Lay Lake - Alabama Department of Conservation and Natural ...
Lay Lake - Alabama Department of Conservation and Natural ...
Lay Lake - Alabama Department of Conservation and Natural ...
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<strong>Lay</strong> <strong>Lake</strong><br />
Management Report<br />
Spring 2011<br />
Prepared by<br />
Michael P. Holley<br />
District Fisheries Biologist<br />
E. Dan Catchings<br />
District Fisheries Supervisor<br />
Kevin Baswell<br />
Biologist Aide<br />
<strong>Alabama</strong> <strong>Department</strong> <strong>of</strong> <strong>Conservation</strong><br />
<strong>and</strong> <strong>Natural</strong> Resources<br />
Division <strong>of</strong> Wildlife <strong>and</strong><br />
Freshwater Fisheries<br />
September 1, 2011
Introduction<br />
The objective <strong>of</strong> the <strong>Lay</strong> <strong>Lake</strong> reservoir management plan is to collect quantitative<br />
biological data to assist district biologists in developing management strategies to enhance the<br />
fishery. The reservoir is sampled periodically to evaluate population metrics <strong>of</strong> major sport fish<br />
species. A summary <strong>of</strong> the morphometric, physical, <strong>and</strong> chemical characteristics <strong>of</strong> <strong>Lay</strong> <strong>Lake</strong> is<br />
listed in Table 1 <strong>of</strong> this report. Readers are encouraged to utilize the tables <strong>and</strong> figures located in<br />
the appendix <strong>of</strong> the report. An angler’s guide to fish management reports is available on the<br />
Outdoor <strong>Alabama</strong> website at the link below.<br />
http://www.outdooralabama.com/fishing/freshwater/where/reservoirs/guide.pdf.<br />
Methods<br />
Largemouth bass <strong>and</strong> <strong>Alabama</strong> bass were targeted during electr<strong>of</strong>ishing surveys on April<br />
12, 2011 <strong>and</strong> April 13, 2011 at eight r<strong>and</strong>omly selected sites (Figures 1-2). Target species<br />
(largemouth bass <strong>and</strong> <strong>Alabama</strong> bass) were measured to the nearest millimeter, <strong>and</strong> weighed to<br />
the nearest gram. Otoliths were extracted to obtain age, growth <strong>and</strong> mortality estimates, <strong>and</strong><br />
were read whole view by District 2 personnel. Suspect ages, or otoliths that were not readable in<br />
whole view prompted sectioning (Maceina 1988) to authenticate age. Fish collection <strong>and</strong> general<br />
data analyses were conducted according to the guidelines <strong>of</strong> the <strong>Alabama</strong> Reservoir Management<br />
Manual (Cook 1999). Simulation modeling was conducted using FAST s<strong>of</strong>tware to determine<br />
whether a 15 inch minimum length limit could improve the largemouth bass fishery.<br />
Results <strong>and</strong> Discussion<br />
Largemouth Bass<br />
A total <strong>of</strong> 308 largemouth bass (Tables 2- 3) were collected at a rate <strong>of</strong> 77.0/hr during<br />
electr<strong>of</strong>ishing surveys, which is similar to the lake average (77.7/hr) catch rate (Table 2). Catch<br />
rates improved for most size groups <strong>of</strong> largemouth bass from 2008 to 2011. The only notable<br />
exception was preferred size fish, which had below average catch rates (Table 2).<br />
The PSD for largemouth was 67, <strong>and</strong> relative stock density indices exceeded the lake<br />
average for stock (RSD S-Q) <strong>and</strong> quality (RSD Q-P) size groups (Figure 4). Abundance <strong>of</strong><br />
preferred (RSD P-M) <strong>and</strong> memorable (RSD M-T) largemouth bass fell below the lake average<br />
2
(Figure 4). RSD S-Q <strong>and</strong> RSD M-T values were below the statewide average for <strong>Alabama</strong><br />
reservoirs. RSD Q-P <strong>and</strong> RSD P-M abundance approached the upper 75 th percentile for <strong>Alabama</strong><br />
reservoirs (Figure 4).<br />
Largemouth bass ranged from 162 mm to 556 mm total length, <strong>and</strong> from 1 to 9 years old<br />
(Table 3; Figures 3 <strong>and</strong> 5). The von Bertalanffy growth curve predicted that it took about 1.4<br />
years for largemouth bass to reach a harvestable size <strong>of</strong> 10 inches, <strong>and</strong> about 2.0 years to reach<br />
12 inches (Figure 5). Growth has remained relatively stable at <strong>Lay</strong> <strong>Lake</strong> over the last 5 samples<br />
(Figure 6). Relative weight was excellent, <strong>and</strong> exceeded the lake average for most size groups <strong>of</strong><br />
largemouth bass; only memorable size largemouth fell below the lake average (Table 2).<br />
Total annual mortality for age-2 to age-9 largemouth bass from catch-curve analysis was<br />
43% (R 2 =0.96; P
Simulation Modeling - Largemouth Bass<br />
Total annual mortality from catch-curve analysis, suggested that fishing mortality may be<br />
moderately high for largemouth bass. In order to determine whether the largemouth bass fishery<br />
could be improved with a minimum length limit, simulation modeling was conducted to compare<br />
the current fishery (where anglers generally only keep largemouth bass 12 inches <strong>and</strong> larger) to<br />
the predicted benefits <strong>of</strong> a 15 inch minimum length limit. At our estimated natural mortality rate<br />
<strong>of</strong> 0.36, the calculated exploitation rate was 16%, which is only moderate for most black bass<br />
populations, but generally higher than reported from recent creel surveys. If a minimum length<br />
limit <strong>of</strong> 15 inches was implemented, our model predicted that about 35% more memorable size<br />
(20 inches) largemouth bass would be available to anglers (Figure 12). However, the model also<br />
predicted that about 39% fewer bass would be available for anglers to keep or weigh-in a<br />
tournament (Figure 13). Mean total length <strong>of</strong> fish in the population was predicted to increase<br />
from 391 mm to 435 mm (just over 15 inches to approximately 17 inches). The yield <strong>of</strong><br />
largemouth bass would only decrease about 13% if a 15 inch minimum length limit was<br />
implemented, since fewer, but bigger bass would be available to keep or weigh-in.<br />
Summary<br />
The black bass populations in <strong>Lay</strong> <strong>Lake</strong> display good growth rates, <strong>and</strong> excellent relative<br />
weights. Preferred <strong>and</strong> memorable size largemouth bass were slightly less abundant than the<br />
lake average, however, stock <strong>and</strong> quality size largemouth bass were abundant <strong>and</strong> should recruit<br />
to larger sizes in forthcoming years. Year-class strength <strong>of</strong> largemouth bass has been relatively<br />
constant since 2002. <strong>Alabama</strong> bass catch rates were high, <strong>and</strong> several year-classes were<br />
represented with constant recruitment since 2004. Moderately high total annual mortality<br />
prompted the evaluation <strong>of</strong> minimum length limits to possibly improve the largemouth bass<br />
fishery. Although a 15 inch minimum length limit could improve size structure, a substantial<br />
decline in number <strong>of</strong> bass available to anglers was predicted. The current number <strong>of</strong> memorable<br />
size largemouth bass was low at <strong>Lay</strong> <strong>Lake</strong>, <strong>and</strong> if this trend continues, a minimum length limit,<br />
such as 15 inches may be beneficial in improving numbers <strong>of</strong> big largemouth bass. Total annual<br />
mortality estimates suggested that, even though catch <strong>and</strong> release may be high from creel<br />
4
surveys, fishing mortality may be increasing through hooking mortality <strong>and</strong> tournament mortality<br />
for both largemouth bass <strong>and</strong> <strong>Alabama</strong> bass.<br />
Conclusions<br />
1. Re-sample the black bass populations in a timely manner, to follow trends in size structure,<br />
age <strong>and</strong> growth, <strong>and</strong> mortality <strong>of</strong> black bass populations.<br />
2. No changes to the current black bass regulations should be implemented at this time.<br />
5
Literature Cited<br />
Cook, S.F. 1999. <strong>Alabama</strong> reservoir management manual. <strong>Alabama</strong> <strong>Department</strong> <strong>of</strong><br />
<strong>Conservation</strong> <strong>and</strong> <strong>Natural</strong> Resources. Montgomery, AL.<br />
Anderson, R.O., <strong>and</strong> R.M. Neumann. 1996. Length, weight, <strong>and</strong> associated structural indices.<br />
Pages 447-482 in B.R. Murphy <strong>and</strong> D.W. Willis, editors. Fisheries Techniques. 2 nd edition.<br />
American Fisheries society, Bethesda, Maryl<strong>and</strong>.<br />
Haffner, J. B., J. L. Moss, <strong>and</strong> J. M. Piper. 2008. Management report for <strong>Lay</strong> <strong>Lake</strong>. <strong>Alabama</strong><br />
<strong>Department</strong> <strong>of</strong> <strong>Conservation</strong> <strong>and</strong> <strong>Natural</strong> Resources. Montgomery, AL.<br />
Jenkins, R. M. 1967. The influence <strong>of</strong> some environmental factors on the st<strong>and</strong>ing crop <strong>and</strong><br />
harvest <strong>of</strong> fishes in U.S. reservoirs. Pages 298-321 in Reservoir Fishery Resources<br />
Symposium. Southern Division American Fisheries Society, Bethesda, Maryl<strong>and</strong>, USA.<br />
Maceina, M.J. 1988. A simple grinding procedure for sectioning otoliths. North American<br />
Journal <strong>of</strong> Fisheries Management 8:141-143.<br />
Ryder, R. A. 1965. A method for estimating the potential fish production <strong>of</strong> north-temperate<br />
lakes. Transactions <strong>of</strong> the American Fisheries Society. 94:213-218.<br />
Welch , P. S. 1948. Limnological Methods. McGraw-Hill. pp. 93-94.<br />
6
Appendix 1<br />
Tables <strong>and</strong> Figures<br />
7
Table 1. Morphometric, physical <strong>and</strong> chemical characteristics <strong>of</strong> <strong>Lay</strong> <strong>Lake</strong>.<br />
Surface area<br />
12,000 acres<br />
Drainage area<br />
9,087 sq. mi.<br />
Full pool elevation<br />
396 feet-msl<br />
Mean annual fluctuation<br />
0.5 feet<br />
Shoreline distance<br />
289 miles<br />
Shoreline development index 19 (Welch 1948)<br />
Mean depth<br />
22 feet<br />
Maximum depth<br />
88 feet<br />
Outlet depth (upper) 362 feet<br />
(lower) 342 feet<br />
Thermocline depth None<br />
Stratification index 10.16<br />
Total dissolved solids<br />
118 mg/l<br />
Morphoedaphic index<br />
5.4 TDS/mean depth(ft)<br />
(Ryder 1965)<br />
Growing season<br />
210 frost free days<br />
(Jenkins 1967)<br />
Year impounded 1914<br />
8
Table 2. Relative stock density, catch per effort, <strong>and</strong> relative weight <strong>of</strong> black basses from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
NO. OF TOTAL SUBSTOCK RSD S-Q RSD Q-P RSD P-M RSD M-T TOTAL<br />
SPECIES YEAR SAMPLES EFFORT NO. CPE SSR NO. CPE PCT. Wr NO. CPE PCT. Wr NO. CPE PCT. Wr NO. CPE PCT. Wr NO. CPE<br />
LARGEMOUTH BASS 2011 8 4.00 17 4.2 6 96 24.0 33 103 113 28.3 39 99 76 19.0 26 99 6 1.5 2 94 308 77.0<br />
LARGEMOUTH BASS 2008 4 2.02 9 4.5 7 26 12.9 20 96 44 21.8 34 97 59 29.2 45 91 2 0.9 2 90 140 69.3<br />
LARGEMOUTH BASS 2004 3 1.51 24 15.9 24 35 23.1 35 93 31 20.5 31 93 33 21.8 33 94 1 0.6 1 93 124 81.9<br />
LARGEMOUTH BASS 2001 3 1.11 20 18.0 18 38 34.2 33 94 33 29.7 29 97 38 34.3 33 99 5 4.5 4 106 134 120.7<br />
LARGEMOUTH BASS 1997 4 1.68 4 2.4 4 28 16.6 27 94 49 29.1 48 94 23 13.7 23 93 2 1.2 2 94 106 63.0<br />
LARGEMOUTH BASS 1992 4 1.80 7 3.9 7 14 7.8 13 94 49 27.2 47 99 37 20.6 36 105 4 2.2 4 105 111 61.7<br />
LARGEMOUTH BASS 1987 3 2.00 38 19.0 37 20 10.0 19 39 19.5 38 38 19.0 37 6 3.0 6 141 70.5<br />
LAKE AVERAGE 9.7 15 37 18.4 26 96 51 25.2 38 97 43 22.5 33 97 4 2.0 3 97 152.0 77.7<br />
ALABAMA BASS 2011 8 4.00 0 0.0 0 20 5.0 16 111 39 9.8 31 108 58 14.5 46 109 10 2.5 8 109 127 31.8<br />
ALABAMA BASS 2008 8 4.05 0 0.0 0 8 2.0 6 109 42 10.4 29 111 77 19.0 53 110 17 4.2 12 103 144 35.6<br />
ALABAMA BASS 2004 10 5.03 5 1.0 6 9 1.8 11 99 27 5.4 33 97 31 6.2 37 100 16 3.2 19 97 88 17.5<br />
ALABAMA BASS 2001 6 2.88 8 2.8 6 27 9.4 21 102 32 11.1 25 103 48 16.7 38 103 19 6.6 15 108 134 46.5<br />
ALABAMA BASS 1997 8 3.69 3 0.8 3 6 1.6 6 103 31 8.4 29 105 47 12.7 44 104 23 6.2 21 107 110 29.8<br />
ALABAMA BASS 1992 10 5.00 0 0.0 0 13 2.6 23 121 18 3.6 32 107 18 3.6 32 109 8 1.6 14 109 57 11.4<br />
ALABAMA BASS 1987 10 5.00 4 0.8 4 27 5.4 28 33 6.6 34 27 5.4 28 9 1.8 9 100 20.0<br />
LAKE AVERAGE 0.8 3 16 4.0 16 108 32 7.9 30 105 44 11.2 40 106 15 3.7 14 106 109 27.5<br />
9
Table 3. Number-at-age, catch-at-age, <strong>and</strong> mean total length <strong>of</strong> largemouth bass collected from<br />
<strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
Age Year Class Number Percent CPE Mean TL SE Range<br />
1 2010 69 22.4 17.3 224 3.3 162-286<br />
2 2009 93 30.2 23.3 298 2.8 226-377<br />
3 2008 81 26.3 20.3 357 4.2 223-440<br />
4 2007 31 10.1 7.8 414 6.1 340-490<br />
5 2006 16 5.2 4.0 421 13.1 350-493<br />
6 2005 8 2.6 2.0 479 15.1 393-531<br />
7 2004 4 1.3 1.0 505 28.8 424-547<br />
8 2003 3 1.0 0.8 500 6.2 493-512<br />
9 2002 3 1.0 0.8 487 34.7 449-556<br />
Total 308 100.0 77.0<br />
10
Table 4. Number-at-age, catch-at-age, <strong>and</strong> mean total length <strong>of</strong> <strong>Alabama</strong> bass collected from<br />
<strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
Age Year Class Number Percent CPE Mean TL SE Range<br />
1 2010 4 3.1 1.0 200.0 10.6 183-231<br />
2 2009 20 15.7 5.0 271.2 5.1 243-321<br />
3 2008 51 40.2 12.8 344.5 3.5 272-402<br />
4 2007 27 21.3 6.8 388.6 5.9 278-441<br />
5 2006 11 8.7 2.8 406.6 6.0 372-428<br />
6 2005 9 7.1 2.3 417.3 11.4 348-453<br />
7 2004 4 3.1 1.0 449.3 11.4 419-470<br />
8 2003 1 0.8 0.3 438.0<br />
Total 127 100.0 31.8<br />
11
Figure 1. <strong>Lay</strong> <strong>Lake</strong> sampling locations north <strong>of</strong> “the narrows”, Spring 2011.<br />
12
Figure 2. <strong>Lay</strong> <strong>Lake</strong> sampling locations south <strong>of</strong> “the narrows”, Spring 2011.<br />
13
Number<br />
60<br />
Age - 9<br />
50<br />
40<br />
30<br />
20<br />
Age - 8<br />
Age - 7<br />
Age - 6<br />
Age - 5<br />
Age - 4<br />
Age - 3<br />
Age - 2<br />
Age - 1<br />
10<br />
0<br />
75 100 125 150 175 200 225 250 275 300 325 350 375 400 425 450 475 500 525 550<br />
Length Group (mm)<br />
Figure 3. Age-length frequency for largemouth bass collected from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
14
RSD (%)<br />
60<br />
50<br />
40<br />
<strong>Lake</strong> Average<br />
30<br />
2011<br />
20<br />
Statewide average<br />
10<br />
0<br />
S-Q Q-P P-M M-T Trophy<br />
Incremental RSD Categories<br />
Figure 4. Relative stock density <strong>of</strong> largemouth bass collected from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
Orange bars are the 2011 sample, red bars are the <strong>Lay</strong> <strong>Lake</strong> average, <strong>and</strong> yellow<br />
diamonds are the statewide means.<br />
15
Length(mm)<br />
600<br />
500<br />
400<br />
300<br />
200<br />
Linf = 534.552<br />
K = 0.302<br />
to = -0.760<br />
100<br />
0<br />
0 1 2 3 4 5 6 7 8 9<br />
Age<br />
Figure 5. Von Bertalanffy growth curve for largemouth bass collected from <strong>Lay</strong> <strong>Lake</strong>,<br />
Spring 2011.<br />
16
Mean Total Length<br />
550<br />
525<br />
500<br />
475<br />
450<br />
425<br />
400<br />
375<br />
350<br />
325<br />
300<br />
275<br />
250<br />
2011<br />
<strong>Lay</strong> <strong>Lake</strong> Average<br />
225<br />
200<br />
175<br />
150<br />
0 1 2 3 4 5 6 7 8 9 10 11<br />
AGE<br />
Figure 6. Mean lengths-at-age for largemouth bass collected from <strong>Lay</strong> <strong>Lake</strong>.<br />
17
Number<br />
100<br />
Z = -0.5637<br />
S = 57% (95% CI 51 - 64)<br />
r 2 = 0.9580<br />
10<br />
1<br />
2 3 4 5 6 7 8 9<br />
Age<br />
Figure 7. Catch-curve for largemouth bass, ages 2-9, collected from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
18
Number<br />
30<br />
25<br />
20<br />
15<br />
10<br />
Age - 8<br />
Age - 7<br />
Age - 6<br />
Age - 5<br />
Age - 4<br />
Age - 3<br />
Age - 2<br />
Age - 1<br />
5<br />
0<br />
175 200 225 250 275 300 325 350 375 400 425 450<br />
Length Group (mm)<br />
Figure 8. Age-length frequency for <strong>Alabama</strong> bass collected from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
19
RSD (%)<br />
70<br />
60<br />
50<br />
40<br />
<strong>Lake</strong> Average<br />
2011<br />
Statewide average<br />
30<br />
20<br />
10<br />
0<br />
S-Q Q-P P-M M-T Trophy<br />
Incremental RSD Catergories<br />
Figure 9. Relative stock density <strong>of</strong> <strong>Alabama</strong> bass collected from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
Orange bars are the 2011 sample, red bars are the <strong>Lay</strong> <strong>Lake</strong> average, <strong>and</strong> yellow<br />
diamonds are the statewide means.<br />
20
Length(mm)<br />
500<br />
450<br />
400<br />
350<br />
300<br />
250<br />
200<br />
150<br />
100<br />
Linf = 462.497<br />
K = 0.394<br />
to = -0.397<br />
50<br />
0<br />
1 2 3 4 5 6 7 8<br />
Age<br />
Figure 10. Von Bertalanffy growth curve for <strong>Alabama</strong> bass collected from <strong>Lay</strong> <strong>Lake</strong>, Spring<br />
2011.<br />
21
Number<br />
100<br />
Z = -0.7311<br />
S = 48% (39-59)<br />
r 2 = 0.9602<br />
10<br />
1<br />
3 4 5 6 7 8<br />
Age<br />
Figure 11. Catch-curve for <strong>Alabama</strong> bass, ages 3-8, collected from <strong>Lay</strong> <strong>Lake</strong>, Spring 2011.<br />
22
Number at Memorable Size<br />
30<br />
25<br />
20<br />
15<br />
10<br />
12" 305 mm<br />
15" 381 mm<br />
5<br />
0<br />
0 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8<br />
Exploitation<br />
Figure 12. The predicted number <strong>of</strong> largemouth bass reaching memorable size in <strong>Lay</strong> <strong>Lake</strong> over<br />
a range <strong>of</strong> exploitation rates <strong>and</strong> minimum length limits. The vertical light-blue line<br />
represents the predicted current fishery.<br />
23
Number Available<br />
450<br />
400<br />
350<br />
300<br />
250<br />
200<br />
150<br />
100<br />
50<br />
0<br />
12' 305 mm<br />
15" 381 mm<br />
0 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8<br />
Exploitation<br />
Figure 13. The predicted number <strong>of</strong> largemouth bass available to anglers in <strong>Lay</strong> <strong>Lake</strong> over<br />
a range <strong>of</strong> exploitation rates at different minimum length limits. The vertical light<br />
blue line represents the predicted current fishery.<br />
24