654 Table 1 Veget<strong>at</strong>ion cover <strong>of</strong> <strong>the</strong> plant communities on <strong>marsh</strong>es, comb<strong>in</strong>ed with a table <strong>of</strong> <strong>the</strong> frequency and cover <strong>of</strong> vascular plants. Roman numerals <strong>in</strong>dic<strong>at</strong>e frequency: I1%,+
655 In order to measure electric conductivity, surface w<strong>at</strong>er was sampled ei<strong>the</strong>r <strong>in</strong> <strong>the</strong> root zone <strong>of</strong> plants (Hippuris tetraphylla, Carex mackenzeii, Dupontia psilosantha communities) or <strong>in</strong> pools th<strong>at</strong> were present <strong>in</strong> <strong>the</strong> <strong>marsh</strong> (all o<strong>the</strong>r plant communities; sample sizes are mentioned <strong>in</strong> Table 1). Soil samples were collected <strong>in</strong> <strong>the</strong> root<strong>in</strong>g zone <strong>of</strong> plants (0–8 cm), dried to a constant weight <strong>at</strong> 70°C and homogenised us<strong>in</strong>g a porcela<strong>in</strong> mortar and pistil and sieved with a mesh width <strong>of</strong> 0.25 mm. Every plant community was sampled once on each transect where present (Table 2). The total nitrogen and carbon contents (% dry m<strong>at</strong>ter) were determ<strong>in</strong>ed by gas chrom<strong>at</strong>ography on an autom<strong>at</strong>ed nitrogen/carbon analyser (ANA-1500) <strong>at</strong> <strong>the</strong> Ecoanalit Labor<strong>at</strong>ory (Institute <strong>of</strong> Biology, Komi Science Centre, Russia). To get <strong>in</strong>form<strong>at</strong>ion about <strong>the</strong> <strong>use</strong> <strong>of</strong> <strong>the</strong> area by o<strong>the</strong>r herbivores, we collected d<strong>at</strong>a on <strong>the</strong> presence <strong>of</strong> o<strong>the</strong>r herbivorous birds and mammals. Sight<strong>in</strong>gs and counts <strong>of</strong> birds and mammals were not collected system<strong>at</strong>ically, but were noted. Dur<strong>in</strong>g 10 days, three l<strong>in</strong>es <strong>of</strong> 20 mo<strong>use</strong>traps, baited with carrot and peanut butter, were set <strong>in</strong> <strong>the</strong> proximity <strong>of</strong> <strong>the</strong> village and transect 1 <strong>in</strong> order to trap small mammals. The taxonomy <strong>of</strong> bird species follows <strong>the</strong> guidel<strong>in</strong>es <strong>of</strong> <strong>the</strong> Dutch Commission for Avian System<strong>at</strong>ics (CSNA). St<strong>at</strong>istics As soil and veget<strong>at</strong>ion properties were measured <strong>in</strong> different plots with different sample sizes, <strong>the</strong>se d<strong>at</strong>a are not appropri<strong>at</strong>e for detailed st<strong>at</strong>istical analyses. Hence, <strong>the</strong> results are presented <strong>in</strong> a descriptive manner. All analyses were performed us<strong>in</strong>g <strong>the</strong> st<strong>at</strong>istical package SPSS for W<strong>in</strong>dows ver.11.0.1. Barnacle goose dropp<strong>in</strong>g density was log-transformed to reach normality criteria. Results Classific<strong>at</strong>ion <strong>of</strong> <strong>the</strong> veget<strong>at</strong>ion On basis <strong>of</strong> <strong>the</strong> veget<strong>at</strong>ion classific<strong>at</strong>ion, 11 plant communities were recognised with<strong>in</strong> <strong>the</strong> <strong>salt</strong> <strong>marsh</strong> (Table 1). Three plant communities not presented <strong>in</strong> Table 1 are <strong>the</strong> D. psilosantha community (a community with a dom<strong>in</strong>ance <strong>of</strong> D. psilosantha), a community similar to <strong>the</strong> C. subsp<strong>at</strong>hacea/dicotyledons community with a dom<strong>in</strong>ance <strong>of</strong> Plantago schrenkii and a community th<strong>at</strong> we call ‘‘beach wall <strong>in</strong> colony’’. The l<strong>at</strong>ter was situ<strong>at</strong>ed on an elev<strong>at</strong>ed sandy ridge <strong>at</strong> <strong>the</strong> coast and was characterised by a low veget<strong>at</strong>ion cover (10%), <strong>of</strong> which 77% <strong>of</strong> <strong>the</strong> cover was gram<strong>in</strong>oids (36% Pucc<strong>in</strong>ellia phryganodes, 41% C. subsp<strong>at</strong>hacea). The rema<strong>in</strong>der was P. schrenkii (23%); and many nests <strong>of</strong> <strong>barnacle</strong> <strong>geese</strong> were found <strong>in</strong> this community. Mosses and lichens are not presented <strong>in</strong> Table 1, although <strong>the</strong>y were recorded as Table 2 Soil properties (N:C r<strong>at</strong>io <strong>of</strong> sediment, fraction nitrogen <strong>in</strong> sediment, electrical conductivity <strong>of</strong> <strong>the</strong> groundw<strong>at</strong>er) and veget<strong>at</strong>ion properties (total aboveground biomass, tiller density, veget<strong>at</strong>ion height) <strong>of</strong> all <strong>salt</strong>-<strong>marsh</strong> plant communities. D<strong>at</strong>a given are means ±SE, with sample size given <strong>in</strong> brackets. For tiller density and canopy height, <strong>the</strong> results <strong>of</strong> a posthoc test are given. Different letters denote significant differences <strong>at</strong> <strong>the</strong> P