 700, 24.07.2006; 

Strumica Region: Monospitovsko Blato, 210, 23.05.2008; Suva Gora 

Mt.: Konjski Grob, 1030, 31.07.2010; Sveti Nikole Region: Gorobinci 

village, 450, 20.03.2006; Šar Planina Mt.: Gorno Jelovce village, 1216, 

17.05.2007; Mountain hut "Ljuboten", 1650, 25.08.2007; Novo Selo 

village, 1340, 20.07.2009; Rogačevo village, 900, 30.07.2008; Selce 

Keč, 1200, 20.07.2009; Vrutok village, 1000, 30.07.2008; Štip Region: 

Bogoslovec village, 755, 12.06.2009; Erdželia village, 260, 11.06.2009; 

Leskovica village, 400, 21.03.2006; Veles Region: Babuna River, 

Gradište, 170, 20.09.2010; Bogomila village, 680, 16.09.1999; Dolno 

Vranovci village, 260, 16.09.1999; Kišino village, 400, 10.06.2009; Slp 

village, 410, 20.04.2009; Veles, Kameno Brdo, 420, 20.09.2010; Vlaina 


Natrix natrix (Linnaeus, 1758) 

Azot Region: Brezica River, near the pond, 410, 01.05.2001; Bilino 

Mt.: Podržikonj village, 1000, 18.03.2007; Bistra Mt.: Galičnik 

village, 1200, 17.05.2006; Lazaropole village, 1400, 18.05.2006; 

Mavrovi Anovi, 1600, 17.05.2006; Bitola Region: Golemo Konjari 

village, 610, 28.04.2006; Lavci village, 960, 29.04.2006; Mogila 

village, 580, 29.04.2006; Topolčani village, 600, 28.04.2006; Delčevo 

Region: Crnik village, 983, 17.09.2009; Grad village, 800, 17.09.2009; 


820, 23.07.2008; Dešat Mt.: Lake Dolno Gaševo Ezero, 1780, 

05.07.2004; Suva Bara, 2100, 05.07.2004; Žirovnica village, 1200, 

28.07.2008; Galičica Mt.: Leskoec village, 900, 10.08.2004; Suvo Pole, 

1300, 24.07.2006; Gradsko Region: Ulanci village, 180, 14.09.2009; 

Ilinska Planina Mt.: Belica village, 900, 03.08.2010; Golemo Ilino 

village, 979, 17.07.2008; Jablanica Mt.: Modrič village, 1080, 

27.07.2008; Jakupica Mt.: Kapina village, 800, 08.09.2010; Nežilovo 

village, 1000, 16.09.1999; Selište village, 1200, 08.09.2010; 

Karaorman Mt.: Botun village, 950, 29.08.2008; Čardak, 1150, 

27.08.2010; Kavadarci Region: Rosoman village, 200, 13.09.2009; 

Kičevo Region: Izvor village, 1000, 25.07.2008; Srbjani village, 870, 

26.07.2008; Kočani Region: Obleševo village, 305, 09.10.2008; Korab 

Mt.: Kobilino Pole, 1900, 04.07.2004; Strezimir, 1200, 29.07.2008; 

Kratovo Region: Kratovo, 712, 17.03.2007; Kuklica village, 700, 

16.06.2011; Stracin village, 592, 16.03.2007; Tatićev Kamen hill, 1013, 

10.08.2012; Kriva Palanka Region: Čuka, 1320, 21.05.2011; 

Ivankovci, 600, 12.06.2011; Ogut village, 900, 20.05.2011; Tlminci 

village, 618, 12.06.2011; Trnovo village, 1100, 20.05.2011; Uzem 

village, 950, 13.06.2011; Zlatkovci, 617, 11.06.2011; Kumanovo 

Region: Beljakovce village, 470, 14.06.2011; Dragomance village, 

501, 09.08.2012; Pelince village, 526, 20.03.2007; Lake Ohrid Region: 

Trpejca village, 730, 30.07.2006; Lake Prespa Region: Carev Dvor 

village, 860, 11.03.2010; Ezerani village, 860, 11.03.2010; Golem 

Grad Island, 870, 01.05.2009; Konjsko village, 950, 22.07.2005; 

Oteševo village, 910, 27.07.2006; Stenje village, 860, 23.07.2005; 

Makedonski Brod Region: Belica village, 940, 24.07.2008; Dolno 

Botušje village, 970, 03.06.2012; Kosovo village, 680, 04.06.2012; 

Samokov village, 900, 24.07.2008; Slatina village, 720, 24.07.2008; 

Suvodol village, 600, 24.07.2008; Volče village, 1000, 04.06.2012; 

Maleševski Planini Mts.: Lake Berovsko Ezero, 980, 23.08.2009; 

Ramna reka River, 1000, 06.08.2010; Mangovica Mt.: Sudić village, 

UNCORRECTED PROOF



14.07.2011; Negotino Region: Eneševo village, 290, 16.08.2010; 

Pepelište village, 193, 15.08.2010; Osogovski Planini Mts.: Bobovski 

Rid, 750, 24.08.2008; Bukovec, 1423, 15.05.2008; Goli Rid, 1216, 

16.05.2008; Gorno Gradče vilage, 650, 20.07.2007; Jamište village, 

906, 18.07.2008; Juručki Grobišta, 1643, 30.08.2008; Luti Rid, 1062, 


11.10.2008; Sokolarci village, 550, 16.04.2009; Štalkovica village, 717, 

14.08.2008; Vlaški Kolibi, 1633, 24.08.2008; Pelister Mt.: Crveni 

Steni, 1900, 13.07.2001; Jorgov Kamen, 1550, 13.07.2001; Rotino 

village, 1450, 15.07.2001; Plačkovica Mt.: Zrnovci village, 590, 


Prilep Region: Brailovo village, 950, 14.03.2010; Lažani village, 630, 


village, 750, 13.08.2008; Skopje Region: Brazda village, 450, 

22.03.2009; Čaršija, 270, 06.04.2003; Divlje village, 800, 14.08.2007; 

Jabolci village, 700, 15.04.2009; Lepenec River, 15.05.2001; Orman 

village, 450, 26.04.2002; Slaviško pole Region: Ginovci village, 500, 

20.04.2007; Psača village, 540, 18.03.2007; Rankovce village, 490, 

12.06.2011; Rankovce village, 480, 20.04.2007; Stogovo Mt.: 

Kodžadžik village, 1200, 15.05.2007; Papradnik village, 700, 

24.07.2006; Strumica Region: Gabrovo village, Dubrava, 210, 

11.06.2007; Gabrovo village, Pluštici, 210, 27.05.2008; 

Monospitovsko Blato, 210, 28.03.2008; Zubovo village, Matnik, 210, 

14.06.2007; Šar Planina Mt.: Dedel Beg, 1900, 01.07.2006; Lešnica, 

1450, 16.07.2009; Novo Selo village, 1340, 20.07.2009; Torbeški Most, 

1350, 01.07.2006; Vrutok village, 1000, 30.07.2008; Štip Region: 

Radanja village, 400, 21.03.2006; Veles Region: Babuna River, 

Gradište, 170, 20.09.2010; Bekirlija village, 200, 02.05.2005; Bogomila 

village, 680, 16.09.1999; Dolno Vranovci village, 260, 16.09.1999; 

Kišino village, 250, 02.05.2005; Slp village, 410, 20.04.2009; Veles, 

Kameno Brdo, 420, 20.09.2010. 

Natrix tessellata (Laurenti, 1768) 

Azot Region: Babuna River, 400, 01.05.2001; Bistra Mt.: Mavrovi 

Anovi, 1300, 17.05.2006; Bogdanci Region: Gjavato village, 100, 

03.05.2005; Čučer-Sandevo Region: Brest village, 1146, 09.08.2012; 

Debar Region: Globočica Lake, 620, 27.09.2003; Jakupica Mt.: 

Nežilovo village, 900, 12.07.1999; Kavadarci Region: Lake Tikveško 

Ezero, Kotursko, 280, 25.08.2004; Rosoman village, 200, 13.09.2009; 

Kratovo Region: Kuklica village, 700, 16.06.2011; Tatićev Kamen 

hill, 1013, 10.08.2012; Kriva Palanka Region: Čuka, 1320, 21.05.2011; 

Golema Crcorija village, 1120, 20.05.2011; Ivankovci, 600, 

12.06.2011; Ogut village, 900, 20.05.2011; Zlatkovci, 617, 11.06.2011; 

Kumanovo Region: Dragomance village, 501, 09.08.2012; Klečevce 

village, 366, 14.06.2011; Kokošinje village, 760, 12.05.2012; Pelince 

village, 526, 20.03.2007; Lake Ohrid Region: Kališta village, 750, 

14.07.2006; Trpejca village, 730, 30.07.2006; Lake Prespa Region: 

Asamati village, 850, 02.08.2004; Ezerani village, 860, 11.03.2010; 

Golem Grad Island, 850, 02.08.2004; Konjsko village, 950, 

22.07.2005; Oteševo village, 910, 27.07.2006; Stenje village, 860, 

23.07.2005; Makedonski Brod Region: Kosovo village, 680, 

04.06.2012; Samokov village, 900, 24.07.2008; Volče village, 1000, 


23.08.2009; Osogovski Planini Mts.: Gorno Gradče vilage, 610, 

15.07.2007; Moždivnjak village, 815, 13.06.2011; Pelister Mt.: 

Magarevo village, 1150, 21.07.2001; Plakenska Planina Mt.: Babino 

village, 680, 19.07.2008; Streževo village, 787, 14.05.2007; Prilep 

Region: Zrze village, 700, 06.06.2012; Probištip Region: Kundino 

village, 750, 13.08.2008; Skopje Region: Čaršija, 250, 07.09.2003; 

Orman village, 370, 30.04.2006; Skopska Crna Gora Mt.: Lipkovo 

village, 580, 22.03.2007; Slaviško pole Region: Rankovce village, 

490, 12.06.2011; Vetunica village, 625, 11.06.2011; Strumica Region: 

Gabrovo village, Dubrava, 210, 11.06.2007; Gabrovo village, 

Pluštici, 210, 27.05.2008; Šar Planina Mt.: Vrutok village, 1000, 


Bogomila village, 680, 16.09.1999; Slp village, 410, 20.04.2009; Veles, 

Kameno Brdo, 420, 20.09.2010. 


Dolichophis caspius (Gmelin, 1789) 

Azot Region: Vojnica village, 28.06.2006; Berovo Region: Blatec 



19.03.2010; Bilino Mt.: Podržikonj village, 1000, 18.03.2007; Bistra 

Mt.: Galičnik village, 1200, 17.05.2006; Bitola Region: Dobruševo 

village, 595, 22.09.2006; Golemo Konjari village, 610, 28.04.2006; 

Lavci village, 960, 29.04.2006; Mogila village, 580, 29.04.2006; 

Topolčani village, 600, 28.04.2006; Čučer-Sandevo Region: Brest 

village, 1146, 09.08.2012; Delčevo Region: Crnik village, 983, 

17.09.2009; Grad village, 800, 17.09.2009; Vetroviti Rid, 753, 

16.09.2009; Demir Hisar Region: Demir Hisar, 820, 23.07.2008; 

Železnec village, 830, 20.07.2008; Galičica Mt.: Bolno village, 1000, 

12.03.2010; Čokolsko, 730, 30.07.2006; Konjsko village, 1100, 

24.08.2007; Leskoec village, 950, 10.08.2004; Volkolegalo, 1100, 


Planina Mt.: Belica village, 900, 03.08.2010; Golemo Ilino village, 

979, 17.07.2008; Jablanica Mt.: Modrič village, 1080, 27.07.2008; 

Jakupica Mt.: Nežilovo village, 1000, 16.09.1999; Karaorman Mt.: 

Botun village, 950, 29.08.2008; Čardak, 1150, 27.08.2010; Kavadarci 

Region: Lake Tikveško Ezero, Kotursko, 280, 25.08.2004; Rosoman 

village, 200, 13.09.2009; Kičevo Region: Izvor village, 1000, 

25.07.2008; Srbjani village, 870, 26.07.2008; Kitka Mt.: Gorno 

Količani village, 800, 01.09.2006; Kočani Region: Obleševo village, 

305, 09.10.2008; Kozjak Mt.: Bešište village, 900, 22.05.2007; 

Malotino village, 30.04.2008; Kratovo Region: Kratovo, 712, 

17.03.2007; Kuklica village, 700, 16.06.2011; Stracin village, 592, 

16.03.2007; Šopsko rudare village, 580, 16.06.2011; Talašmance 


Palanka Region: Tlminci village, 618, 12.06.2011; Trnovo village, 



village, 501, 09.08.2012; Kokošinje village, 760, 12.05.2012; Orašac 

village, 500, 21.03.2007; Pčinja River, Bislimska Gorge, 08.08.2004; 

Pelince village, 526, 20.03.2007; Lake Dojran Region: Čaušli village, 

720, 10.06.2006; Lake Ohrid Region: Trpejca village, 800, 25.08.2007; 

Lake Prespa Region: Asamati village, 850, 02.08.2004; Carev Dvor 

village, 860, 11.03.2010; Ezerani village, 860, 11.03.2010; Konjsko 

village, 950, 22.07.2005; Oteševo village, 910, 27.07.2006; Stenje 

village, 850, 28.07.2003; Makedonska Kamenica Region: Dramče 

village, 800, 31.03.2009; Makedonski Brod Region: Belica village, 

940, 24.07.2008; Makedonski Brod, 700, 24.07.2008; Samokov village, 

900, 24.07.2008; Slatina village, 720, 24.07.2008; Suvodol village, 600, 

24.07.2008; Volče village, 1000, 04.06.2012; Maleševski Planini Mts.: 

Ramna reka River, 1000, 06.08.2010; Mariovo Region: Vitolište 

village, 1000, 20.09.2006; Negotino Region: Eneševo village, 290, 

16.08.2010; Pepelište village, 193, 15.08.2010; Nidže Mt.: Gradešnica 

village, 950, 17.06.2007; Skočivir village, 650, 18.07.2002; Osogovski 

Planini Mts.: Buov Rid, 848, 30.08.2008; Dobrevo village, 603, 

13.08.2008; Golak hill, 330, 10.10.2008; Golem Govedarnik, 1047, 

30.08.2008; Gorno Gradče vilage, 610, 21.04.2007; Jamište village, 

1062, 18.07.2008; Katanska Maala, 850, 31.03.2009; Kunovo village, 

1266, 15.07.2008; Leski village, 910, 21.07.2007; Luti Rid, 1062, 


11.10.2008; Rajčani village, 770, 12.04.2008; Sokolarci village, 550, 

16.04.2009; Straovica village, 1190, 01.08.2008; Šlegovo village, 798, 

16.05.2008; Tursko Rudari village, 550, 14.08.2008; Varovište village, 


Magarevo village, 1150, 19.10.2002; Niže Pole village, 1064, 


Plakenska Planina Mt.: Babino village, 680, 19.07.2008; Brežani 

village, 915, 18.07.2008; Streževo village, 787, 14.05.2007; Prilep 

Region: Brailovo village, 950, 14.03.2010; Čelopeci village, 593, 

13.03.2010; Desovo village, toward the lake, 28.08.2004; 

Krivogaštani village, 670, 14.03.2010; Lažani village, 630, 13.03.2010; 

Prisad village, Sivec mine, 246, 17.06.2012; Rusjaci village, 620, 




Skopje Region: Blace village, 600, 22.03.2009; Brazda village, 450, 

22.03.2009; Čaršija, 250, 14.04.2002; Dervenska Klisura gorge, 570, 


15.04.2009; Orman village, 250, 14.09.2002; Sopište village, 710, 

15.04.2009; Volkovo village, 350, 23.05.2004; Skopska Crna Gora



Mt.: Čučer village, 600, 22.03.2009; Lipkovo village, 580, 22.03.2007; 

Slaviško pole Region: Ginovci village, 500, 20.04.2007; Luda Mogila 

hill, 673, 23.05.2006; Psača village, 540, 18.03.2007; Rankovce village, 


625, 11.06.2011; Stogovo Mt.: Papradnik village, 700, 24.07.2006; 

Strumica Region: Novo Selo village, Rakovec, 210, 14.06.2007; Sveti 

Nikole Region: Gorobinci village, 450, 20.03.2006; Šar Planina Mt.: 

Gorno Jelovce village, 1216, 17.05.2007; Rogačevo village, 900, 

30.07.2008; Selce Keč, 1200, 20.07.2009; Vrutok village, 1000, 

30.07.2008; Štip Region: Bogoslovec village, 755, 12.06.2009; 

Erdželia village, 260, 11.06.2009; Leskovica village, 400, 21.03.2006; 

Radanja village, 400, 21.03.2006; Sofilari village, 320, 21.03.2006; 

Valandovo Region: Udovo village, 150, 30.03.2012; Veles Region: 

Babuna River, Gradište, 170, 20.09.2010; Bogomila village, 680, 

16.09.1999; Dolno Vranovci village, 260, 16.09.1999; Kišino village, 

400, 10.06.2009; Slp village, 410, 20.04.2009; Veles, Kameno Brdo, 

420, 20.09.2010; Vinica Region: Vinica, Kale, 470, 19.09.2010. 

Hierophis gemonensis (Laurenti, 1768) 

Kavadarci Region: Tumba hill, 597 m, 17.06.2010; Lake Prespa 

Region: Stenje village, 870 m, 24.07.2005; Prilep Region: Čepigovo 

village, 646 m, 05.06.2009. 

Zamenis longissimus (Laurenti, 1768) 

Berovo Region: Blatec village, 950, 24.08.2009; Bigla Mt.: Zlatari 


18.03.2007; Bistra Mt.: Galičnik village, 1200, 17.05.2006; Mavrovi 

Anovi, 1600, 17.05.2006; Demir Hisar Region: Demir Hisar, 820, 

23.07.2008; Demir Kapija Region: Studena Glava, 700, 31.05.2010; 

Dešat Mt.: Žirovnica village, 1200, 28.07.2008; Galičica Mt.: Bolno 

village, 1000, 12.03.2010; Dolno Studino, 1450, 25.08.2007; Golem 

Rid, 1079, 18.04.2006; Leskoec village, 980, 26.07.2005; Lipona 

Livada, 1540, 28.07.2005; Pljuska, 940, 25.07.2005; Prisoj, 950, 

26.07.2006; Suvo Pole, 1550, 24.07.2006; Volkolegalo, 1100, 

27.07.2005; Zli Dol, 965, 29.07.2006; German Mt.: Vojnova češma, 

900, 17.03.2007; Ilinska Planina Mt.: Sveti Ilija peak, 1544, 

12.07.2008; Jablanica Mt.: Modrič village, 1080, 27.07.2008; Jakupica 

Mt.: Kapina village, 800, 08.09.2010; Nežilovo village, 1000, 

16.09.1999; Karaorman Mt.: Botun village, 950, 29.08.2008; Popova 

Livada, 1680, 27.08.2010; Kičevo Region: Izvor village, 1000, 

25.07.2008; Kitka Mt.: Kitka hut, 1100, 01.09.2006; Korab Mt.: 

Strezimir, 1200, 29.07.2008; Kožuf Mt.: Konopište village, Žensko 

stream, 12.07.2005; Kratovo Region: Dimonce village, 530, 

15.06.2011; Stracin village, 592, 16.03.2007; Kriva Palanka Region: 

Golema Crcorija village, 1120, 20.05.2011; Uzem village, 950, 

13.06.2011; Kumanovo Region: Pelince village, 526, 20.03.2007; Lake 

Ohrid Region: St. Naum Monastery, 760, 25.08.2007; Lake Prespa 

Region: Stenje village, 870, 24.07.2005; Makedonski Brod Region: 

Dolno Botušje village, 970, 03.06.2012; Kosovo village, 680, 

04.06.2012; Samokov village, 900, 24.07.2008; Slatina village, 720, 

24.07.2008; Maleševski Planini Mts.: Klepalo hill, 1400, 22.08.2009; 

Lake Berovsko Ezero, 980, 23.08.2009; Ramna reka River, 1000, 

06.08.2010; Marjanska Planina Mt.: Mamino Brdo, 650, 03.05.2005; 

Nidže Mt.: Kopanki, 1550, 12.07.2002; Osogovski Planini Mts.: 

Bukovec, 1423, 15.05.2008; Dobrevo village, 603, 13.08.2008; Goli 

Rid, 1216, 16.05.2008; Ivova Glava, 1318, 30.08.2008; Jamište village, 

1062, 18.07.2008; Lisje Brdo, 1569, 14.07.2007; Luto Brdo, 1332, 

01.08.2008; Pavlevi Nivi, 1366, 15.07.2008; Plavica, 1297, 16.05.2008; 

Popova Čuka hill, 597, 10.10.2008; Ramni Nivi, 830, 15.05.2008; 

Rusov Grob, 850, 11.10.2008; Sokolarci village, 550, 16.04.2009; 

Toranica mine, 1068, 19.03.2007; Tursko Rudari village, 550, 

14.08.2008; Pelister Mt.: Kunina Stena, 1700, 19.10.2002; Niže Pole 

village, 10.08.2002; Tutunica, 1150, 10.08.2002; Plakenska Planina 

Mt.: Konjarnik, 1275, 19.07.2008; Streževo village, 787, 14.05.2007; 

Prilep Region: Brailovo village, 950, 14.03.2010; Skopje Region: 

Blace village, 600, 22.03.2009; Brazda village, 450, 22.03.2009; 

Dervenska Klisura gorge, 570, 30.03.2007; Divlje village, 800, 

14.08.2007; Orman village, 370, 30.04.2006; Skopska Crna Gora Mt.: 

Biljak peak, 1250, 22.03.2009; Čučer village, 600, 22.03.2009; Pržar 

peak, 1200, 22.03.2009; Strumica Region: Monospitovsko Blato, 210, 

19.06.2007; Murtino village, Tunevica, 210, 23.05.2008; Šar Planina 

Mt.: Dedel Beg, 1460, 01.07.2006; Gorno Jelovce village, 1216, 

17.05.2007; Lešnica, 1450, 16.07.2009; Novo Selo village, 1340, 

20.07.2009; Rogačevo village, 900, 30.07.2008; Vrutok village, 1000, 

30.07.2008; Veles Region: Bogomila village, 680, 16.09.1999; Slp 

village, 410, 20.04.2009. 

Zamenis situla (Linnaeus, 1758) 

Gradsko Region: Ulanci village, 180, 14.09.2009; Katlanovo Region: 

Katlanovska banja, 330, 04.11.2012; Kavadarci Region: Rosoman 

village, 200, 13.09.2009; Lake Dojran Region: Crničani village, 180, 

02.08.2002; Dojran, 150, 27.03.2008; Negotino Region: Eneševo 

village, 298, 27.05.2008; Skopje Region: Kačanička klisura gorge, 

300, 31.03.2007; Treska River gorge, Matka, 650, 12.09.2004; 

Valandovo Region: Udovo village, 150, 30.03.2012; Veles Region: 

Babuna River, Gradište, 170, 20.09.2010; Bekirlija village, 230, 

08.04.2007; Veles, Kameno Brdo, 420, 20.09.2010. 

Elaphe quatuorlineata (Lacepède, 1789) 

Azot Region: Krajnici village, Krajnici-Vojnica, 28.06.2006; Bogdanci 

Region: Gjavato village, 100, 03.05.2005; Demir Kapija Region: 

Došnica village, 300, 01.06.2009; Galičica Mt.: Pljuska, 940, 

26.07.2005; Gevgelija Region: Kovanska Reka River, 60, 25.05.2008; 

Gradsko Region: Ulanci village, 180, 14.09.2009; Kavadarci Region: 

Rosoman village, 200, 13.09.2009; Kumanovo Region: Klečevce 

village, 366, 14.06.2011; Pelince village, 526, 20.03.2007; Lake Dojran 

Region: Crničani village, 180, 27.03.2008; Dojran, 150, 27.03.2008; 

Lake Prespa Region: Golem Grad Island, 870, 25.04.2008; Oteševo 

village, 910, 27.07.2006; Stenje village, 870, 24.07.2005; Negotino 

Region: Eneševo village, 290, 16.08.2010; Pepelište village, 193, 

15.08.2010; Osogovski Planini Mts.: Golak hill, 330, 10.10.2008; 

Gorno Gradče vilage, 610, 15.07.2007; Zletovska reka River, 800, 


30.03.2007; Kačanička klisura gorge, 300, 31.03.2007; Katlanovsko 

Blato, 340, 30.05.2009; Treska River gorge, Matka, 340, 12.09.2004; 

Strumica Region: Čam Čiflik, 500, 26.05.2008; Monospitovsko Blato, 

210, 23.05.2008; Valandovo Region: Sobri village, 170, 25.05.2008; 

Veles Region: Babuna River, 180, 27.05.2001; Babuna River, 

Gradište, 170, 20.09.2010; Bekirlija village, 300, 10.06.2009; Kišino 

village, 400, 10.06.2009; Slp village, 410, 20.04.2009; Veles, Kameno 

Brdo, 420, 20.09.2010. 

Platyceps najadum (Eichwald, 1831) 

Galičica Mt.: Leskoec village, 900, 28.07.2003; Gevgelija Region: 

Miravci village, 250, 03.05.2005; Gradsko Region: Ulanci village, 

180, 14.09.2009; Kavadarci Region: Drenovo village, Drenovska 

george, 200, 30.04.2010; Rosoman village, 200, 13.09.2009; Kratovo 

Region: Dimonce village, 530, 15.06.2011; Filipovci village, 

21.04.2010; Lake Dojran Region: Čaušli village, 720, 10.06.2006; Lake 

Ohrid Region: Trpejca village, 750, 29.08.2007; Lake Prespa Region: 

Stenje village, 870, 24.07.2005; Mariovo Region: Beluče village, 

20.04.2007; Brest, Agro "Mariovo", 30.04.2005; Negotino Region: 

Eneševo village, 290, 16.08.2010; Pepelište village, 193, 15.08.2010; 

Nidže Mt.: Skočivir village, 650, 18.07.2002; Osogovski Planini Mts.: 

Dobrevo village, 603, 13.08.2008; Leski village, 910, 21.07.2007; 

Orizari village, 406, 11.10.2008; Popova Čuka hill, 597, 10.10.2008; 

Sokolarci village, 550, 16.04.2009; Tursko Rudari village, 550, 

14.08.2008; Skopje Region: Čaršija, 250, 06.04.2003; Kožle village, 

Prnar, 10.06.2007; Volkovo village, 350, 23.05.2004; Strumica 

Region: Monospitovo village, 210, 28.05.2008; Veles Region: Babuna 

River, 170, 20.09.2010; Bekirlija village, 400, 10.06.2009; Kišino 

village, 400, 10.06.2009; Slp village, 410, 20.04.2009; Veles, 420, 

20.09.2010. 

Malpolon monspessulanus Hermann, 1804 

Gradsko Region: Ulanci village, 180, 14.09.2009; Kavadarci Region: 

Rosoman village, 200, 13.09.2009; Kumanovo Region: Beljakovce 

village, 470, 15.06.2011; Pelince village, 526, 20.03.2007; Lake Ohrid 

Region: Trpejca village, 750, 20.06.2009; Lake Prespa Region: Stenje 

village, 860, 17.04.2006; Mariovo Region: Manastir village, road to 

Vitološte village, 13.04.2012; Negotino Region: Eneševo village, 290, 

16.08.2010; Pepelište village, 193, 15.08.2010; Osogovski Planini 

UNCORRECTED PROOF



Mts.: Sokolarci village, 550, 13.04.2008; Valandovo Region: Udovo 

village, 150, 30.03.2012; Veles Region: Babuna River, Gradište, 170, 


20.04.2009; Sopot village, 08.04.2009; Veles, Kameno Brdo, 420, 

20.09.2010. 

Telescopus fallax (Fleischmann, 1831) 

Gradsko Region: Ulanci village, 196, 13.09.2009; Lake Dojran 

Region: Crničani village, 180, 09.06.2009; Skopje Region: Kačanička 

klisura gorge, 300, 17.03.2008; Orman village, 260, 19.05.2004; 

Treska River gorge, Matka, 340, 22.06.2009. 

Eryx jaculus (Linnaeus, 1758) 

Lake Dojran Region: Nikolić village, 150 m, 05.05.2005. 

Vipera ammodytes (Linnaeus, 1758) 

Babuna Mt.: Trojaci village, 582, 12.03.2010; Berovo Region: Blatec 



19.03.2010; Bistra Mt.: Daboec, 1000, 27.08.2010; Galičnik village, 

Selce-Galičnik, 30.06.2012; Straža pass, 1200, 22.07.2010; Bitola 

Region: Dobruševo village, 595, 22.09.2006; Golemo Konjari village, 

610, 28.04.2006; Lavci village, 960, 29.04.2006; Topolčani village, 

600, 28.04.2006; Bogdanci Region: Gjavato village, 100, 03.05.2005; 

Čučer-Sandevo Region: Brest village, 1146, 09.08.2012; Delčevo 

Region: Crnik village, 983, 17.09.2009; Grad village, 800, 17.09.2009; 


820, 23.07.2008; Vranešnica village, 1100, 24.07.2008; Demir Kapija 

Region: Došnica village, 300, 01.06.2009; Koprišnica village, 500, 

29.06.2005; Dešat Mt.: Žirovnica village, 1200, 28.07.2008; Galičica 

Mt.: Bolno village, 1000, 12.03.2010; Gevgelija Region: Gorničet 

village, 210, 26.05.2008; Kovanska Reka River, 60, 25.05.2008; 

Gradečka planina Mt.: Salandžak, 840, 26.07.2010; Gradsko Region: 

Ulanci village, 180, 14.09.2009; Ilinska Planina Mt.: Belčišta village, 

Belčiško Blato, 850, 25.07.2008; Belica village, 900, 01.09.2008; Lazoi, 

1150, 17.07.2008; Jakupica Mt.: Kapina village, 800, 08.09.2010; 

Selište village, 1200, 08.09.2010; Karaorman Mt.: Botun village, 950, 

29.08.2008; Pesočani village, 950, 27.08.2010; Kavadarci Region: 

Lake Tikveško Ezero, Kotursko, 08.04.2006; Ljubaš hill, 580, 

25.07.2010; Tumba hill, 597, 17.08.2010; Kičevo Region: Srbjani 

village, 870, 26.07.2008; Kitka Mt.: Kitka hut, 1100, 01.09.2006; 

Kočani Region: Obleševo village, 305, 09.10.2008; Korab Mt.: 

Strezimir, 1200, 29.07.2008; Kozjak Mt.: Bešište village, 900, 

22.05.2007; Kožuf Mt.: Kalugerska Čuka, 737, 03.05.2005; Lukar, 

1047, 17.07.2005; Sermenin village, 400, 26.05.2008; Kratovo Region: 

Kovačevci, 650, 16.06.2011; Kratovo, 712, 17.03.2007; Krilatica 

village, 698, 11.06.2011; Kuklica village, 700, 16.06.2011; Šopsko 

rudare village, 580, 16.06.2011; Talašmance village, 639, 11.06.2011; 


1320, 21.05.2011; Drenje village, 680, 13.06.2011; Golema Crcorija 

village, 1120, 20.05.2011; Ogut village, 900, 20.05.2011; Tlminci 

village, 618, 12.06.2011; Trnovo village, 1100, 20.05.2011; Zlatkovci, 



village, 501, 09.08.2012; Klečevce village, 366, 14.06.2011; Kokošinje 

village, 760, 12.05.2012; Mlado Nagoričane village, 398, 20.03.2007; 

Orašac village, 500, 21.03.2007; Pelince village, 526, 20.03.2007; 

Vaksince village, 600, 30.05.2008; Lake Dojran Region: Crničani 

village, 180, 02.08.2002; Lake Prespa Region: Carev Dvor village, 

860, 11.03.2010; Ezerani village, 860, 11.03.2010; Makedonska 

Kamenica Region: Dramče village, 800, 31.03.2009; Makedonski 

Brod Region: Belica village, 940, 24.07.2008; Dolno Botušje village, 

970, 03.06.2012; Kosovo village, 680, 04.06.2012; Krapa village, 1046, 

24.07.2010; Makedonski Brod, 700, 24.07.2008; Samokov village, 900, 

24.07.2008; Suvodol village, 600, 24.07.2008; Volče village, 1000, 

04.06.2012; Maleševski Planini Mts.: Klepalo hill, 1400, 22.08.2009; 

Ramna reka River, 1000, 29.03.2008; Negotino Region: Eneševo 

village, 290, 16.08.2010; Pepelište village, 193, 15.08.2010; Osogovski 

Planini Mts.: Kapetanska Maala, 1141, 31.03.2009; Katanska Maala, 

850, 31.03.2009; Moždivnjak village, 700, 18.03.2007; Sredni Rid, 

850, 19.09.2010; Toranica mine, 1068, 19.03.2007; Zletovska reka 

River, 800, 05.06.2008; Plačkovica Mt.: Zrnovci village, 590, 


Prilep Region: Brailovo village, 950, 14.03.2010; Čelopeci village, 

593, 13.03.2010; Krivogaštani village, 670, 14.03.2010; Lažani village, 

630, 13.03.2010; Rusjaci village, 620, 12.03.2010; Zrze village, 700, 

06.06.2012; Radoviš Region: Babino Platno, 547, 21.03.2006; Selečka 

Planina Mt.: Čanište village, 520, 23.05.2007; Skopje Region: Blace 

village, 600, 22.03.2009; Brazda village, 450, 22.03.2009; Čaršija, 270, 

06.04.2003; Dervenska Klisura gorge, 570, 30.03.2007; Jabolci village, 

700, 15.04.2009; Kačanička klisura gorge, 300, 17.03.2008; 

Katlanovsko Blato, 340, 30.05.2009; Sopište village, 710, 15.04.2009; 

Taorska Klisura gorge, 300, 29.06.2007; Skopska Crna Gora Mt.: 

Biljak peak, 1250, 22.03.2009; Čučer village, 600, 22.03.2009; Lipkovo 

village, 580, 22.03.2007; Pržar peak, 1200, 22.03.2009; Slaviško pole 

Region: Ginovci village, 500, 20.04.2007; Psača village, 540, 

18.03.2007; Rankovce village, 490, 12.06.2011; Rankovce village, 480, 

20.04.2007; Stogovo Mt.: Kodžadžik village, 1200, 15.05.2007; 

Papradnik village, 700, 24.07.2006; Strumica Region: Čam Čiflik, 

500, 26.05.2008; Monospitovo village, Selište, 210, 28.05.2008; 

Monospitovsko Blato, 210, 19.06.2007; Novo Selo village, 250, 

24.05.2008; Novo Selo village, Rakovec, 210, 14.06.2007; Suva Gora 

Mt.: Konjski Grob, 1030, 31.07.2010; Šar Planina Mt.: Gajre village, 

750, 10.04.2003; Gorno Jelovce village, 1216, 17.05.2007; Novo Selo 

village, 1340, 20.07.2009; Rogačevo village, 900, 30.07.2008; Torbeški 

Most, 1400, 29.07.2008; Vrutok village, 1000, 30.07.2008; Štip Region: 

Bogoslovec village, 755, 12.06.2009; Erdželia village, 260, 11.06.2009; 

Leskovica village, 400, 21.03.2006; Radanja village, 400, 21.03.2006; 

Sofilari village, 320, 21.03.2006; Valandovo Region: Sobri village, 

170, 25.05.2008; Udovo village, 150, 30.03.2012; Veles Region: 




Vinica, Kartal, 670, 19.09.2010; Vinica, Kale, 470, 19.09.2010; Vlaina 


Vipera berus (Linnaeus, 1758) 

Jakupica Mt.: Begovo Pole, 1900, 17.07.1999; Karaorman Mt.: 

Popova Livada, 1714, 27.08.2010; Kožuf Mt.: Dudica Peak, Dudički 

Kolibi, 20.07.2005; Konjska Glava, 1750, 20.07.2005; Top peak, 1781, 

10.07.2005; Nidže Mt.: Belo Grotlo, 1750, 16.07.2002; Pelister Mt.: 

Lake Golemo Ezero, 2100, 19.07.2001; Široka, 1960, 10.07.2001; Viroi, 

2100, 19.10.2002; Plakenska Planina Mt.: Bojkoec, 1620, 03.10.2008; 

Stogovo Mt.: Bijak Dorok peak, 1900, 15.05.2007; Šar Planina Mt.: 

Lukovo Pole, 1600, 13.07.2012. 

Vipera ursinii (Bonaparte, 1835) 

Šar Planina Mt.: Lukovo pole, 1600 m, 13.07.2012. 

UNCORRECTED PROOF



Appendix II – Literature data 

Emys orbicularis (Linnaeus, 1758) 

Demir Kapija Region: Mravnica (Dolfein, 1921); Jablanica Mt.: 

Dušman (Sterijovski et al. 2010a); Lake Ohrid Region: springs of 

Crni Drim River, St. Naum Monastery (Dimovski 1960, 1963, 1966a, 

1981); Skopje Region: without precise locality (Dimovski 1960, 1963, 

1966a, 1981). 

Mauremys rivulata (Valenciennes, 1833) 

Demir Kapija Region: Mravnica (Dolfein, 1921); Gevgelija Region: 

Gevgelija (Radovanović 1951); Lake Dojran Region: Dojran 

(Dolfein, 1921; Karaman 1928, 1937, 1939; Radovanović 1951); 

Strumica Region: Udovo village (Radovanović 1951). 

Testudo graeca Linnaeus, 1758 

Demir Kapija Region: Demir Kapija, Mravnica (Dolfein, 1921); 

Gevgelija Region: Gevgelija (Radovanović 1951; Džukić 1972); 

Gradsko Region: Nogaevci village, Ulanci village (Dimovski 1960, 

1963, 1966a, 1971); Jablanica Mt.: Gorna Belica village (Sterijovski et 

al. 2010a); Kožuf Mt.: Smrdliva Voda (Sterijovski et al. 2010b); Lake 

Dojran Region: Dojran (Dolfein, 1921; Radovanović 1951); Lake 

Ohrid Region: without precise locality (Karaman 1928, 1939); Lake 

Prespa Region: without precise locality (Karaman 1928, 1939); 

Negotino Region: Djidimirci village, Eneševo village, Mušanci 

village (Dimovski 1960, 1963, 1966a, 1971); Skopje Region: without 

precise locality (Dimovski 1960, 1963, 1966a, 1971); Skopje 

(Karaman 1928, 1939); Štip Region: Adjibegovo, between Štip and 

Krivolak, Bogoslovec, Delisinci village (Dimovski 1960, 1963, 1966a, 

1971); Strumica Region: Strumica (Džukić 1972); Vardar River 

valley, without precise locality (Radovanović 1951). 

Testudo hermanni Gmelin, 1789 

Demir Kapija Region: Demir Kapija, Mravnica (Dolfein, 1921); 

Gevgelija Region: Gevgelija (Radovanović 1951); Gradsko Region: 

Ulanci village (Dimovski 1960, 1963, 1966a, 1971, 1981); Jakupica 

Mt.: Oreše village (Sterijovski et al. 2002); Kožuf Mt.: Mihajlovo 

(Sterijovski et al. 2010b); Lake Dojran Region: Dojran (Dolfein, 1921; 

Radovanović 1951); Lake Ohrid Region: without precise locality 

(Karaman 1922, 1937, 1939; Radovanović 1951); Elšani village, 

Ljubaništa village, Peštani village, St. Naum Monastery, Trpejca 

village (Dimovski 1960, 1963, 1966a, 1971, 1981); Lake Prespa 

Region: Oteševo village, Stenje village (Dimovski 1960, 1963, 1966a, 

1971, 1981); Negotino Region: Djidimirci village, Eneševo village, 

Mušanci village (Dimovski 1960, 1963, 1966a, 1971, 1981); Šar 

Planina Mt.: Lisec village (Džukić 1972); Tetovo (Radovanović 

1951); Skopje Region: without precise locality (Dimovski 1960, 1963, 

1966a, 1971, 1981); Skopje (Karaman 1922, 1937, 1939); Štip region: 

between Štip and Krivolak, Bogoslovec (Dimovski 1960, 1963, 

1966a, 1971, 1981); Ovče Pole, Štip (Karaman 1922, 1937, 1939); 

Vardar River valley, without precise locality (Radovanović 1951); 

Veles Region: Veles (Karaman 1922, 1937, 1939). 

Anguis fragilis Linnaeus, 1758 

Demir Kapija Region: Gradec (Dolfein, 1921); Galičica Mt.: without 

precise locality (Dimovski 1960, 1963, 1966a, 1981); Jakupica Mt.: 

without precise locality (Karaman 1928, 1931, 1939); Korab Mt.: 

without precise locality (Karaman 1928, 1931, 1939); Ogražden Mt.: 

Ogražden peak (Sterijovski & Stamatoski 2002); Šar Planina Mt.: 

without precise locality (Karaman 1928, 1931, 1939); Skopje Region: 

without precise locality (Dimovski 1960, 1963, 1966a, 1981). 

Pseudopus apodus (Pallas, 1775) 

Lake Dojran Region: Dojran (Dolfein, 1921); lower flow of the 

Vardar River, without precise locality (Radovanović 1951); 

Strumica Region: Udovo village (Dolfein, 1921). 


Ablepharus kitaibelii Bibron & Bory, 1833 

Bigla Mt.: Gopeš (Dolfein, 1921); Galičica Mt.: Prespa side 

(Dimovski 1959a, 1960, 1963, 1966a, 1981); Jakupica Mt.: Čeples hut 

(Sterijovski et al. 2002); Nežilovo village (Dimovski 1959a, 1960, 

1963, 1966a, 1981); Papradište village (Ljubisavljević et al. 2002); 

Konečka Planina Mt.: Ogražden Mt.: Nivičino village (Sterijovski & 

Stamatoski 2002); Pešternica village (Ljubisavljević et al. 2002); 

Skopje Region: Vodno Mt. (Dimovski 1959a, 1960, 1963, 1966a, 

1981); Veles Region: Babuna hill, Han Abdipaša (Dolfein, 1921). 

Cyrtopodion kotschyi (Steindachner, 1870) 

Demir Kapija Region: Kalučkovo hill (Dolfein, 1921); Gevgelija 

Region: Gevgelija (Karaman 1928, 1937, 1939; Radovanović 1951; 

Džukić 1972); Lake Dojran Region: Dojran (Dolfein, 1921; Karaman 

1928, 1937, 1939); Skopje Region: Skopje (Karaman 1928, 1937, 1939; 

Dimovski 1960, 1963, 1966a); Strumica Region: Udovo village 

(Dolfein, 1921); Valandovo Region: Valandovo (Karaman 1928, 

1937, 1939); Veles Region: Veles (Karaman 1928, 1937, 1939). 

Algyroides nigropunctatus (Duméril & Bibron, 1839) 

Debar Region: Debar (Radovanović 1951); Lake Ohrid Region: 

Peštani village (Dimovski 1963, 1964, 1966a,b, 1981); Lake Prespa 

Region: Oteševo village (Dimovski 1963, 1964, 1966a,b, 1981); 

Skopje Region: without precise locality (Radovanović 1951); Treska 

River gorge, Matka (Karaman 1931, 1937, 1939; Dimovski 1960, 

1963, 1964, 1966a,b). 

Lacerta agilis Linnaeus, 1758 

Bistra Mt.: Golem Brzovec peak, Koritnik bačila, Mal Brzovec peak, 

Maskarevec, Ploča (Sterijovski et al. 2003); Korab Mt.: without 

precise locality (Dimovski 1959a, 1964); Pelister Mt.: without 

precise locality (Dolfein, 1921); Pelister Mt.: Golemo Ezero Lake 

(Dimovski 1959a, 1964); Malo Ezero Lake (Dimovski 1959a, 1964). 

Lacerta trilineata Bedriaga, 1886 

Bistra Mt.: Tresonče village, Boškov most (Sterijovski et al. 2003); 

Demir Kapija Region: Demir Kapija (Dolfein, 1921); Galičica Mt.: 

Prespa side (Dimovski 1959b, 1960, 1963, 1966a, 1971, 1981); 

Gradsko Region: Ulanci village (Dimovski 1959b, 1960, 1963, 1966a, 

1971, 1981); Katlanovo Region: railroad station Pčinja, railroad 

station Rajko Žinzifov (Dimovski 1959b, 1960, 1963, 1966a, 1971, 

1981); Lake Dojran Region: Dojran (Karaman 1922, 1928, 1931, 1937, 

1939); Lake Ohrid Region: without precise locality (Karaman 1922, 

1928, 1931, 1937, 1939); Negotino Region: Djidimirci village, 

Eneševo village, Mušanci village (Dimovski 1959b, 1960, 1963, 

1966a, 1971, 1981); Ogražden Mt.: Nivičino village, Boraj hill 

(Sterijovski & Stamatoski 2002); Skopje Region: without precise 

locality (Dimovski 1959b, 1960, 1963, 1966a, 1971, 1981); Skopje 

(Karaman 1922, 1928, 1931, 1937, 1939); Skopska Crna Gora Mt.: 

Banjani village (Karaman 1922, 1928, 1931, 1937, 1939); Štip Region: 

between Štip and Krivolak, Bogoslovec (Dimovski 1959b, 1960, 

1963, 1966a, 1971, 1981); Ovče Pole, Štip (Karaman 1922, 1928, 1931, 

1937, 1939); Veles Region: Veles (Dolfein, 1921). 

Lacerta viridis (Laurenti, 1768) 

Bistra Mt.: Škrka (Sterijovski et al. 2003); Demir Kapija Region: 

Gradec (Dolfein, 1921); Galičica Mt.: Ohrid side, Prespa side 

(Dimovski 1959a, 1960, 1963, 1966a, 1981); Jablanica Mt.: Skala 

(Sterijovski et al. 2010a); Kožuf Mt.: Mala Rupa (Dolfein, 1921); 

Konopište village, Konjska Glava, Mala Rupa (Sterijovski et al. 

2010b); Šar Planina Mt.: without precise locality (Dolfein, 1921); 

Ogražden Mt.: Lake Turija Ezero, Nivičino village, Boraj hill 

(Sterijovski et al. 2002); Skopje Region: without precise locality 

(Dimovski 1959a, 1960, 1963, 1966a, 1981); Skopje (Džukić 1972); 

Treska River gorge (Karaman 1931); Vodno Mt. (Dimovski 1959a, 

1960, 1963, 1966a, 1981); Skopska Crna Gora Mt.: Blagoveštenja 

monastery (Karaman 1931). 

Podarcis erhardii (Bedriaga, 1882) 

Bitola Region: Bitola (Karaman 1922, 1928, 1931, 1939; Radovanović 

1951); Brod: without precise locality (Karaman 1922, 1928, 1931, 

1939); Delčevo Region: Delčevo (Karaman 1922, 1928, 1931, 1939); 

Gevgelija Region: Gevgelija (Karaman 1922, 1928, 1931, 1939;



Radovanović 1951); Gradsko Region: Gradsko (Karaman 1922, 

1928, 1931, 1939); Nogaevci village (Dimovski 1959b, 1960, 1963, 

1966a,b, 1971, 1981); Jakupica Mt.: without precise locality 

(Karaman 1922, 1928, 1931, 1939); Katlanovo Region: railroad 

station Pčinja, railroad station Rajko Žinzifov (Dimovski 1959b, 

1960, 1963, 1966a,b, 1971, 1981); Kočani Region: Kočani (Karaman 

1922, 1928, 1931, 1939); Kratovo Region: Kratovo (Karaman 1922, 

1928, 1931, 1939); Kumanovo Region: Kumanovo, Mlado 

Nagoričane village (Karaman 1922, 1928, 1931, 1939); Lake Dojran 

Region: Dojran (Karaman 1922, 1928, 1931, 1939; Radovanović 

1951); Lake Ohrid Region: without precise locality (Karaman 1922, 

1928, 1931, 1939; Radovanović 1951); Ljubaništa village (Dimovski 

1959b, 1960, 1963, 1966a,b, 1971, 1981); St. Naum Monastery 

(Džukić 1972); Lake Prespa Region: without precise locality 

(Karaman 1922, 1928, 1931, 1939; Radovanović 1951); Golem Grad 

Island (Dimovski 1959b, 1960, 1963, 1966a,b, 1971, 1981); Kališta 

village (Sterijovski et al. 2010a); Negotino Region: Djidimirci 

village, Mušanci village (Dimovski 1959b, 1960, 1963, 1966a,b, 1971, 

1981); Prilep Region: Prilep (Karaman 1922, 1928, 1931, 1939); Šar 

Planina Mt.: Tetovo (Radovanović 1951; Džukić 1972); Skopje 

Region: without precise locality (Dimovski 1959b, 1960, 1963, 

1966a,b, 1971, 1981); Skopje (Karaman 1922, 1928, 1931, 1939; 

Radovanović 1951); Treska River gorge, Matka (Dimovski 1959b, 

1960, 1963, 1966a,b, 1971, 1981); Vodno Mt. (Karaman 1922, 1928, 

1931, 1939); Skopska Crna Gora Mt.: Blagoveštenja monastery 

(Karaman 1922, 1928, 1931, 1939); Štip Region: Adjibegovo, 

between Štip and Krivolak, Bogoslovec village, Bregalnica River 

valley, Delisinci village (Dimovski 1959b, 1960, 1963, 1966a,b, 1971, 

1981); Ovče Pole (Karaman 1922, 1928, 1931, 1939; Dimovski 1959b, 

1960, 1963, 1966a,b); Štip (Karaman 1922, 1928, 1931, 1939); 

Strumica Region: Strumica (Karaman 1922, 1928, 1931, 1939); Veles 

Region: Veles (Karaman 1922, 1928, 1931, 1939). 

Podarcis muralis (Laurenti, 1768) 

Berovo Region: Berovo (Karaman 1928, 1931, 1939); Bistra Mt.: 

Koritnik Bačila, Rosoki village, Rosočka reka River, Tresonče 

village (Sterijovski et al. 2003); Debar Region: Debar (Karaman 

1928, 1931, 1939); Demir Kapija Region: Gradec (Dolfein, 1921); 

Jakupica Mt.: Salakova Mt. (Karaman 1928, 1931, 1939); Oreše 

village, Papradište village, Solunska Glava peak (Sterijovski et al. 

2002); Kičevo Region: Kičevo (Karaman 1928, 1931, 1939); Kožuf 

Mt.: Dudica peak, Kapetnova češma, Konopište village, Konjska 

Glava, Mala Rupa, Mihajlovo, Porta, Šarenka, Todorčevica, Zelen 

Breg (Sterijovski et al. 2010b); Kriva Palanka Region: Kriva Palanka 

(Karaman 1928, 1931, 1939); Lake Ohrid Region: without precise 

locality (Karaman 1928, 1931, 1939; Radovanović 1951); Šar Planina 

Mt.: without precise locality (Dolfein, 1921); Gostivar (Karaman 

1928, 1931, 1939); Tetovo (Džukić 1972); Skopje Region: without 

precise locality (Dimovski 1959a, 1960, 1963, 1966a); Skopje (Džukić 

1972); Vodno Mt. (Dimovski 1959a, 1960, 1963, 1966a); Žeden hill 

(Karaman 1928, 1931, 1939); Skopska Crna Gora Mt.: without 

precise locality (Karaman 1928, 1931, 1939); Veles Region: Veles 

(Dolfein, 1921). 

Podarcis tauricus (Pallas, 1814) 

Bitola Region: Bitola (Karaman 1922, 1928, 1931, 1937, 1939); 

Galičica Mt.: Ohrid side, Prespa side (Dimovski 1959a, 1960, 1963, 

1966a, 1971, 1981); Galičica Mt.: without precise locality (Karaman 

1922, 1928, 1931, 1937, 1939); Gradsko Region: Ulanci village 

(Dimovski 1959a, 1960, 1963, 1966a, 1971, 1981); Lake Dojran 

Region: Dojran (Karaman 1922, 1928, 1931, 1937, 1939); Lake Ohrid 

Region: without precise locality (Karaman 1922, 1928, 1931, 1937, 

1939; Radovanović 1951); St. Naum Monastery (Džukić 1972); Lake 

Prespa Region: Resen (Karaman 1922, 1928, 1931, 1937, 1939); 


village (Dimovski 1959a, 1960, 1963, 1966a, 1971, 1981); Šar Planina 

Mt.: without precise locality (Dolfein, 1921); Tetovo (Radovanović 

1951); Skopje Region: without precise locality (Dimovski 1959a, 

1960, 1963, 1966a, 1971, 1981); Skopje (Karaman 1922, 1928, 1931, 

1937, 1939; Radovanović 1951); Vodno Mt. (Dimovski 1959a, 1960, 

1963, 1966a, 1971, 1981); Štip Region: between Štip and Krivolak, 

Bogoslovec (Dimovski 1959a, 1960, 1963, 1966a, 1971, 1981); Štip 

(Karaman 1922, 1928, 1931, 1937, 1939); Strumica Region: Strumica 

(Karaman 1922, 1928, 1931, 1937, 1939). 

Zootoca vivipara von Jacquin, 1787 

Bistra Mt.: Medenica peak (Sterijovski et al. 2003); Korab Mt.: 

without precise locality (Karaman 1939; Radovanović 1951; 

Dimovski 1964); Šar Planina Mt.: without precise locality (Doflein 

1921; Karaman 1939; Radovanović 1951); Rudoka Mt. (Džukić 

1972); Titov Vrv peak (Dimovski 1964; Džukić 1972). 

Typhlops vermicularis (Daudin, 1803) 

Demir Kapija Region: Demir Kapija (Dimovski 1959b); Katlanovo 

Region: mouth of Pčinja River (Karaman 1931, 1937, 1939); railroad 

station Pčinja (Radovanović 1941, 1951, 1957; Dimovski 1959b); 

railroad station Rajko Žinzifov (Dimovski 1959b); Kavadarci 

Region: Mravince village (Buresch & Zonkow 1934); Lake Dojran 

Region: Dojran (Doflein 1921); Skopje Region: Gazi Baba, springs of 

Rašče River (Dimovski 1959b); Taor village (Karaman 1931, 1937, 

1939); Vodno Mt. (Radovanović 1941, 1951, 1957; Džukić 1972); 

Veles Region: Topolka River gorge (Karaman 1931, 1937, 1939). 

Coronella austriaca Laurenti, 1768 

Bistra Mt.: Mavrovi Anovi (Karaman 1928, 1931, 1939); Golem 

Brzovec peak (Sterijovski et al. 2003); Galičica Mt.: without precise 

locality (Dimovski 1960, 1963, 1966a, 1981); Jakupica Mt.: without 

precise locality (Doflein 1921; Karaman 1928, 1931, 1939); Kožuf 

Mt.: Konjsko village, Mala Rupa (Doflein 1921); Konopište village 

(Sterijovski et al. 2010b); Ogražden Mt.: Bezgaštevo village 

(Sterijovski & Stamatoski 2002); Skopje Region: without precise 

locality (Dimovski 1960, 1963, 1966a, 1981); Skopje (Džukić 1972); 

Treska River gorge (Karaman 1928, 1931, 1939). 

Natrix natrix (Linnaeus, 1758) 

Bistra Mt.: Rosoki village, Rosočka reka River (Sterijovski et al. 

2003); Galičica Mt.: Ohrid side, Prespa side (Dimovski 1960, 1963, 

1966a, 1971, 1981); Jakupica Mt.: Čeples hut (Sterijovski et al. 2002); 

Kožuf Mt.: Mala Rupa (Doflein 1921); Konjska reka River, Dlabok 

Dol (Sterijovski et al. 2010b); Lake Ohrid Region: without precise 

locality (Karaman 1922, 1928, 1931, 1939); Lake Prespa Region: 

Resen (Karaman 1922, 1928, 1931, 1939); Skopje Region: without 

precise locality (Dimovski 1960, 1963, 1966a, 1971, 1981); 

Katlanovsko Lake (Karaman 1922, 1928, 1931, 1939); Skopje 

(Karaman 1922, 1928, 1931, 1939; Džukić 1972); Štip Region: 

between Štip and Krivolak (Dimovski 1960, 1963, 1966a, 1971, 

1981); Veles Region: Veles (Karaman 1922, 1928, 1931, 1939). 

Natrix tessellata (Laurenti, 1768) 

Bistra Mt.: Rosoki village, Rosočka reka River (Sterijovski et al. 

2003); Demir Kapija Region: Gradec (Doflein 1921); Galičica Mt.: 

Ohrid side, Prespa side (Dimovski 1960, 1963, 1966a, 1971, 1981); 

Gradsko Region: Ulanci village (Dimovski 1960, 1963, 1966a, 1971, 

1981); Jakupica Mt.: Babuna River (Sterijovski et al. 2002); Lake 

Dojran Region: Dojran (Karaman 1922, 1928, 1931, 1939); Lake 

Ohrid Region: without precise locality (Karaman 1922, 1928, 1931, 

1939); Kališta village (Sterijovski et al. 2010a); Lake Prespa Region: 

without precise locality (Karaman 1922, 1928, 1931, 1939); Negotino 

Region: Djidimirci village, Eneševo village, Mušanci village 

(Dimovski 1960, 1963, 1966a, 1971, 1981); Skopje Region: without 

precise locality (Dimovski 1960, 1963, 1966a, 1971, 1981); 

Katlanovsko Lake (Karaman 1922, 1928, 1931, 1939); Štip Region: 

between Štip and Krivolak, Bogoslovec, Bregalnica River valley, 

Ovče Pole (Dimovski 1960, 1963, 1966a, 1971, 1981); Veles Region: 

Veles (Doflein 1921; Karaman 1922, 1928, 1931, 1939). 

Dolichophis caspius (Gmelin, 1789) 

Bistra Mt.: Tresonče village (Sterijovski et al. 2003); Bitola Region: 

Bitola (Karaman 1928, 1931, 1939); Galičica Mt.: Ohrid side, Prespa 

side (Dimovski 1960, 1963, 1966a, 1971, 1981); Gradsko Region: 

Ulanci village (Dimovski 1960, 1963, 1966a, 1971, 1981); Jablanica 

Mt.: Šum (Džukić 1972); Negotino Region: Djidimirci village, 

UNCORRECTED PROOF



Eneševo village, Mušanci village (Dimovski 1960, 1963, 1966a, 1971, 

1981); Ogražden Mt.: Ogražden mine (Sterijovski & Stamatoski 

2002); Šar Planina Mt.: Tetovo (Džukić 1972); Skopje Region: 

without precise locality (Dimovski 1960, 1963, 1966a, 1971, 1981); 

(Karaman 1928, 1931, 1939; Džukić 1972); Skopska Crna Gora Mt.: 

Blagoveštenja monastery (Karaman 1928, 1931, 1939); Štip Region: 

between Štip and Krivolak, Bogoslovec (Dimovski 1960, 1963, 

1966a, 1971, 1981); Štip (Karaman 1928, 1931, 1939). 

Hierophis gemonensis (Laurenti, 1768) 

Debar Region: Debar (Petkovski et al. 2000/2001). 

Zamenis longissimus (Laurenti, 1768) 

Bistra Mt.: Mavrovi Anovi (Karaman 1931, 1939); Galičica Mt.: 

Ohrid side, Prespa side (Dimovski 1960, 1963, 1966a, 1981); 

Jakupica Mt.: Papradište village (Sterijovski et al. 2002); Kožuf Mt.: 

Mala Rupa (Doflein 1921); Mihajlovo (Sterijovski et al. 2010b); 

Skopje Region: without precise locality (Dimovski 1960, 1963, 

1966a, 1981); Skopska Crna Gora Mt.: Blagoveštenja monastery 

(Karaman 1931, 1939). 

Zamenis situla (Linnaeus, 1758) 

Demir Kapija Region: Gradec (Doflein 1921); Katlanovo Region: 

railroad station Pčinja, railroad station Rajko Žinzifov (Dimovski 

1959b, 1960, 1963, 1966a,b, 1971); Skopje Region: without precise 

locality (Dimovski 1959b, 1960, 1963, 1966a,b, 1971); Katlanovsko 

Lake (Karaman 1931); Treska River gorge, Matka (Dimovski 1959b, 

1960, 1963, 1966a,b, 1971); Skopska Crna Gora Mt.: Banjani village 

(Karaman 1931); Štip Region: between Štip and Krivolak (Dimovski 

1959b, 1960, 1963, 1966a,b, 1971); Vardar River valley: without 

precise locality (Radovanović 1951). 

Elaphe quatuorlineata (Lacepède, 1789) 

Demir Kapija Region: Demir Kapija (Doflein 1921); Galičica Mt.: 

Ohrid side (Dimovski 1960, 1963, 1966a, 1971, 1981); Gradsko 

Region: Nogaevci village, Ulanci village (Dimovski 1960, 1963, 

1966a, 1971, 1981); Negotino Region: Djidimirci village, Eneševo 

village, Mušanci village (Dimovski 1960, 1963, 1966a, 1971, 1981); 

Skopje Region: without precise locality (Dimovski 1960, 1963, 

1966a, 1971, 1981; Karaman 1928, 1931, 1939); Taor village 

(Karaman 1928, 1931, 1939); Štip Region: Adjibegovo, between Štip 

and Krivolak, Bogoslovec, Bregalnica River valley, Delisinci village, 

Ovče Pole (Dimovski 1960, 1963, 1966a, 1971, 1981); Vardar River 

valley: without precise locality (Radovanović 1951). 

Platyceps najadum (Eichwald, 1831) 

Gradsko Region: Ulanci village (Dimovski 1960, 1963, 1966a,b, 

1971); Jablanica Mt.: Višni village (Sterijovski et al. 2010a); Kožuf 

Mt.: Konopište village (Sterijovski et al. 2010b); Lake Dojran 

Region: without precise locality (Karaman 1928, 1931, 1937, 1939); 


village (Dimovski 1960, 1963, 1966a,b, 1971); Skopje Region: 

without precise locality (Dimovski 1960, 1963, 1966a,b, 1971); 

Katlanovsko Lake (Karaman 1928, 1931, 1937, 1939); Markova 

Kruška (Džukić 1972); Skopje (Džukić 1972); Treska River gorge 

(Karaman 1928, 1931, 1937, 1939); Treska River gorge, Matka 

(Dimovski 1960, 1963, 1966a,b, 1971); Štip Region: between Štip and 

Krivolak, Bogoslovec (Dimovski 1960, 1963, 1966a,b, 1971); Vardar 

River valley: without precise locality (Radovanović 1951). 

Malpolon monspessulanus Hermann, 1804 

Lake Dojran Region: without precise locality (Karaman 1928, 1931, 

1937, 1939); Skopje Region: without precise locality (Radovanović 

1951; Dimovski 1960, 1963, 1966a); Katlanovsko Lake (Karaman 

1928, 1931, 1937, 1939). 


Telescopus fallax (Fleischmann, 1831) 

Gradsko Region: Ulanci village (Dimovski 1960, 1963, 1966a, 1971); 

Lake Dojran Region: without precise locality (Karaman 1928, 1937, 

1939); Negotino Region: Djidimirci village, Eneševo village, 

Mušanci village (Dimovski 1960, 1963, 1966a, 1971); Skopje Region: 

without precise locality (Dimovski 1960, 1963, 1966a, 1971); Skopje 

(Karaman 1928, 1937, 1939; Džukić 1972); Štip Region: between Štip 

and Krivolak, Bogoslovec (Dimovski 1960, 1963, 1966a, 1971); 

Vardar River valley, without precise locality (Radovanović 1951). 

Eryx jaculus (Linnaeus, 1758) 

Gradsko Region: Ulanci village (Dimovski 1971); Kočani Region: 

Lepopelci village (Petrušev et al. 1990); Lake Dojran Region: Dojran 

(Doflein 1921); Osogovski Planini Mt.: Orizari village (Petrušev et 

al. 1990); Štip Region: between Štip and Krivolak (Dimovski 1971); 

Veles Region: Babuna River gorge (Džukić & Grubač 1988); Bašino 

village (Karaman 1955). 

Vipera ammodytes (Linnaeus, 1758) 

Bilina Mt.: Podržikonj village (Kriva Palanka) (Jelić et al. 2012); 

Bistra Mt.: Galičnik village, Lazaropole village, Tresonce village 

(Sterijovski et al. 2003; Jelić et al. 2012); Bitola Region: Mogila 

village (Jelić et al. 2012); Debar Region: Debar (Jelić et al. 2012); 

Demir Kapija Region: Demir Kapija (Kavadarci) (Doflein 1921, Jelić 

et al. 2012); Dešat Mt.: Bituše village (Jelić et al. 2012); Galičica Mt.: 

Elšani village, Trpejca village (Dimovski 1959a, 1959b, 1960, 1963, 

1966a,b, 1971, 1981; Jelić et al. 2012); Čokolsko, Gradište auto camp 

Konjsko village (Ohrid side), Leskoec village, Peštani village, 

Pljuska, Prisoj, Sir Han, Suvo Pole, Volkolegalo, Zli Dol (Jelić et al. 

2012); German Mt. (Kriva Palanka) (Jelić et al. 2012); Gevgelija 

Region: Kovanci village, Miravci village (Jelić et al. 2012); Gradeška 

planina Mt.: Konče (Jelić et al. 2012); Gradsko Region: Nogaevci 

village, Ulanci village (Dimovski 1959a, 1959b, 1960, 1963, 1966a,b, 

1971, 1981); Ilinska Planina Mt.: Golemo Ilino (Jelić et al. 2012); 

Jablanica Mt.: Kališta, Krstec, Mala Krasta, Modrič village 

(Sterijovski et al. 2010a; Jelić et al. 2012); Karaorman Mt.: 

Tašmaruništa village (Jelić et al. 2012); Kavadarci Region: Dabnište 

village, Krnjevo village, Rosoman village (Veles) (Jelić et al. 2012); 

Kičevo Region: Kičevo, Izvor village (Jelić et al. 2012); Kožuf Mt.: 

Citate peak, Dren village (Marjanska Planina Mt.), Konopište 

village (Jelić et al. 2012); Kratovo Region: Stracin village (Jelić et al. 

2012); Kožuf Mt.: Čavdarka, Huma village, Konopište village, 

Konjsko village, Mihajlovo (Sterijovski et al. 2010b); Kruševo 

Region: Dolno Divjaci village (Jelić et al. 2012); Kumanovo Region: 

Pčinja River, Ramno village, Strnovac village (Jelić et al. 2012); Lake 

Dojran Region: Dojran (Karaman 1928, 1931, 1937, 1939); Čaušli 

village (Jelić et al. 2012); Lake Prespa Region: Carina village, Golem 

Grad Island, Oteševo village, Stenje village (Dimovski 1959a, 1959b, 

1960, 1963, 1966a,b, 1971, 1981); Konjsko village (Jelić et al. 2012); 

Makedonski Brod Region: Slatina village (Jelić et al. 2012); 

Maleševski Planini Mts.: Lake Berovsko Ezero – near (Jelić et al. 

2012); Mariovo Region: Dunje village, Staravina village, Suvodol, 

Vitolište village (Jelić et al. 2012); Negotino Region: Dubrovo 

village (Jelić et al. 2012); Nidže Mt.: Skočivir village (Jelić et al. 

2012); Ogražden Mt.: Ogražden mine (Jelić et al. 2012); Osogovski 

Planini Mts.: Dobrevo village (Probištip), Gradče (Kočani), Jamiška 

reka River (Kratovo), Jamište village (Kratovo), Jastrebnik village 

(Kočani), Kriva Jabuka (Kratovo), Kunovo village (Kratovo), Luti 

Rid (Kriva Palanka), Orizari village (Istibanja), Popova Čuka hill 

(Istibanja), Ramni Nivi (Kratovo), Rusov Grob (Istibanja), Sinkovica 

(Kočani), Šlegovo village (Kratovo), Sokolarci village (Kočani), 

Sokolovci village (Istibanja), Štalkovica village (Zletovo), Tursko 

Rudare village (Zletovo), Varovište village (Kriva Palanka) (Jelić et 

al. 2012); Prilep Region: Sivec – above (Jelić et al. 2012); Pelister Mt.: 

Brajčino village (Jelić et al. 2012); Plakenska Planina Mt.: Babino 

village, Brežanska reka River, Virovo village (Jelić et al. 2012); 

Probištip Region: Kundino village (Jelić et al. 2012); Šar Planina Mt.: 

Tetovo (Džukić 1972); Selce Keč, Tetovo – above (Jelić et al. 2012); 

Katlanovo Region: railroad station Pčinja, railroad station Rajko 

Žinzifov (Dimovski 1959a, 1959b, 1960, 1963, 1966a,b, 1971, 1981); 

Skopje Region: Skopje (Karaman 1928, 1931, 1937, 1939; Džukić 

1972); Treska River gorge, Matka, Vodno Mt. (Dimovski 1959a, 

1959b, 1960, 1963, 1966a,b, 1971, 1981); Tabuška Reka river (Džukić 

1972); Creševo village, Divlje village, Kučkovo village, Orman 

village, Lepenec River (Jelić et al. 2012); Skopska Crna Gora Mt.: 

Blagoveštenja monastery (Karaman 1928, 1931, 1937, 1939; Džukić



1972); Slaviško pole Region: Luda Mogila hill (Stracin), Petralica 

village (Kriva Palanka) (Jelić et al. 2012); Štip Region: Adjibegovo 

village, between Štip and Krivolak, Bogoslovec village, Bregalnica 

River valley, Delisinci village, Ovče Pole (Dimovski 1959a, 1959b, 

1960, 1963, 1966a,b, 1971, 1981); Stogovo Mt.: Osino (Jelić et al. 

2012); Suva Gora Mt.: Zdunje village (Reka) (Jelić et al. 2012); Sveti 

Nikole Region: Gorobinci village (Jelić et al. 2012); Veles Region: 

Veles (Doflein 1921); Bekirlija village (Štip), Lozja, River Babuna 

(Jelić et al. 2012); Žeden Mt.: Grupčin (Tetovo) (Jelić et al. 2012). 

Vipera berus (Linnaeus, 1758) 

Bistra Mt.: Golem Brezovec peak - bellow (Sterijovski et al. 2003; 

Jelić et al. 2012); Dešat Mt.: Lake Gorno Gasevo Ezero, Golema 

stena - bellow (Jelić et al. 2012); Galičica Mt.: Asan Gjura (Dimovski 

1963, 1964, 1966a, 1981); Magaro peak, Tepeno (Jelić et al. 2012); 

Jablanica Mt.: Crven kamen - near, Čumin Vrv peak, Golemo Kule, 

Krstec, Lake Podgorečko Ezero, Mečkina Dupka (Sterijovski et al. 

2010a; Jelić et al. 2012); Kožuf Mt.: Dudica peak, Orlov Vrv peak 

(Sterijovski et al. 2010b); Jakupica Mt.: Golešnica Mt. (Doflein 1921); 

Golemo Kule (Jelić et al. 2012); Solunska Glava peak (Dimovski 

1963, 1964, 1966a, 1981); Korab Mt.: Nistrovo Bačilo (Dimovski 

1963, 1964, 1966a, 1981); Kobilino Pole, Kabaš (Jelić et al. 2012); 

Kožuf Mt.: Dudica peak, Džibarica, Orlov Vrv peak, Porta, Zelen 

Beg (Jelić et al. 2012); Maleševski Planini Mt.: Čengino (Jelić et al. 

2012); Nidže Mt.: Čemerikata (Jelić et al. 2012); Osogovski Planini 

Mt.: without precise locality (Dimovski 1963, 1964, 1966a, 1981); 

Juručki Grobišta, Sultan Tepe peak, Vlaški Kolibi (Jelić et al. 2012); 

Pelister Mt.: without precise locality (Dimovski 1963, 1964, 1966a, 

1981); Bolnici, Crvena reka River, Golemo Stopalo peak, Kunina 

Stena, Lake Malo Ezero, Pelister peak (Jelić et al. 2012); Plakenska 

Planina Mt.: Straža (Jelić et al. 2012); Salakova Mt. (Karaman 1931); 

Šar Planina Mt.: without precise locality (Dimovski 1963, 1964, 

1966a, 1981); Karabunar, Lake Belo Ezero, Lake Karanikolićko 

Ezero, Plat, Popova Šapka, Rasangjule, Vakaf (Jelić et al. 2012). 

Vipera ursinii (Bonaparte, 1835) 

Bistra Mt. (Sterijovski 2005); Korab Mt.: Kobilino Pole (Dimovski 

1964); Šar Planina Mt.: Kobilica Mt. (Doflein 1921; Karaman 1938- 

1939); Dedel Beg, Fudan, Idrizova Dupka, Karabunar, Kobilica, 

Kraište, Kuči Baba, Lake Karanikolićko Ezero, Ljubanski Uši, Mala 

Smreka, Mountain hut "Ljuboten", Popova Šapka, Radika, 

Rasangjule, Rudoka, Selišta, Šija, Vakaf, Venec, Vojvodino, 

Vratnički bačila (Jelić et al. 2012). 

UNCORRECTED PROOF

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