2011 - Monte L. Bean Life Science Museum - Brigham Young ...

Monte L. Bean
Life Science Museum
2011 Annual Report
Brigham Young University

Annual Report
January–December 2011
Monte L. Bean
Life Science Museum
Brigham Young University

4 5
Mission Statement
Table of Contents
The Monte L. Bean Life Science Museum at Brigham
Young University is a dynamic repository and trustee for a
remarkable group of biological collections. These collections
are used to celebrate the role of Jesus Christ as Creator, while
enhancing student learning and mentoring and promoting
faculty teaching and research. They also serve as a unique venue
for inviting the public and scientific community to explore and
contemplate intricate biological relationships and processes.
Mission Statement ......................................... 4
Introduction ................................................... 6
Dedication-
Dr. Wilmer W Tanner ....................... 7-8
Donors .......................................................... 17
The Lytle Ranch Preserve ........................... 18
We accomplish our mission by
• Collecting and properly maintaining biological specimens and
associated data to effectively support current and future research
efforts;
• Providing and developing database options to better support
research on the biodiversity and ecological complexities of the
earth’s ecosystems;
• Producing and sharing quality research products in order to
increase scientific knowledge and understanding;
• Facilitating an ongoing dialogue about issues and concerns
related to faith and science as different but complementary ways
of knowing;
• Educating our students and the public about the natural
processes essential to sustaining the biological diversity and
ecological complexity of the earth;
• Providing a forum for educating our students and the public
about best earth stewardship practices;
• Promoting and facilitating quality learning and
mentoring experiences for our students;
• Engaging the public effectively through
compelling exhibits and innovative education
programs in order to promote understanding and
appreciation for the diversity and complexity of the
earth’s biological heritage;
• Using the museum’s resources to develop and implement K-12
science education programs based on the Utah State Core
Curriculum in order to enhance the education of our local public
and private school children while providing powerful, “handson”
training experiences for pre-service primary and secondary
teachers in the School of Education.
Historical Highlight-
D Elden Beck ................................... 8-9
Expansion ............................................ 10
New Exhibits .................................... 10-11
Yellow Perch
Perca flavescens
Museum
Governing
Authority
Cecil O. Samuelson
Brent W. Webb
Alan R. Harker
Rodney J. Brown
Larry L. St. Clair
Leigh A. Johnson
Marta J. Adair
MLBM
Brigham Young University
Provo, Utah 84602
http://mlbean.byu.edu
801-422-5052
Education Programming
Nature Experienceships .......................... 11
Date Night ................................................ 11
Development of an iPad
Dichotomous Key ............................... 12
Bio-Box Initiative Update ........................ 12
Partnership School Districts .................... 12
Live Animal Webcast Initiative .............. 12
CUSEF ........................................................13
Western North American Naturalist .......... 13
Museum Sponsored Lectures and Events
Jon Gelhaus ............................................ 14
Mark Belk ................................................. 14
Screening of “Green Fires” Introduction
by Stanley Temple ............................... 14
Collections-Based Research
Visiting Fulbright Scholar ......................... 15
Studying Fishes Across Latin America ... 15
Research at the Lytle Ranch Preserve ...16
Wildfire and Small Mammals ................. 16
Gambel’s Quail and Kit Fox ................... 16
Cynthia Troxell Lichen
Collection Donated ................................ 16
Visitor Services
Museum Store .......................................... 19
Museum Volunteers ............................... 19
Touch Screen Installed ............................ 19
Museum Newsletter ................................ 19
Museum Staff Awards ............................. 19
Financial Report ........................................... 20
Credits ......................................................... 20
2011 Research Publications .................. 21-27

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Dr. Larry St Clair
Museum Director
Introduction
This has been an extraordinary year. In March, we received
word that the Board of Trustees had granted permission
to begin formal planning for the museum expansion. We
immediately began working with the University’s Physical
Facilities team to initiate planning and to prepare and submit
the documents required to organize the design and construction
team. By May, our application to proceed had been reviewed
and approved, and we sent out a call for proposals to 10
architectural firms. By June, we had narrowed our list of potential
architects to four and had made arrangements to visit with
each of these prospective firms. Later in June, the decision was
made to hire Jacoby Architects from Salt Lake City and Wadman
Construction from Ogden. From the beginning, Jacoby and
Wadman have worked together with the museum’s expansion
steering committee to effectively plan for the museum expansion.
By the end of 2011, Jacoby had the design and development
documents ready for review. The plan is to complete the
construction documents by early spring 2012 and begin
construction as soon as possible. The plans call for a 30,000 square
foot addition to the existing building, to be located to the east, with
a 10,000 square foot basement to house the museum exhibit team;
a 12,000 square foot main level with a spectacular gallery to honor
President Boyd K. Packer; and then an 8,000 square foot 3rd level with
new exhibits, including a remarkable series of exhibits that illustrate the
major ecosystems of Utah. The project also calls for limited remodeling
in the existing building to add office space, storage space, and a
spectacular exhibit highlighting the Fred and Sue Morris bird collection,
which will be located immediately adjacent to President Packer’s
gallery. To say the least, the museum family is delighted!
In October, the museum’s first director and dear friend, Dr. Wilmer
Tanner, passed away. Dr. Tanner would have been 102 years old
in December. His positive influence on the museum has known no
limits. He gave freely of his time and personal resources to support
and build the museum. It seemed that the museum was always on
Dr. Tanner’s mind. Until the last six months of his life, Dr. Tanner was in
the museum working at least two days a week. Three days before he
passed away, he called and indicated that he was planning to come
to the museum and wondered if he could spend some time with me
while he was here. He called back later that day and indicated that
he wasn’t feeling well and would come later in the week. His example
and dedication to the museum and BYU were exemplary in every
way. In my more than 20 years of working at the museum, I have not
known anyone who loved and cared about the museum more than
Dr. Tanner. He is missed.
Dedication
On October 28th, two months short of his
102nd birthday, Dr. Wilmer W. Tanner
passed away, leaving a long lifetime of
accomplishments. Wilmer was born on
December 17, 1909, in Fairview, Utah. His
parents were John Myron and Lois Ann
Tanner, descendants of early Utah pioneers,
and they resided on a ranch in the open
and meadow-bottomed valley at Indianola
in Sanpete County. They had five children:
Vasco, Ray, Lena, Wilmer, and Jean. Wilmer
began his college studies at BYU
in the winter of 1932. He had to
drop out, however, and return to
ranching for a while to help pay off
family debts. Wilmer returned in the
fall of 1933, and despite a constant
struggle for financial support,
he enjoyed university life. By the
winter of 1934, he was certain
that his future was somewhere in
science, and he graduated with
a BA in the spring of 1936 with a
major and minor in Zoology and
Geology, respectively. In 1938, he
also completed a MS degree in
Zoology at BYU.
While at BYU, Wilmer met Helen Brown,
and on January 4, 1935, they were
married. Years later he declared in his
personal history that his children (Lynn, David,
and Mary Ann) and his 60-year marriage to
Helen were his most enduring achievements.
Wilmer attributed his academic and
professional success to Helen’s constant
encouragement and, in some cases, her
active participation; he was devastated
by her death in 1995. He nevertheless
maintained his academic and professional
activities and eventually married again at
age 90. His second wife was Otella Tyndal
Devey. Otella died in 1999.
In the fall of 1946, Wilmer started his
graduate program at the University of
Kansas. For financial reasons, Helen and the
children remained in Provo for the first year of
Wilmer’s graduate studies. They finally arrived
in Lawrence in June of 1947. Wilmer’s thesis
advisor was Edward Harrison Taylor. Wilmer
completed his research by June of 1949, and
in August, he defended his research and
filed his dissertation (published three years
later; Tanner 1952). Around the same time, he
Dr. Wilmer W.
Tanner
received a contract to teach Zoology at BYU
and moved his family back to Provo.
Wilmer’s early academic career was
characterized by heavy teaching
loads (three classes per semester). At that
time, the zoology faculty included his
brother Vasco, D Elden Beck, and C. Lynn
Hayward. Wilmer’s group in the Department
of Zoology & Entomology was soon joined by
entomologists Stephen L. Wood and Dorald
M. Allred; as a group, they taught
a range of courses that spanned
the entire breadth of zoology.
Wilmer and other BYU
faculty conducted what
research they could on their
own time and out of their own
pockets, all the while still carrying
heavy teaching loads. Wilmer’s
regular classes included, among
others, general biology, general
zoology, vertebrate anatomy,
herpetology, and genetics.
Financial constraints in his early
academic career limited his travel
to only regional meetings and events, and
even these had to be covered from personal
funds.
In the early 1970s a committee was
organized to explore the possibility of a
collections facility separate from the Botany
and Zoology departments. Eventually, Wilmer
became chairman of the committee and
worked closely with Steve Wood and Stan
Welsh to outline goals and procedures for
such a facility. The committee was actively
working on the concept of a collections
facility when a letter arrived from Monte
L. Bean, a wealthy businessman
from the Seattle area, in which
Monte offered his extensive
trophy collection to the
university. Monte’s
wife Birdie had been
encouraging him to
donate his collection
of trophy animals
for a number of
years. After some
preliminary
contacts, Wilmer
and Monte
Striped-Bass
Morone saxatillis

8 9
Brook Trout
Salvelinus fontinalis
Dr. D Elden Beck
Dedication cont.
met to discuss options, and soon after, Monte’s collection of trophy
mounts arrived and were displayed temporarily in a portion of the
main reading room of the old Grant Library building. What was now
required was a building large enough and sufficiently modern to
effectively house the university’s extensive life science
research collections. Wilmer and Monte, and members
of the committee, worked together to define what would
be required, and Monte both listened and agreed. Plans
were drawn up, presented to Monte and Birdie, and
construction was begun in the mid-1970s. Money for the
construction was donated by the Bean family, who also
established an endowment fund for the museum. The
Monte L. Bean Life Science Museum was completed in
the autumn of 1977 and officially opened to the public in
the early spring of 1978.
Wilmer served as first director of the museum, with Dorald Allred
and Dick Murdock as assistant directors. Forced by law to retire
at age 65, he continued to visit the museum almost daily to follow its
growth and development. He continued to publish until the time of his
death, with some papers to be published posthumously. Three days
prior to his passing, Wilmer contacted the present museum director
about meeting to discuss future publications. His body wore out, but
his mind never did!
Note: Much of the above narrative was taken from two obituaries written by Dr.
S.L. Welsh and Dr. J.W. Sites.
Historical Highlight
Historical Highlight – D Elden Beck was born April 11, 1906,
in Spanish Fork, Utah, to Ruth Eleanor Davis and Mitchell
Robertson Beck. Because his father had a difficult time finding
work, the family eventually moved to McGill, Nevada, where his
father worked in the mines. It was there that young Elden met
people from a variety of cultures and ethnic backgrounds—
Chinese, African Americans, Greeks, and others. The young men
from these backgrounds became his closest friends, and from
those interactions came one of Dr. Beck’s finest qualities—his
tolerance, respect, and kindness to those of different ethnic and
cultural backgrounds.
Early on, Beck struggled in school, except in biology—where
he excelled! However, his determination, his ability to work
hard, and his faith led him to enroll at BYU in the fall of 1925. He lived
in Spanish Fork with his grandmother Grace Matley and commuted
every day to BYU on the Orem electric interurban train. The terminal
was near the southwest corner of Center Street and University
Avenue in Provo. Those early years at BYU were extremely difficult.
Not being a gifted student, Beck had to work harder. According to
Dr. Vasco Tanner, a lifetime friend and mentor, Beck arrived early at
the laboratory and was one of the last to leave at night. The students
and teachers with whom he worked influenced his growing interest in
entomology. He was frequently chosen to participate in field trips as
a collector.
In 1929 he
received a
BA degree,
and in
1930, an
MA degree
in zoology
and entomology.
Subsequently, he was accepted for further
graduate studies at Iowa State College in
Ames, Iowa.
In 1933, Dr. Beck was appointed head
of the Biology Department at Dixie
College, married Florence Robinson, and
was awarded his PhD degree from the
Department of Entomology at Iowa State
College. Upon moving to St. George, he
immediately became immersed in the
community. He studied landscape art
with his talented wife, Florence, and their
mutual interest in painting resulted in the
establishment of a Fine Arts Festival in 1934
at the college, which has continued to this
day. Dr. Beck also cultivated a profound
interest in photography, a passion that lasted
throughout his life. Most of his photographs,
negatives, and slides are stored in the
archival collection in the Harold
B. Lee Library at Brigham
Young University. In 1938,
he was asked to return
to BYU as an assistant
professor of zoology and
entomology. He left Dixie
College with mixed emotions; his heart was
filled with a deep love and appreciation for
the people and places of southern Utah.
From 1942 to 1944
(during WWII),
Dr. Beck worked in
the United States Medical
Entomological Services. Part of that service
was in Guadalcanal in the Solomon Islands.
It was in these remote jungles that Dr. Beck
distinguished himself as director of mosquito
control activities. He also spent time
collecting thousands of insects, reptiles, etc.,
all of which were carefully shipped to BYU to
become a part of the university’s research
collections. After the war, he returned to his
scholarly and professional pursuits at BYU,
where he was recognized as a devoted
teacher who also had a sense of humor.
Among his accomplishments while at BYU,
Dr. Beck produced an impressive list of more
than 60 scientific
publications.
In addition to
his academic
and professional
responsibilities
at BYU, Dr. Beck
was also involved
in various
service-oriented
projects. Some
of his accomplishments were related to
his academic work. For example, he was
assigned by the Utah County Commissioners
and the Health Department to develop
a mosquito abatement program for Utah
County. And, from 1956 to 1957, he
worked with the United Nations World
Health Organization to eradicate
serious malaria-related problems
in Taiwan.
After a long struggle with
emphysema, Dr. Beck died
on August 9, 1967. Prior to his death, he
established a scholarship fund for zoology
students at BYU, which started in 1970 and
continues to the present day.
Note: Much of the above narrative was prepared
by Janet Beck Clark, Dr. Beck’s daughter.
Yellow Perch
Perca flavescens

N
600 EAST
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BYU Broadcasting
Current Museum
Rendering of the
museum addition
Expansion!
Museum
Addition
After more than 10 years of fundraising, the university’s board of
trustees authorized preliminary planning for a major addition to the
museum. In April, the first meetings of the museum expansion steering
committee were held, and by May, the university had received
permission to both formally plan and construct the addition!
In June, we solicited proposals from several architects, and
after visiting four of the respondents, we selected Jacoby
Architects of Salt Lake City. Jacoby has an impressive portfolio
of work that crosses the state and includes several new
buildings at Utah State University. Wadman Construction
out of Ogden was selected as the contractor. Throughout
the summer months and into the fall, we have worked with
both the architects and the contractor on the programming,
design, and development documents for the museum
addition.
The addition will include 30,000 square feet distributed
over three floors, matching the three floors in the existing
building. The basement will house the museum’s exhibit
planning, design, construction, and installation team in one
contiguous area. The ground level will include the President
Boyd K. Packer Gallery where we will exhibit President
Packer’s magnificent bird carvings and other artwork.
President Packer’s gallery will also provide a wonderful venue
for declaring the message of Jesus Christ as the Creator!
The ground level will also house the museum’s education
team, including two classrooms, a new live-animal facility,
and offices and work space for our education group. The
upper level will accommodate a series of new exhibits surrounding a
beautiful center atrium. The expansion will also allow the museum to
better accommodate the invaluable research collections, which will
remain in the existing building with additional space and improved
security. We hope to have construction documents ready by the
spring of 2012 and for construction to begin in the late spring or early
summer!
New Exhibits
The museum’s exhibits generally fall into several simple
categories—permanent, long-term but subject to change, and
temporary—and often celebrate a particular event or explore a
specific current issue. Two of the exhibits below fall into the more
permanent category, while one is long-term but will eventually be
changed, and the last is clearly temporary.
Renovated Arctic Exhibit –
During the summer, Ken Packer and the exhibits crew moved and
completely renovated the Arctic exhibit that had occupied the
northwest corner of the center atrium for the last several years. The
new location, against the north stairwell of the main level, provided
more space and brought the exhibit into the center atrium gallery.
Again our exhibits team made impressive use of the high-density
Styrofoam to create a remarkably
realistic Arctic scene that conveys
targeted information about
several Arctic animals.
Into Africa –
During 2011, the museum’s
exhibits staff completely
remodeled the Bean Memorial
Room and installed a new exhibit
highlighting some of the unique
ecosystems features of Africa.
Randy Baker and his students
hand-painted a remarkable mural
that stretches around the exterior
wall of the exhibit area from floor
to ceiling! Using high-density
styrofoam, Ken Packer and Clark
Brereton developed a unique
modular foreground plan to
support the education element of
the exhibit, which highlights several
of the major ecological zones
of Africa. The exhibit opened in
August and will be a permanent
feature of the expanded museum.
Expansion Exhibit –
Shortly after the museum
received permission to expand,
we put together plans for a
free-standing “Expansion Exhibit”
for the north main-level entrance
to the museum. This exhibit
highlighted the Bean family’s early
gift of the existing building, as well
as basic information about the
museum addition. In particular, the
exhibit included details about the
President Boyd K. Packer Gallery,
a major feature of the addition
that highlights President Packer’s
remarkable bird carvings!
Renovation of the Cougar Exhibit in
the Wilkinson Student Center –
The museum’s exhibit team
was invited to redo the
“Cougar” exhibit at the
north entrance to the Wilkinson
Student Center. Our exhibits group
completely reworked the old rock
face and added two new cougars
and a deer to the display. The
exhibit adds a new and impressive
highlight to the Wilkinson Center
north entrance!
Education
Programming
Nature Experienceships –
During 2011, the museum
sponsored four successful
Nature Experienceships. Two
of these experienships were
with Merrill Webb, once in April
and then again in September.
Our birding exprienceship
continues to be one of the Merrill Webb- Birding Experienceships
more popular field trips so we
plan to continue offering
two birding experiences
each year. Our other two
experiencships were in August
and October. We had a
good size group learn about
mammals with Duke Rogers;
and for the second time,
Sam St. Clair led a field trip Sam St. Clair- Quaking Aspen
to Provo Canyon to look at
and talk about the Quaking
Aspen community. Our nature
experienceship program
provides our patrons with a
unique opportunity to interact
with experts in specific life
science areas. Anyone 12
years and older is invited!
Date Night –
Duke Rogers- Mammals
The museum held a “Clue”
themed date night in February and October. Both
events were sold out! This activity continues to be
popular with our patrons. Date nights include dinner
and various types of engaging activities.
Northern Pike
Exox lucius

12 13
Rainbow Trout
Oncorhynchus mykiss
Life Science
School Programs
Look what’s happening this year!
2011-2012
Monte L. Bean Life Science Museum
Brigham Young University • Provo, Utah
mlbean.byu.edu (801) 422-5052
Education
Programming Cont.
Development of an iPad dichotomous key –
During 2011, the iPad app, “IdentifyMe,” was launched and
submitted to the App Store. It was designed to be used with the
iPad, iPhone, or iPod touch. On-site patrons who have a
3G network data plan can download the “Identify Me”
app to their own devices. This app can be used to identify
specially labeled specimens throughout the museum
by carefully observing each specimen and answering a
series of questions. The museum also purchased five iPads
preloaded with the “Identify Me” app that are available
for checkout at the museum store for $3. In conjunction
with the new “Into Africa” exhibit, Matt Meese and Katy
Knight also added a dichotomous key to the app that
highlights several of the animals in the exhibit. In addition,
they prepared 16 short videos for the “more info” section
of the exhibit. These videos are available on both YouTube
and the museum’s website (mlbean@byu.edu). They can also be
accessed using any smart phone with a data plan.
Bio-Box Initiative Update –
The Adaptations and Ecosystems Bio-Boxes continue to be a
popular way for local science teachers to bring real specimens,
activities, and educational resource materials to their students. The
boxes were checked out 22 times during the school year. Bio-Boxes
are the next best thing to a field trip to the museum! We continue
to make progress on the Utah’s Wetlands, Forests, and Deserts Box.
We currently have 20 activities that tie directly to the Utah 4th grade
science core curriculum. We hope to have our third Bio-Box ready
sometime in 2012.
Partnership School Districts –
We produced and distributed another Teachers’ Guide
this year that highlights the museum’s 2011–2012 school
programs. This document provides each of the teachers
in the partnership districts with pertinent information about
the various ways they can access the museum’s unique
and valuable resources. The museum partners with our five
local school districts to provide teachers and students with
enhanced support for learning about the life sciences.
We also set up a listserv with the help of Danny Yeo of Life
Sciences Computer Support, so now it is much easier to send
emails to all of the teachers in the partnership districts.
Live-Animal Webcast Initiative –
During 2011, we completed three trial runs with our new
“webcast” live-animal shows. This new program allows
teachers to bring the museum’s live-animal shows in real
time to their classroom using readily available remote video
technology. The museum education team has been proactively
developing this program in anticipation of the museum expansionrelated
closure next year. They also plan to increase their efforts with
the live animal shows as part of the museum’s outreach program.
Central Utah Science and Engineering Fair –
This year, the Central Utah Science and Engineering Fair team
moved to the museum. The Cluff Building had been home to this
program for many years. However, with construction of the new life
sciences building, the Cluff Building was scheduled for demolition,
and the CUSEF team moved to the Bean Life Science Museum.
The program has been under the direction of John Gardner and
Lisa Clarke for many years. Lisa Clarke has been involved with the
program for more than 10 years – growing the fair from a one-day
event with 200 participants to a three-day event with nearly 900
participants! This past spring, the fair hosted 875 participants over
three days in the Harmon Continuing Education Building at BYU.
Students received cash awards and prizes of more than $65,000 –
as well as more than $500,000 in scholarships to local colleges and
universities. Five students were also selected to represent Central
Utah at the 2012 Intel International Science and Engineering
Fair in Pittsburgh. Next spring, Lisa will step down and assume
full-time duties as a mother, and Nettina Smith will assume CUSEF
management responsibilities.
Western North American Naturalist
In 2011, the museumsponsored
natural
history journal, Western
North American Naturalist,
continued in its 72nd year of
publication. The WNAN office
handled 119 manuscript
submissions and published
661 pages among 3
monograph articles, 7 special
feature articles, 42 regular
articles, 16 notes, and 2 book
reviews. Usage at BioOne.
org, WNAN’s major online
distributor, exceeded 8,000
full-article downloads, and
revenues from this source
continue to increase year to
year.
In August 2011, WNAN
published volume 5 of the
monograph series, which
volume included articles on
bats of Mesa Verde National
Park, birds of Glen Canyon
National Recreation Area,
and cyanobacteria in the
Mojave Desert. Another
debut in December 2011 was
2011 Central Utah Science
and Engineering Fair
a special feature on Lake
Suckers. Additional special
features, with a goal of one
per year, are planned to
highlight little-studied taxa or
ecological systems. WNAN
also granted its 4th annual
“Outstanding Natural History
Paper” to Andrew Kulmatiski,
K.H. Beard, L.A. Meyerson,
J.R. Gibson, and K.E. Mock
for their contribution entitled
“Nonnative Phragmites
australis invasion in Utah
wetlands.”
Largemouth Bass
Micropterus salmoides

14 15
King Salmon
Oncorhynchus tshawytscha
Jon Gelhaus
Mark Belk
Museum Sponsored
Lectures and Events
Every year the museum sponsors at least two
lectures: the fall and spring Tanner lectures.
The museum also periodically hosts DVD
screening events highlighting topics that support
the museum’s mission statement. Occasionally,
the museum will also sponsor special lectures
to celebrate the opening of new exhibits or
to highlight a special event. During 2011, the
museum sponsored two Tanner lectures and
one DVD screening. Note: The Tanner lectures
can be viewed on the museum’s website
(mlbean.byu.edu).
Spring Tanner Lecture –
Mongolia was the focus of the spring 2011 Bean Life Science
Museum John Tanner Lecture, given by Dr. Jon K. Gelhaus of
the Academy of Natural Sciences of Philadelphia. He and his team
have been surveying the aquatic invertebrates of this remote country
for more than 10 years as they document the distribution of these
animals. Dr. Gelhaus and his team use the distribution data to assess
the environmental quality of streams, rivers, lakes, and wetlands
in this quickly developing nation of nomads. Mining and grazing
are having noticeable impacts on the landscape and lifestyles of
these people, and the Mongolian Aquatic Insect Survey, under
the guidance of Dr. Gelhaus, has been training native Mongolians
in modern systematic and ecological methods for identifying and
classifying both the animals and the watercourses. In addition, during
his visit, Dr. Gelhaus also helped curate and update identifications for
the museum’s Crane fly collection.
Fall Tanner Lecture –
In his fall Tanner lecture, Dr. Mark C. Belk discussed how knowledge
of the biology of living things allows us to better understand and
appreciate the living world. He used examples from his study of the
evolutionary ecology of fishes to illustrate the beauty and wonder
so commonly observed in the natural world. He described the main
challenges that fishes face by focusing on the unique adaptations of
fishes in terms of how they feed, avoid predation, and find a mate.
Examples included protrusible jaws, electrosensory capabilities, poisonous
spines, live-bearing, and spontaneous sex change. Dr. Belk concluded by
emphasizing to the audience the need for each of us to be personally
responsible as good stewards of the earth’s living community.
Screening of “Green Fires” DVD with introduction by Stanley Temple –
During winter semester 2011, the museum sponsored a screening
of the new DVD “Green Fires.” We were fortunate to have Dr.
Stanley Temple, from the Aldo Leopold Foundation at the University
of Wisconsin, attend and introduce this new and intriguing DVD.
More than 200 people attended the “Green Fires” screening. “Green
Fires” celebrates the life and contributions of the early American
conservationist Aldo Leopold. This event was cosponsored by the
Charles Redd Center on campus.
Collections-Based Research –
At the core of the museum’s mission statement is a
clear commitment to collections-based research. The
museum’s nine research-quality collections are used by
faculty, students, and visiting scientists to explore and
better understand the diversity and complexities of life on
earth. Each collection is cared for by the museum’s curatorial
staff. The productivity of the museum curatorial team has
been consistently impressive, and, in 2011, the team
secured millions of dollars of external research funds and
published more than 80 peer-reviewed journal articles!
Visiting Fulbright Scholar –
During fall semester 2011, Dr. David
Muranyi of the Hungarian
Natural History Museum, Budapest,
Hungary, was a postdoctoral
researcher at the museum. He is the
most outstanding young plecopterologist
in Europe and his visit benefited both him
and the museum. His research project
was on the stoneflies of the Balkans, and
he emphasized the genera Amphinemura
and Isoperla. In addition, he studied the
winter stonefly family Capniidae, and his
findings will completely revolutionize the taxonomy of this family
in the Holarctic realm. While in Provo, Dr. Muranyi also studied
an interesting series of large perlid stonefly nymphs that we had in
our holdings, taken from waterfalls in Iran in 2004. He brought a nice
collection of Palearctic stoneflies with him that he donated to BYU,
and we shared with him stonefly species that he needed from North
America. The Muranyi family, David, Szilvi, and Andris (2 years old), lived
in Wymount Terrace from September 2011 to January 2012. They had many
great experiences here in Utah.
Dr. David Muranyi
Studying Fishes Across Latin
America –
For some, catching fish is a relaxing hobby.
For Jerry Johnson, Assistant Curator of Fishes,
collecting fishes is just short of an obsession. Johnson
spends time every year exploring rivers and streams
in Central and South America sampling all kinds of
freshwater species. It’s more than just fun though.
Johnson and his students are pursuing important
research questions focused on the origins of fish biodiversity.
This research has taken them from the coasts of Mexico all
the way to Patagonia at the southern tip of South America.
Although Johnson loves fishes, he’s even more excited to be
able to share his passion with BYU students. “I love being in the
field collecting. But one of the best parts of my job as a curator
is allowing students to experience the thrill of conducting field
work.” Most of the fishes collected in this research end up back
at the Bean Museum in our research collections, where students
and scientists can use them for teaching and research. Now
that’s the kind of work you can get hooked on!

16 17
Collections-Based
Research – cont.
Research at the Lytle Ranch Preserve –
Currently, there are several long-term research projects
operating at the Lytle Preserve. One project involves an
impressive team of several young BYU faculty members examining
the effects of fire in the Mojave Desert. The Mojave Desert has
only recently been exposed to fire, and this exposure is due to
the introduction of invasive species that provide sufficient fuel to
support wildfires. There is no background data on how the Mojave
Desert ecoregion will respond to this new kind of impact. Randy
Larsen and Brock McMillan from the Department of Plant and Wildlife
Sciences are two of several BYU faculty members actively conducting
research at the Lytle Preserve. In this section, we highlight Brock and
Randy’s student-centered research.
Wildfire and Small Mammals –
Dr. Brock McMillan, Department of Plant and Wildlife Sciences,
and his students are involved in a collaborative effort to look
at the interactive effects of fire and small mammals on various
functional attributes of the Mojave Desert ecoregion. They have
erected 20 small-mammal exclosures and used controlled burns
to (1) determine the effects of wildfire on small mammals, (2)
examine the impact of small mammals on seed and seedling
survival, and (3) explore the interactive effects of fire and small
mammals on seeds and seedlings. To monitor the small mammal
community, they have been live-trapping for three consecutive
nights each April, July, and October. In order to effectively assess the
effect of small mammals on plants, they have also been conducting
seed and seedling predation experiments.
Gambel’s Quail and Kit Fox –
Dr. Randy Larsen’s research is a collaborative effort with state
and federal agencies, as well as nonprofit organizations, to
evaluate the influence of water developments in the Mojave Desert
on wildlife. More specifically, Randy and his students are interested
in the ecology and behavior of Gambel’s Quail and kit fox (two
aridland specialists) in relation to available drinking water. The
goal of their research is to determine whether or not wildlife water
developments represent a positive, negative, or neutral influence
on habitat use and the population dynamics of both species. To
assess the influence of water on these two species, they are using
radiotelemetry (Gambel’s Quail) and remote cameras
(Gambel’s Quail and kit fox). Both lines of research are
time intensive and require lengthy stays at the Lytle
Preserve during the summer field season.
Cynthia Troxell Lichen Collection Donated –
In the fall, Dan Troxell emailed Larry St. Clair (curator
of nonvascular cryptogams – lichens, bryophytes,
etc.) and inquired about donating his wife’s lichen collection
to the herbarium. Cynthia had been a graduate student at the
University of Colorado at the same time Larry was working on
his PhD at CU. They interacted briefly as Cindy finished up her
MS degree, and then she moved on to Louisiana with her husband’s employment. Cindy
continued to collect lichens wherever she and Dan lived. In early 2011, Cindy was diagnosed
with cancer and passed away in the summer. As Dan sorted through Cindy’s lichen
collection, he realized the specimens could be a valuable resource if placed in the right
hands. He worked with Bob Egan at the University of Nebraska and Larry St. Clair here in the
museum to see that Cindy’s collection was properly secured. Cindy’s mother also donated
funds to help with the care of the collection. Those funds were placed in the herbarium
endowment. Cindy and Dan’s gift to the museum has become an important part of the
lichen collection, which now includes more than 100,000 specimens.
Donors –
The museum’s donors play a central role in funding the operations of the museum. The
university generously supports many of the basic functions of the museum, but our donors
provide funds essential for supporting the three core programs of the museum: education,
exhibits, and research collections. Without the support of our generous donors, the capacity
of the museum to support these core programs would be drastically limited.
Mellor Family Donate to the Museum’s Education Enhancement Program –
In September, Reed and Lareen Mellor, along with their two sons, Gabe and Jade, donated
$10,000 to the museum’s education enhancement program. These funds will be used to
expand and strengthen the museum’s education programming so that we can better serve
the thousands of school children who visit the museum each year. The Mellor family gives
generously to a variety of worthwhile causes, and Reed and Lareen actively involve their
sons in the selection process. We deeply appreciate the generosity and support of the Mellor
family!
Donors to the Education Partnership –
One of the more recent initiatives of the museum has been the education partnership.
This program invites corporate partners to consider providing funds to enhance the
museum’s education programming. Every year, the museum’s education team interacts
with thousands of school children, both in the museum and through our outreach programs.
Generous corporate support is essential to the museum’s efforts to share the unique resources
of the museum with our public and private school partners. In 2011, we received generous
donations from Craig and Jodie Allred of Springville and Tahitian Noni International. These
corporate sponsors allow us to bring the best of the life sciences to our school-age children!
Donor Box at North Entrance –
In the fall of 2009, the museum placed a modest “donation box” at the north entrance to
the museum with an invitation to our patrons to give back to the museum. When Monte L.
and Birdie Bean donated the money to build the museum, they specifically requested that
access to the museum always be free, and we have honored that request faithfully over
the years. However, extending an opportunity to our patrons to offer a modest donation has
yielded impressive results! Since the “donation box” was set out in the fall of 2009, almost
$8,000 has come to us as pennies, dimes, and quarters! These funds are dedicated to
visitor services and will consistently come back to help our patrons better engage with the
museum’s resources. Thanks to everyone!
Brook Trout
Salvelinus fontinalis

18 19
The Lytle Ranch Preserve –
The Lytle Ranch Preserve is a 600 acre natural preserve located
along Beaver Dam Wash at the Utah–Nevada border in
Washington County. The preserve has unique ecological and
biological features, as well as a fascinating history. The preserve is
administered by the M.L. Bean Life Science Museum and serves as a
field station for faculty and student learning and research as well as
a remote—but remarkable—location for observing nature!
Lytle Preserve Use Policy –
At any one time, there are now 5–7 active research
projects underway at the Lytle Preserve involving
7–10 faculty researchers and 75–100 students. With this
increased activity and traffic, it became expedient that
we develop and implement a use and fee policy for the
preserve. This task was completed in 2011. The fees are
modest and are used to improve or develop appropriate
support facilities/resources at the preserve. Also, this policy
provides the preserve manager, Heriberto Madrigal, with
clear guidelines and expectations for researchers and
visitors at the Lytle Preserve. The Use and Fee Policy is
posted on the museum’s website (mlbean.byu.edu).
First Annual Lytle Ranch Preserve Research Meeting –
In December, the museum administration met with the various
groups actively conducting research at the Lytle Preserve. The
intent of this meeting was to provide better coordination and
oversight for the various research projects currently in place at the
preserve. Each researcher was asked to (1) prepare a 5–10 minute
presentation highlighting their research efforts at the preserve; and
(2) submit a written report documenting their 2011 activities at the
preserve and outlining their plans for 2012. The meeting provided
everyone with valuable information while accommodating an open
discussion about concerns and needs.
Flood-related Recovery Efforts and Plans –
In December of 2010, a catastrophic flood caused major damage
to the Lytle Ranch Preserve. The irrigation system and old ranch
house were seriously damaged, and the hay fields and adjacent
orchards were inundated with sediment. Through the winter and
early spring months, Heriberto and Ken worked to put critical
resources back online while we began to work with our campus
planning team to assess the damage and develop a recovery
plan. By spring, Heriberto had the irrigation system functional,
and by summer, we had started working with a local St. George
engineering firm (Bowen and Collins) to develop a comprehensive
flood mitigation plan. It was determined that the old ranch house
and trailer were beyond recovery and that it was imperative that we
move all of the preserve’s critical infrastructure out of the flood plain.
From fall into early winter, we developed a solid flood mitigation plan
and early drawings for a replacement ranch house. The prospects
are promising and the plans and associated costs will be reviewed
by the Church in the winter and early spring of 2012.
Visitor Services –
One of the key operational aspects of the museum is the museum’s commitment to
provide each patron with a significant, memorable experience. In some way, every
member of the museum family contributes to this important assignment. Our goal is to ensure
that every patron is treated with respect and kindness. Thanks to the museum employees,
students, and volunteers who make the museum experience extraordinary!
Museum Store –
In spite of a struggling economy, sales in the store were once again impressive in 2011.
Revenue generated by the sale of store merchandise covered the wages for our five
student employees. We sincerely appreciate our student employees and staff whose diligent
efforts ensure that the store merchandise is of high quality and directly supports the mission
of the museum. We look forward to the new and larger museum store which will be a key
element in the museum addition.
Museum Volunteers –
Volunteers play a crucial role in the operation of the museum, and in 2011, the museum
was the recipient of over 5,700 volunteer hours! Volunteers serve in a variety of capacities,
including help with the research collections, visitor services, education, and exhibit design
and development.
Touch Screen Installed –
In order to better facilitate patron access to the museum’s extensive collection of
educational videos, a touch-screen interface was designed and installed by Broderick
Klemetson, one of the museum’s student employees. This wall-mounted control system allows
patrons to select and view videos on an adjacent large screen LCD TV.
Museum Newsletter –
In 2011, two issues of the museum newsletter were published and mailed to more than
5,000 museum patrons while another 1,500 patrons received copies through email. The
newsletter is also available on the museum web site and in either foyer of the museum.
Museum Staff Awards –
Each year, the museum staff nominates colleagues for three types of museum awards:
curation contributions, service efforts of the full-time museum staff, and contributions by our
student employees. This year, a total of six individuals were honored:
Curator/Collection Manager:
Dr. Jack Sites, Curator of Reptiles and Amphibians
Staff/Administrative:
Patty Jones, Assistant to the Director
Student Employees:
McKenzie Giles (Secretary)
Hilary Norton (Museum Store)
Angela Jensen (Herbarium)
John Whitehead (Custodial)
Common Carp
Cyprinus carpio

20 21
Rainbow Trout
Oncorhynchus mykiss
Albino Rainbow Trout
Oncorhynchus mykiss
Annual Financial Report –
slowly improving 2011 economy resulted in a slight increase
A in the museum’s endowments in terms of both principal and
spendable income. The earnings from our more than 30 endowments
made it possible to continue to effectively care for and support the
museum’s various operations, which include exhibits, education, and
research collections. The curatorial team was also able to secure an
impressive amount of external funding, which contributes directly to
the care and upkeep of the museum’s research collections. We are
also deeply grateful to the college and university administrations for
their consistent support for many of the museum’s basic operational
expenses, which include security, physical facilities, equipment
funding, salaries for our full-time staff, and supply money to help
meet the everyday needs of the museum. In turn, the museum staff
consistently demonstrates an impressive commitment to carefully
and thoughtfully managing the museum’s resources while producing
quality products and opportunities for our patrons.
Credits –
Again, sincere thanks to Randy Baker
for the beautiful design and layout
for this year’s annual report; likewise, I
am deeply grateful to Janene Auger
and the Western North American
Naturalist team for their careful and
thoughtful editing of the report narrative.
At the risk of being redundant, I cannot
say enough about all of my colleagues
at the Bean Life Science Museum. They
are unselfish and absolutely dedicated
to the success and improvement of the
museum. Their numbers are small, but
their efforts are consistently remarkable!
Bluegill
Lepomis macrochirus
2011 Research Publications
Janene Auger, Assistant Editor WNAN
Hatch, K.A., B.L. Roeder, R.S. Buckman, B.H. Gale, S.T. Bunnell, D.L. Eggett, J. Auger, L.A.
Felicetti, and G.V. Hilderbrand. 2011. The correlation between gross fecal analysis of
archived black bear scats and isotopic signatures. Ursus 22(2):133–140.
Wojda, S.J., M. McGee-Lawrence, R.A. Gridley, J. Auger, H.L. Black, and S.W. Donahue.
2011. Yellow-bellied marmots (Marmota flaviventris) preserve bone strength and
microstructure during hibernation. Bone [http://dx.doi.org/10.1016/j.bone.2011.10.013].
Ian T. Baldwin, Faculty Research Affiliate
Diezel, C., D. Kessler, and I.T. Baldwin. 2011. Pithy protection: Nicotiana attenuata’s
Jasmonic acid-mediated defenses are required to resist stem-boring weevil larvae. Plant
Physiology 155:1936–1946.
Stitz, M., I.T. Baldwin, and E. Gaquerel. 2011. Diverting the flux of the JA pathway in
Nicotiana attenuata compromises the plant’s defense metabolism and fitness in nature
and glasshouse. PLoS One 6(10):e25925.
Stork, W.F.J., A. Weinhold, and I.T. Baldwin. 2011. Trichomes as dangerous lollipops: do
lizards also use caterpillar body and frass odor to optimize their foraging Plant Signaling
and Behavior 6(12):1893–1896.
van Doorn, A., G. Bonaventure, I. Rogachev, A. Aharoni, and I.T. Baldwin. 2011. JA-Ile
signaling in Solanum nigrum is not required for defense responses in nature. Plant, Cell and
Environment 34:2159–2171.
Weinhold, A., and I.T. Baldwin. 2011. Trichome-derived O-acyl sugars are a first meal
for caterpillars that tags them for predation. Proceedings of the National Academy of
Sciences of the United States of America 108(19):7855–7859.
Richard W. Baumann, Emeritus Curator of Insects:
Baumann, R.W., and B.C. Kondratieff. 2011. Collecting endemic and rare stoneflies
(Plecoptera) in California, U.S.A. Perla 29:13–19.
Lee, J.J., and R.W. Baumann. 2011. Mesocapnia aptera (Plecoptera: Capniidae) a new
wingless winter stonefly from northern California, U.S.A. Illiesia 7:192–196.
Shepard, W.D., and R.W. Baumann. 2011. Canopy fogging in the Valdivian
forests of southern Chile produces stoneflies (Plecoptera). Illiesia 7:127–
132.
Stark, B.P., and R.W. Baumann. 2011. Records of Anacroneuria
(Plecoptera: Perlidae) from Bolivia and Paraguay with descriptions of
three new species. Illiesia 7:182–191.
Zwick, P., and R.W. Baumann. 2011. Nemoura jejudoensis a new species
of stonefly and the redescription of Amphinemura baei Ham and Lee
(Plecoptera, Nemouridae) from Jeju Island, Korea. Illiesia 7:148–155.
Shawn M. Clark, Collection Manager, Arthropods
Barney, R.J., S.M. Clark, and E.G. Riley. 2011. Annotated list of the leaf beetles
(Coleoptera: Chrysomelidae) of Kentucky: subfamily Cryptocephalinae. Journal of
the Kentucky Academy of Science 72(1):3–23.
Barney, R.J., S.M. Clark, and E.G. Riley. 2011. Annotated list of the leaf beetles

22 23
Channel Catfish
Ictalurus punctatus
(Coleoptera: Chrysomelidae) of Kentucky: subfamily Eumolpinae.
Journal of the Kentucky Academy of Science 71(1–2):3–18.
Chaboo, C.S., and S.M. Clark. 2011. The beetle spawn of John Wilcox:
Shawn Clark. Chrysomela Newsletter 53:14–16.
Keith A. Crandall, Curator of Crustaceans
Bybee, S.M., H.D. Bracken-Grissom, R.A. Hermansen,
M.J. Clement, K.A. Crandall, and D.L. Felder.
2011. Directed next generation sequencing for
phylogenetics: an example using Deacapoda
(Crustacea). Zoologischer Anzeiger 250:497–506 [http://
dx.doi.org/doi:10.1016/j.jcz.2011.05.010].
Bybee, S., H. Bracken-Grissom, B. Haynes, R.
Hermansen, R. Byers, M. Clement, J. Udall, and K.A.
Crandall. 2011. Targeted amplicon sequencing (TAS): a
scalable next-gen approach to multi-locus, multi-taxa
phylogenetics. Genome Biology and Evolution 3:1312–
1323 [http://dx.doi.org/10.1093/gbe/evr106].
Yang, C.-H., H. Bracken-Grissom, D. Kim, K.A. Crandall, and T.-Y. Chan.
2012. Phylogenetic relationships, character evolution, and taxonomic
implications within the slipper lobsters (Crustacea: Decapoda:
Scyllaridae). Molecular Phylogenetics and Evolution 62(2012):237–250
[http://dx.doi.org/doi:10.1016/j.ympev.2011.09.019].
Barber, B.R., P.J. Unmack, M. Pérez-Losada, J.B. Johnson, and K.A.
Crandall. 2011. Different processes lead to similar patterns: a test
of codivergence and the role of sea level and climate changes
in shaping a southern temperate freshwater assemblage. BMC
Evolutionary Biology 11:343 [http://dx.doi.org/doi:10.1186/1471-2148-
11-343].
Sinclair, E.A., A. Madsen, T. Walsh, J. Nelson, and K.A. Crandall. 2011.
Cryptic genetic divergence in the giant Tasmanian freshwater
crayfish, Astacopsis gouldi (Decapoda: Parastacidae); implications
for conservation. Animal Conservation 14:87–97 [http://dx.doi.
org/10.1111/j.1469-1795.2010.00395.x].
Michael W. Hastriter, Research Associate
Hastriter, M.W., and R.D. Sage. 2011. Description of a new species
of Ectinorus (E. spiculatus) (Siphonaptera, Rhopalopsyllidae) from
Argentina and a review of the subgenus Ichyonus Smit, 1987. ZooKeys
124:1–18.
Richard A. Heckmann, Emeritus Curator:
Heckmann R.A., A. Khan, and O.M. Amin. 2011. Juveniles of the
Genus Oligacanthorhynchus Travassos, 1915 (Acanthocephala:
Oligacanthorhynchidae Southwell and Macfie, 1925) from snakes in
Karachi, Pakistan. Pakistan Journal of Zoology 43:604–606.
Heckmann, R.A. 2011. Comments on Amoeba and Coccidia two
overlooked and important groups of parasites and other parasites of
fishes. Proceedings of Parasitology 51:1–27.
El Naggar, A., R.A. Heckmann, and S.A. Mohamoud. 2011. A survey and history of helminth fish
parasites from the Nile River, Egypt. Proceedings of Parasitology 52:59–85.
Amin, O.M., R.A. Heckmann, and N.V. Ha. 2011. Description of two new species of
Rhadinorynchus (Acanthocephala, Rhadinorynchidae) from marine fish in Halong Bay,
Vietnam, with a key to species. Acta Parasitologica 56:67–77.
Heckmann, R.A., G.I. McCallister, and Z. McCallister. 2011. Prevalence and fine structure of
mites attached to mosquitoes from west central Colorado, USA. Proceedings of Parasitology
51:123–141.
Heckmann, R.A., O.M. Amin, Y. Tepe, S. Dusen, and M.C. Oguz. 2011. Acanthocephalus ranae
(Acanthocephala, Echinorhynchidae) from amphibians in Turkey, with special references to
new morphological features revealed by SEM and histopathology. Scientia Parasitologica
12:23–32.
Amin, O.M., M.C. Oguz, R.A. Heckmann, Y. Tepe, and Y. Kvach. 2011. Acanthocephaloides
irregularis n. sp. (Acanthocephala; Arhythmacanthidae) from marine fishes off the Ukranian
Black Sea coast. Systematic Parasitology 80:125–135.
Amin, O.M., R.A. Heckmann, and A.M. El-Naggar. 2011. Revisiting the morphology of
Acanthocephalus lucii (Acanthocephala: Echinorhynchidae) in Europe, using SEM. Scientia
Parasitologica 12:197–201.
Amin, O.M., R.A. Heckmann, and A.M. El-Naggar. 2011. The morphology of a unique
population of Corynosoma strumosum (Acanthocephala, Polymorphidae) from the Caspian
seal, Pusa caspica, in the land-locked Caspian Sea using SEM, with special notes on
histopathology. Acta Parasitologica 5b:1230.
El-Naggar, A.M., R.A. Heckmann, A.A. Ashour, M.H. Rizk, S.H. Mohamed, and S.A. Mohmoud.
2011. Ultrastructural studies for spermigonesis of Acanthostomum aswaninesis (Digena,
Acanthostomatidae) a helminth parasite of Bagrus bayad (Forsskal 1775) (Osteichthyes)
Scientia Parasitologica 12:85–92.
El-Naggar, A.M., R.A. Heckmann, A.A. Ashour, M.H. Rizk, S.H. Mohamed, and S.A. Mohmoud.
2011. Additional Information for spermiogenesis and sperm ultrastructure of Acanthostomum
aswaninesis (Digenea, Acanthostomatodiae), a helminth parasite of Bagrus bayad (Forsskal
1775) (Osteichthyes). Proceedings of Parasitology 52:1–18.
Jerald B. Johnson, Assistant Curator of Fishes
Barber, B.R., P.J. Unmack, M. Pérez-Losada, J.B. Johnson, and K.A. Crandall. 2011. Different
processes lead to similar patterns: a test of codivergence and the role of sea level and
climate changes in shaping a southern temperate species assemblage. BMC Evolutionary
Biology 11:343 [http://dx.doi.org/10.1186/1471-2148-11-343].
Belk, M.C., E.E. Nance, and J.B. Johnson. 2011. Life history of Brachyrhaphis parismina:
variation among and within populations. Copeia 2011:372–378.
Adams, M., S.D. Wedderburn, P.J. Unmack, M.P. Hammer, and J.B. Johnson. 2011. Use of
congeneric assessment to reveal the linked genetic histories of two threatened fishes in the
Murray-Darling Basin, Australia. Conservation Biology 25:767–776.
Zúñiga-Vega, J.J., M. Suarez-Rodriguez, H. Espinosa-Perez, and J.B. Johnson. 2011.
Morphological and reproductive variation among populations of the fish Poecilia butleri.
Journal of Fish Biology 79:1029–1046.
Johnson, J.B., and J.C. Bagley. 2011. Ecological drivers of life-history divergence in Poeciliid
fishes. Pages 38–49 in J. Evans, A. Pilastro, and I. Schlupp, editors, Ecology and evolution of
livebearing fishes. University of Chicago Press, Chicago, IL.

24 25
Brook Trout
Salvelinus fontinalis
Leigh A. Johnson, Curator of Vascular Plants
Sérsic, A.N., A. Cosacov, A.A. Cocucci, L.A. Johnson, R.
Pozner, J. Avila, J.W. Sites Jr., and M. Morando. 2011. Emerging
phylogeographic patterns of plants and terrestrial vertebrates from
Patagonia. Biological Journal of the Linnean Society 103:475–494.
Leavitt, S.D., L.A. Johnson, T. Goward,
and L.L. St. Clair. 2011. Species
delimitation in taxonomically
difficult lichen–forming fungi: an
example from morphologically and
chemically diverste Xanthoparmelia
(Parmeliaceae) in North America.
Molecular Phylogenetics and
Evolution 60:317–332.
Leavitt, S.D., J.D. Fankhauser, D.H.
Leavitt, L.D. Porter, L.A. Johnson, and
L.L. St. Clair. 2011. Complex patterns
of speciation in cosmopolitan ‘‘rock
posy’’ lichens – discovering and
delimiting cryptic fungal species in
the lichen–forming Rhizoplaca melanophthalma species–complex
(Lecanoraceae, Ascomycota). Molecular Phylogenetics and
Evolution 59:587–602.
Leavitt, S.D., L.A. Johnson, and L.L. St. Clair. 2011. Species
delimitation and evolution in morphologically and chemically
diverse communities of the lichen–forming genus Xanthoparmelia
(Parmeliaceae, Ascomycota) in western North America. American
Journal of Botany 98:175–188.
Randy T. Larsen, Assistant Curator of Birds
Boyd, C., S. Petersen, W. Gilgert, R. Rodgers, S. Fuhlendorf, R.T. Larsen,
D. Wolfe, K.C. Jensen, P. Gonzales, M. Nenneman, R. Danvir, D.
Dahlgren, and T. Messmer. 2011. Looking toward a brighter future for
lekking birds [featured article]. Rangelands 34:1–10.
Larsen, R.T., J.A. Bissonette, A.C. Robinson, and J.T. Flinders. 2011.
Does small-perimeter fencing inhibit mule deer or pronghorn use of
water developments Journal of Wildlife Management 75:1417–1425.
Pierce, J.E., R.T. Larsen, J.T. Flinders, and J.C. Whiting. 2011.
Fragmentation of sagebrush communities: does an increase in
habitat edge impact pygmy rabbits Animal Conservation 14:314–
321.
C. Riley Nelson, Assistant Curator of Insects
Gates, T.A., J.R. Horner, R.R. Hanna, and C.R. Nelson. 2011. New
unadorned hadrosaurine hadrosaurid (Dinosauria, Ornithopoda)
from the Campanian of North America. Journal of Vertebrate
Paleontology 31(4):798–811.
Duke S. Rogers, Curator of Mammals
Corley, M.S., N. Ordóñez-Garza, D.S. Rogers, and R.D. Bradley. 2011. Molecular evidence for
paraphyly in Nyctomys sumichrasti: support for a new genus of vesper mice Occasional
Papers, The Museum of Texas Tech University 306:1–9.
Rogers, D.S., R.N. Leite, and R.J. Reed. 2011. Molecular phylogenetics of an endangered
species: the Tamaulipan woodrat (Neotoma angustapalata). Conservation Genetics 12:1035–
1048.
Rickart, E.A., R.J. Rowe, S.L. Robson, L.F. Alexander, and D.S. Rogers. 2011. Shrews of the Ruby
Mountains, northeastern Nevada. Southwestern Naturalist 56:95–102.
Dennis Shiozawa, Curator of Fishes
Shiozawa, D.K., R.P. Evans, P. Unmack, A. Johnson, and J. Mathis. 2011. Cutthroat trout
phylogenetic relationships with an assessment of associations among several subspecies.
Pages 158–166 in Wild Trout X: conserving wild trout. Proceedings of a Symposium. Trout
Unlimited, Arlington, VA.
Rasmussen, J.E., M.C. Belk, E. Habit, D.K. Shiozawa, R.D. Hepworth, and A. Anthony. 2011.
Variation in size-at-age between native cutthroat and introduced brown trout in allopatry
and sympatry: implications for competitive interaction. Aquatic Biology 13:285–292.
Houston, D.D., D.K. Shiozawa, and B.R. Riddle. 2011. The roles of Neogene geology and
late Pleistocene lake levels in shaping the genetic structure of the Lahontan redside shiner
Richardsonius egregious (Teleostei: Cyprinidae). Biological Journal of the Linnean Society
104:163–176.
Jack Sites, Curator of Reptiles and Amphibians
Werneck, F.P., G.C. Costa, G.R. Colli, D.E. Prado, and J.W. Sites Jr. 2011. Revisiting the historical
distribution of the seasonally dry tropical forest: new insights based on paleodistribution
modeling and palynological evidence. Global Ecology and Biogeography 20:272–288.
Nuñez, J.J., N. Koontz, F. Rabanal, F. Fontanella, and J.W. Sites Jr. 2011. Amphibian
phylogeography in the Antipodes: refugia and postglacial colonization explain mitochondrial
haplotype distributions in the Patagonian frog Eupsophus calcaratus (Cycloramphidae).
Molecular Phylogenetics and Evolution 58:343–352.
Breitman, M.F., L.J. Avila, J.W. Sites Jr., and M. Morando. 2011. Lizards from the end of the
world: phylogenetic relationships of the Liolaemus lineomaculatus section (Squamata:
Iguania: Liolaemini). Molecular Phylogenetics and Evolution 59:364–376.
Sérsic, A.N., A. Cosacov, A.A. Cocucci, L.A. Johnson, R. Poznar, L.J. Avila, J.W. Sites Jr., and M.
Morando. 2011. Emerging phylogeographic patterns in plants and terrestrial vertebrates from
Patagonia [invited review]. Biological Journal of the Linnean Society 103:475–494.
Sites, J.W., Jr., T.W. Reeder, and J.W. Wiens. 2011. Phylogenetic insights on evolutionary
novelties in lizards and snakes: sex, birth, bodies, food, and venom [invited review]. Annual
Review of Ecology, Evolution, and Systematics 42:227–244.
Mulcahy, D.G., T.H. Beckstead, and J.W. Sites Jr. 2011. Molecular systematics of the
Leptodeirini (Colubroidea, Dipsadidae) revisited: species tree analyses and multi-locus data.
Copeia 2011:407–417.
Martinez, L.E., L.J. Avila, C.H.F. Pérez, D.R. Pérez, J.W. Sites Jr., and M. Morando. 2011. A
new species of Liolaemus (Squamata, Iguania, Liolaemini) endemic to the Auca Mahuida
volcano, northwestern Patagonia, Argentina. Zootaxa 3010:31–46.
Townsend, T.M., D.G. Mulcahy, B.P. Noonan, J.W. Sites Jr., C.A. Kuczynski, J.J. Wiens, and T.W.

26 27
King Salmon
Oncorhynchus tshawytscha
Reeder. 2011. The molecular systematics of Iguania (Squamata)
inferred from 29 nuclear loci. Molecular Phylogenetics and Evolution
61:363–380.
Olave, M., L.E. Martinez, L.J. Avila, J.W. Sites Jr., and M. Morando.
2011. Evidence of hybridization in the Argentinian lizards Liolaemus
gracilis and L. bibronii (Iguania: Liolaemini): an integrative
approach based on genes and morphology. Molecular
Phylogenetics and Evolution 61:381–391.
Pellegrino, K.C.M., M.T. Rodrigues, D.J. Harris, Y.Y.
Yassuda, and J.W. Sites Jr. 2011. Molecular phylogeny,
biogeography, and insights into the origin of
parthenogenesis in the Neotropical genus Leposoma
(Squamata: Gymnophthalmidae): ancient links
between the Atlantic Forest and Amazonia. Molecular
Phylogenetics and Evolution 61:446–459.
Heideman, N.J.L., J.W. Sites Jr., D.G. Mulcahy, M.G.J.
Hendricks, and S.R. Daniels. 2011. Cryptic diversity and
morphological convergence in threatened species
of fossorial skinks in the genus Scelotes (Squamata:
Scincidae) from the Western Cape Coast of South
Africa: implications for species boundaries, digit
reduction and conservation. Molecular Phylogenetics
and Evolution 61:823–833.
Breitman, M.F., C.H.F. Perez, M. Parra, M. Morando, J.W.
Sites Jr., and L.J. Avila. 2011. New species of lizard from
the magellanicus clade of the Liolaemus lineomaculatus
section (Squamata: Iguania: Liolaemini) from southern
Patagonia. Zootaxa 3123:32–48.
Breitman, M.F., M. Parra, C.H.F. Perez, and J.W. Sites Jr. 2011. Two
new species of lizards from the Liolaemus lineomaculatus section
(Squamata: Iguania: Liolaemini) from southern Patagonia. Zootaxa
3120:1–28.
Mendelson, J.R., III, D.G. Mulcahy, T.S. Williams, and J.W. Sites Jr. 2011.
A phylogeny and evolutionary natural history of Mesoamerican toads
(Anura: Bufonidae: Incilius). Zootaxa 3138:1–34.
Larry St. Clair, Curator of Nonvascular Cryptogams
Glacy, L.A., S.D. Leavitt, and L.L. St. Clair. 2011. A checklist of the
lichens collected during the 2008 Sixth International Association of
Lichenology field trip at Point Reyes National Seashore and selected
locations in Marin and Sonoma counties, California, USA. Evansia
28(1):18–26.
Leavitt, S.D., L. Johnson, and L.L. St. Clair. 2011. Species delimitation
and evolution in morphologically and chemically diverse
communities of the lichen-forming genus Xanthoparmelia
(Parmeliaceae, Ascomycota) in western North America. American
Journal of Botany 98(2):175–188.
Leavitt, S.D., J.D. Fankhauser, D.H. Leavitt, L.D. Porter, L.A.
Johnson, and L.L. St. Clair. 2011. Complex patterns of speciation
in cosmopolitan “rock posy” lichens – Discovering and delimiting
cryptic fungal species in the lichen-forming Rhizoplaca
melanophthalma species-complex (Lecanoraceae, Ascomycota). Molecular Phylogenetics
and Evolution 59:587–602.
Leavitt, S.D., L.A. Johnson, T. Goward, and L.L. St. Clair. 2011. Species delimitation in
taxonomically difficult lichen forming fungi: an example from morpholically and chemically
diverse Xanthoparmelia (Parmeliacea) in North America. Molecular Phylogenetics and
Evolution 60:317–332.
Shrestha, G., and L.L. St. Clair. 2011. A comparison of the lichen floras of four locations in the
Intermountain Western United States. North American Fungi 6(8):1–20.
Leavitt, S.D., and L.L. St. Clair. 2011. Estimating Xanthoparmelia (Parmeliaceae) population
density in subalpine communities in southern Utah, U.S.A. using two distance methods, with
implications for assessing community composition. Bryologist 114(3):625–636.
Clayton White, Emeritus Curator of Birds
Jones, L., H.L. Black, and C.M. White. 2011. Evidence for convergent evolution in gape
morphology of the Bat Hawk (Macheirampus alcinus) with swifts, swallows and goatsuckers.
Biotropica 44:386–393.
Jones, L.R., H.L. Black, C.M. White, N.P. Johnston, M.E. McGee, S.W. Donahue, and D.L. Eggett.
2011. Effects of calcium-loading on egg production in Ring-necked Pheasants. Journal of
Wildlife Management 74:1295–1300.
Talbot, S.L., A.C. Palmer, G.K. Sage, S.A. Sonsthagen, T. Swem, D.J. Brimm, and C.M. White.
2011. Lack of genetic polymorphism among Peregrine Falcons Falco peregrinus of Fiji. Journal
of Avian Biology 42:415–428.
Matz, A., T. Swem, P. Johnson, T. Booms, and C.M. White. 2011. Potential for climate change to
increase contaminants exposure and effects on Gyrfalcons. Pages 161–175 in R.T. Watson, T.J.
Cade, M. Fuller, G. Hunt, and E. Potopov, editors, Gyrfalcons and ptarmigan in a changing
world. Volume 1. The Peregrine Fund, Inc., Boise, ID.
White, C.M. 2011. Thoughts on the interface between Mormonism, the environment, and
evolution: connection and dilemma. Articles and Essays, Dialogue, June 13.
Michael Whiting, Curator of Insects
Cameron, S.L., K. Yoshizawa, A. Mizukoshi, M.F. Whiting, and K.P. Johnson. 2011. Mitochondrial
genome deletions and minicircles are common in lice (Insecta: Phthiraptera). BMC Genomics
12:394–409.
Gontijo, A.M., V. Miguela, M.F. Whiting, R.C. Woodruff, and M. Dominguez. 2011. Intron
retention in the Drosophila melanogaster Rieske iron sulphur protein gene generated a new
protein. Nature Communications 2:323 [http://dx.doi.org/10.1038/ncomms1328].
Beutel, R.G., F. Friedrich, T. Hörnschemeyer, H. Pohl, F. Hünefeld, F. Beckmann, R. Meier, B.
Misof, M.F. Whiting, and L.B. Vilhelmsen. 2011. Morphological and molecular evidence
converge upon a robust phylogeny for the megadiverse Holometabola. Cladistics 27:341–
355.
Whiting, M.F. 2011. Evolution and the gospel: seeking grandeur in this view of life. In: M.D.
Rhodes and J.W. Moody, editors, Converging paths to truth: the Summerhays lectures on
science and religion. Deseret Book, Salt Lake City, UT.