A Review of the Genus Eunice - Smithsonian Institution Libraries

150 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
rior acicular lobes rounded; aciculae emerging at midline.
Pre- and postsetal lobes low, transverse folds. First 4 ventral
cirri thick, tapering. Ventral cirri basally inflated from about
setiger 5. Inflated bases short welts where best developed,
decreasing in size from about setigcr 45; narrow tips tapering.
Posterior ventral cirri thick, tapering, directed posteriorly.
Anterior notopodial cirri tapering and articulated with up to 5
articulations, farther posteriorly cirri decreasing in length,
without articulations.
Limbate setae slim, marginally smooth. Shafts of pectinate
setae (Figure 48j) cylindrical. Blades slightly flaring, flat. One
marginal tooth longer than other teeth; -12 teeth present.
Compound falcigers (Figure 48k) very large, long-shafted.
Shafts tapering, marginally smooth; distal beaks absent.
Appendages with distinctly angled back edge, tapering,
bidentatc. Proximal teeth longer than distal teeth, tapering,
directed laterally. Distal teeth gently curved, nearly erect.
Guards longer than appendages, angular, but distinctly symmetrically
bluntly pointed; mucros absent. Pscudocompound
falcigers and compound spinigcrs absent. Aciculae (Figure 48i)
paired, with light brown or chestnut-colored cores and clear
sheaths, tapering, slender, gently curved tips; tips curved
dorsally; cross-sections round. Subacicular hooks (Figure 48h)
with chestnut-colored cores and clear sheaths, bidentatc. Hooks
first present from setiger 27, present in all sctigcrs thereafter,
paired in some sctigers. Hooks slender, not distinctly larger
than compound falcigers, tapering, with small heads. Proximal
teeth larger than distal teeth, triangular, directed laterally.
Distal teeth short, nearly erect. Paired bosses present on distal
bases of proximal teeth. Bases of proximal teeth much narrower
than shaft at that point.
UNKNOWN MORPHOLOGICAL FEATURES.—Jaw structure; all
features associated with posterior setigers, including branchial
distribution; pygidium and anal cirri.
EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—
Mx III short and forming part of distal arc with left Mx IV.
CHARACTERS USED IN PREPARATION OF KEY NOT
SCORED.—Inappropriate Characters: 34, 56, 58, 59. Unknown
Characters: 1, 2, 13, 14, 36-38,40, 47, 50, 63.
ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—37,1;
38,1.
REMARKS.—Eunice frauenfeldi is assumed to belong to
group B-2 and is listed with similar species in Tables 27 and 32;
the reduction in size of the branchiae towards the posterior end
of the incomplete fragment makes this assumption somewhat
suspect. The problem cannot be resolved without access to
more material from the type locality. The synonymy list given
in Hartman (1964:118) must be revised: Eunice magellanica is
here treated as a distinct species. The only other species with
palmate branchiae listed in Table 32 is E. rubella. The two
species differ in the proportions of the prostomium and in
various shape-related characters best elucidated by comparing
the illustrations of the two species.
72. Eunice fucata Ehlers, 1887
FIGURE 49a-d; TABLES 27,28, 50
Eunice fucata Ehlers. 1887:91-93, pi. 25: figs. 8-20.
Staurocephalus gregaricus Mayer, 1900:1.
Mayeria gregaricus.—Vcrrill, 1900:650.
MATERIAL EXAMINED.—MCZ, 3 posterior fragments from
type lots, Tortugas, Florida.
COMMENTS ON MATERIAL EXAMINED.—The rc-description
is based on the original description, complemented where
possible by observations on the original material.
DESCRIPTION.—One specimen complete, with 250 segments;
total length 195 mm; width ~5 mm; length through
sctiger 10 -10 mm (based on illustration of anterior end).
Posterior body roughly circular in cross-section with no
indication of dorsovcntral flattening.
Eyes present. Ccratostylcs without articulations. A-I to
posterior pcristomial ring; A-I I and A-I 11 to setiger 3. Anterior
per is torn ial ring 3 /" of total pcristomial length. Peristomial cirri
to middle of prostomium, tapering, without articulations.
Maxillary formula 1+1, 4+5, 6+0, 3 + 8, 1 + 1, and 1+1.
Branchiae (Figure 49d) present, pectinate, distinctly longer
than notopodial cirri, not reduced in mid-body region, erect.
Branchiae from setigcr 5, absent on last 70-80 sctigcrs.
Branchiae terminating well before posterior end, present on
more than 65% of total number of setigcrs. Last 30 pairs single
filaments; maximum 12 primary filaments; primary filaments
sometimes pectinate, especially towards posterior end, yielding
up to 24 or more terminal filaments. Branchiae longer than
notopodial cirri except in last 15 branchial setigers. Filaments
digitiform, shorter than notopodial cirri.
Neuropodial acicular lobes obliquely truncate with aciculae
emerging at upper, high end of lobes. Pre- and postsetal lobes
low, transverse folds. Median ventral cirri basally inflated.
Inflated bases missing posteriorly, in fragments available, all
ventral cirri thick, digitiform. Anterior and median notopodial
cirri basally inflated, tapering to digitiform tips, without
articulations. Posterior notopodial cirri short, digitiform.
Limbate setae very narrowly limbate, marginally smooth.
Pectinate setae (Figure 49b) very long in posterior setigers.
Shafts cylindrical. Blades strongly flaring, furled. Marginal
teeth no longer than other teeth, with 20 teeth. Compound
falcigers broken in all sctigers examined. Shafts described as
slightly inflated, marginally smooth. Appendages very short,
triangular. Cutting edge with series of small, irregular teeth,
rather than single proximal teeth. Distal teeth nearly erect.
Guards asymmetrically bluntly to sharply pointed; mucros
absent. Pseudocompound falcigers and compound spinigers
absent. Aciculae (Figure 49c) paired, with dark brown cores
and clear sheaths, tapering to slender lips, gently curved or
straight; cross-sections round. Subacicular hooks (Figure 49a)
with light to medium brown cores and clear sheaths, unidentate.
Hooks distally gently curved, without any traces of other teeth.