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BIAWAK - International Varanid Interest Group

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<strong>BIAWAK</strong> VOL. 4 NO. 4<br />

124<br />

Fig. 1. The evolution and dispersal route of varanid lizards. Laurasian hypothesis is based on fossil records (Estes,<br />

1983) and indicated by grey arrows, whereas Gondwanan hypothesis is based on species diversity (Hutchinson &<br />

Donellan,<br />

Figure 1. The1993) evolution<br />

and<br />

andindicated dispersal route<br />

by<br />

of<br />

white<br />

varanid<br />

arrows.<br />

lizards. Laurasian<br />

Map shows<br />

hypothesis<br />

Late<br />

isCretaceous-Tertiary based on fossil records (Estes,<br />

transition<br />

1983) and<br />

(∼65<br />

indicated<br />

Ma),<br />

inredrawn<br />

grey<br />

after arrows, a whereas paleogeographic Gondwanan hypothesis map available is based on at species http://jan.ucc.nau.edu/~rcb7/globaltext2.html diversity (Hutchinson & Donellan, 1993) and indicated in by white Ron arrows. Blakey, Map shows Geology<br />

Department, Late Cretaceous-Tertiary Northern transition Arizona (∼65University. Ma), redrawn after Laurasian a paleogeographic and Gondwanan map availablefragments at http://jan.ucc.nau.edu/~rcb7/globaltext2.html are indicated in this map with by the<br />

Ron Blakey, at Geology Department, Northern Arizona University. Laurasian and Gondwanan fragments are indicated in this map with the letter<br />

letters “L” and “G”, respectively.<br />

„L“ and „G“, respectively.<br />

L<br />

L<br />

L<br />

G<br />

G<br />

X<br />

G<br />

G<br />

G<br />

G<br />

G<br />

G<br />

Fig. 2. Hypothetical dispersal of ancestral varanids from Asia as illustrated in Estes (1983). <strong>Varanid</strong> lizards are<br />

thought to have originated in central Asia during the Cretaceous (~80 Ma) based on fossils found in Mongolia,<br />

whereas the genus Varanus is thought to have emerged in Africa sometime in the Eocene, more than 37 Ma (Holmes,<br />

2010).<br />

Received: 16 April 2010; Accepted 4 November 2010

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