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Checklist of Butterflies (Papilionoidea) of the Mongolian Altai ...

Checklist of Butterflies (Papilionoidea) of the Mongolian Altai ...

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82 YAKOVLEV: <strong>Butterflies</strong> <strong>of</strong> <strong>the</strong> <strong>Mongolian</strong> <strong>Altai</strong><br />

Khasgt-Khirkhan Mountains E. chastilovi is a bona fide species and its status as a species<br />

is based on <strong>the</strong> following characters: clear external distinctions (detailed in <strong>the</strong> original<br />

description) and some distinctions in genitalia morphology (valve is strongly curved,<br />

with a specific process distally forming an almost flat area with small denticles on it,<br />

and a larger denticle at <strong>the</strong> base <strong>of</strong> <strong>the</strong> process; <strong>the</strong> antevaginal plate is U-shaped, with<br />

ra<strong>the</strong>r massive lateral processes, as opposed to <strong>the</strong> much smaller structures in altajana<br />

Staudinger, 1901). The new subspecies from <strong>the</strong> nor<strong>the</strong>rn part <strong>of</strong> <strong>the</strong> distribution area<br />

(Khovd aimak, Ikh-Nuruu Mountains) is described and differentiated based on wing<br />

pattern elements, smaller size, and slight differences in female genitalia morphology.<br />

Erebia chastilovi nomada ssp. n. (Figs 22i – n, 23d, 24i – o, 25)<br />

Material. Holotype P, W Mongolia, Khovd aimak, Ikh-Nuruu Mountains (between Bodonchin-Gol<br />

and Uenchin-Gol valleys), Ikh-Daba pass, 2900m, 5.vii.2007, leg. R. Yakovlev (ZISP). – Paratypes: 11<br />

PP, 4 OO, same data (ZISP, RYB).<br />

Diagnosis. The new subspecies differs from <strong>the</strong> nominate in its smaller size (<strong>the</strong> length<br />

<strong>of</strong> <strong>the</strong> forewing is 15 – 16.5 mm, and 16 – 18.5 mm in <strong>the</strong> nominate subspecies). The<br />

wing upperside in males has a better defined ochreous pattern, forming a wide area in<br />

<strong>the</strong> forewing and consisting <strong>of</strong> small submarginal dashes in <strong>the</strong> hindwing. The hindwing<br />

underside is more brightly coloured, with a very well-defined marmorate pattern. The<br />

male genitalia have no obvious distinctions. The female genitalia bear signa, strongly<br />

converging distally on <strong>the</strong> corpus bursae.<br />

Habitat. Very dry alpian tundra-steppe.<br />

Etymology. The subspecies name comes from <strong>the</strong> word “nomad”.<br />

Erebia pandrose (Borkhausen, 1788)<br />

Tsagan-Kol Lake, rare and local.<br />

Hyponephele lycaon (Rottemburg, 1775) (Figs 26 – 27)<br />

This species is very variable in Central Asia. Ssp. catalampra (Staudinger, 1895), described<br />

from <strong>the</strong> Central Khangai Mountains, has been traditionally considered present<br />

in Mongolia and is listed in <strong>the</strong> recent book with all western and central <strong>Mongolian</strong><br />

populations (Tschikolovets et al. 2009). However, populations from different areas<br />

<strong>of</strong> <strong>the</strong> <strong>Mongolian</strong> <strong>Altai</strong> are very variable externally and represent very well-delineated<br />

taxa, whose rank needs refinement. In order to refine <strong>the</strong> real rank considerable<br />

additional material and molecular-genetic studies are required. Here taxa from <strong>the</strong><br />

<strong>Mongolian</strong> <strong>Altai</strong> are considered as subspecies. Probably such a pr<strong>of</strong>usion <strong>of</strong> phenotypes<br />

is caused by <strong>the</strong> complete isolation <strong>of</strong> river valleys in <strong>the</strong> south <strong>of</strong> <strong>the</strong> <strong>Mongolian</strong><br />

<strong>Altai</strong>, thus making <strong>the</strong> genetic exchange between populations <strong>of</strong> Hyponephele lycaon<br />

impossible. The extreme west <strong>of</strong> <strong>the</strong> sou<strong>the</strong>rn macroslope <strong>of</strong> <strong>the</strong> <strong>Mongolian</strong> <strong>Altai</strong> is<br />

inhabited with ssp. smirnovi Yakovlev, 2004 stat. n. (LT: Arshantyn-Nuruu Mountains)<br />

(Figs 26a, b). Similar topotypical butterflies were collected in <strong>the</strong> valley <strong>of</strong> Ulyastain-<br />

Gol River. Eastwards in <strong>the</strong> Mogoin-Gol river valley and <strong>the</strong> Hara-Adzragyn-Nuru<br />

Mountains a small series <strong>of</strong> specimens was collected, different from all <strong>the</strong> known<br />

forms <strong>of</strong> Hyponephele lycaon, and it is described here as a new subspecies.

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