THE GENUS SCHISTIDIUM (GRIMMIACEAE, MUSCI) IN HUNGARY ...

THE GENUS SCHISTIDIUM IN HUNGARY 39
Costa and leaf margins smooth. Lamina smooth, in lower part unistratose, in upper part
with bistratose spots. Upper lamina cells rounded, 6–9 μm wide, lower cells wider and
slightly sinuose, basal marginal cells sometimes subhyaline, with slightly thickened
cross-walls. K+ red.
Sporophytes frequent, mostly shallowly immersed. Urn orange to reddish-brown, often
finely striolate, widest below middle, length/width ratio (1.3–)1.4–1.6–1.8 (BLOM
1996), in Hungarian material (0.94–)1.34 ±0.04(–1.93), n = 91. Exothecial cells elongate,
especially in basal part, at least some up to 50 μm (often up to 80 μm) long, arranged in a
regular pattern. Stomata absent. Peristome teeth 220–340 μm, orange, in central and
upper parts mostly strongly perforated to cribrose.
Variability: The exothecial cells in the lower part of the urn sometimes only reach
(50–)55 μm, but in other samples they attain 75–80 μm.
The brown, dense cushions with hair-points only around the sporophytes give this
plant a very characteristic appearance. However, some forms of S. crassipilum can look
quite similar, and in that case the strongly elongate exothecial cells (Fig. 5G, H, compare
with Fig. 13I, J), and the shorter and wider urn of S. brunnescens are diagnostic. Confusion
could also occur with forms of S. helveticum lacking the typical black colour, but again the
pattern of exothecial cells is quite different. For the difference between subsp. brunnescens
and subsp. griseum see the note under the latter.
: S. brunnescens subsp. brunnescens grows nearly exclusively on exposed calcareous
rocks (limestone, dolomite), and is a very typical member of the bryophyte vegetation in
calcareous grasslands. It was found associated with the lichen Fulgensia fulgens (det. V. Otte).
: Bryum argenteum, Didymodon rigidulus, D. vinealis, Grimmia
dissimulata, G. orbicularis, G. pulvinata, Grimmia tergestina, Pseudocrossidium revolutum,
Pseudoleskeella catenulata, Schistidium crassipilum, S. helveticum, Tortella inclinata, Tortula
crinita, Trichostomum sp.
: 150–780 m a.s.l.
: : Comit. Gömör. In rupibus
calc. montis “Baradla-tetõ” prope pag. Aggtelek, 350 m, 03.06.1928, leg. Á. Boros, BP
111757; : Comit. Borsod. In rupibus calcar. montis Bélkõ prope Bélapátfalva,
500–780 m, 03.09.1959, leg. Á. Boros, BP 110944; Lehmschieferfelsen an der
Straßenböschung zwischen Bükkzsérc und Felsõtárkány, ca 400 m, mit S. crassipilum, 27.03.
2008, leg. P. Erzberger and T. Pócs, herb. Erzberger, (B) 12886; :
Glaukonit-Sandsteinfelsen “Noé szõllõje” im Ort Istenmezeje, 230 m, 26.03.2008, leg. P.
Erzberger and T. Pócs, herb. Erzberger, (B) 12861; : Comit. Pest. In
rupibus dolomit. apricis montis Vár-hegy prope Csõvár, 350 m, 06.05.1953, leg. Á. Boros,
BP 111748; : Comit. Pest. In monte Somlyó prope Fóth, 21.05.1901,
leg. Á. Degen, BP 37650; : Comit. Pest. In rupibus dolomiticis merid.
montis Nagykevély prope pag. Pilisborosjenõ, 500 m, 25.03.1947, leg. Á. Boros, BP 111750;
Hung. centr., com. Pest. In rupibus calcareis supra pag. Pomáz, 31.03.1946, leg. L. Vajda,
EGR; : Budakalász, comit. Pest, in monte “Monalovác” ad saxa calcarea,
250 m, III.1911, leg. J. Szurák, revid. M. Péterfi, BP 5645; : Gyermely,
Studia bot. hung. 39, 2008